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  <front>
    <journal-meta>
<journal-id journal-id-type="publisher">BG</journal-id>
<journal-title-group>
<journal-title>Biogeosciences</journal-title>
<abbrev-journal-title abbrev-type="publisher">BG</abbrev-journal-title>
<abbrev-journal-title abbrev-type="nlm-ta">Biogeosciences</abbrev-journal-title>
</journal-title-group>
<issn pub-type="epub">1726-4189</issn>
<publisher><publisher-name>Copernicus GmbH</publisher-name>
<publisher-loc>Göttingen, Germany</publisher-loc>
</publisher>
</journal-meta>

    <article-meta>
      <article-id pub-id-type="doi">10.5194/bg-12-467-2015</article-id><title-group><article-title>Carbon storage versus albedo change: radiative forcing of forest expansion
in temperate mountainous regions of Switzerland</article-title>
      </title-group><?xmltex \runningtitle{Carbon storage versus albedo change}?><?xmltex \runningauthor{J.~Schwaab et al.}?>
      <contrib-group>
        <contrib contrib-type="author" corresp="yes" rid="aff1 aff5">
          <name><surname>Schwaab</surname><given-names>J.</given-names></name>
          <email>jonasschwaab@ethz.ch</email>
        </contrib>
        <contrib contrib-type="author" corresp="no" rid="aff1">
          <name><surname>Bavay</surname><given-names>M.</given-names></name>
          
        <ext-link>https://orcid.org/0000-0002-5039-1578</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff4">
          <name><surname>Davin</surname><given-names>E.</given-names></name>
          
        <ext-link>https://orcid.org/0000-0003-3322-9330</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff2">
          <name><surname>Hagedorn</surname><given-names>F.</given-names></name>
          
        <ext-link>https://orcid.org/0000-0001-5218-7776</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff3">
          <name><surname>Hüsler</surname><given-names>F.</given-names></name>
          
        </contrib>
        <contrib contrib-type="author" corresp="no" rid="aff1 aff6">
          <name><surname>Lehning</surname><given-names>M.</given-names></name>
          
        <ext-link>https://orcid.org/0000-0002-8442-0875</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff1">
          <name><surname>Schneebeli</surname><given-names>M.</given-names></name>
          
        <ext-link>https://orcid.org/0000-0003-2872-4409</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff2">
          <name><surname>Thürig</surname><given-names>E.</given-names></name>
          
        </contrib>
        <contrib contrib-type="author" corresp="no" rid="aff1">
          <name><surname>Bebi</surname><given-names>P.</given-names></name>
          
        </contrib>
        <aff id="aff1"><label>1</label><institution>WSL – Institute for Snow and Avalanche Research SLF, 7260 Davos,
Switzerland</institution>
        </aff>
        <aff id="aff2"><label>2</label><institution>Swiss Federal Institute for Forest, Snow and Landscape Research
WSL, 8903 Birmensdorf, Switzerland</institution>
        </aff>
        <aff id="aff3"><label>3</label><institution>Institute of Geography and Oeschger Center for Climate Change
Research, University of Bern, 3012 Bern, Switzerland</institution>
        </aff>
        <aff id="aff4"><label>4</label><institution>Institute for Atmospheric and Climate Sciences, ETH Zürich, 8092
Zürich, Switzerland</institution>
        </aff>
        <aff id="aff5"><label>5</label><institution>Institute for Spatial and Landscape Planning, ETH Zürich, 8093
Zürich, Switzerland</institution>
        </aff>
        <aff id="aff6"><label>6</label><institution>CRYOS, School of Architecture, Civil and Environmental Engineering,
EPFL, 1015 Lausanne, Switzerland</institution>
        </aff>
      </contrib-group>
      <author-notes><corresp id="corr1">J. Schwaab (jonasschwaab@ethz.ch)</corresp></author-notes><pub-date><day>27</day><month>January</month><year>2015</year></pub-date>
      
      <volume>12</volume>
      <issue>2</issue>
      <fpage>467</fpage><lpage>487</lpage>
      <history>
        <date date-type="received"><day>27</day><month>May</month><year>2014</year></date>
           <date date-type="rev-request"><day>30</day><month>June</month><year>2014</year></date>
           <date date-type="rev-recd"><day>11</day><month>November</month><year>2014</year></date>
           <date date-type="accepted"><day>1</day><month>December</month><year>2014</year></date>
           
      </history>
      <permissions>
<license license-type="open-access">
<license-p>This work is licensed under a Creative Commons Attribution 3.0 Unported License. To view a copy of this license, visit <ext-link ext-link-type="uri" xlink:href="http://creativecommons.org/licenses/by/3.0/">http://creativecommons.org/licenses/by/3.0/</ext-link></license-p>
</license>
</permissions><self-uri xlink:href="https://bg.copernicus.org/articles/.html">This article is available from https://bg.copernicus.org/articles/.html</self-uri>
<self-uri xlink:href="https://bg.copernicus.org/articles/.pdf">The full text article is available as a PDF file from https://bg.copernicus.org/articles/.pdf</self-uri>


      <abstract>
    <p>In this study, we assess the climate mitigation potential from afforestation
in a mountainous snow-rich region (Switzerland) with strongly varying
environmental conditions. Using radiative forcing calculations, we quantify
both the carbon sequestration potential and the effect of albedo change at
high resolution. We calculate the albedo radiative forcing based on remotely
sensed data sets of albedo, global radiation and snow cover. Carbon
sequestration is estimated from changes in carbon stocks based on national
inventories. We first estimate the spatial pattern of radiative forcing (RF) across Switzerland
assuming homogeneous transitions from open land to forest. This highlights
where forest expansion still exhibits climatic benefits when including the
radiative forcing of albedo change. Second, given that forest expansion is
currently the dominant land-use change process in the Swiss Alps, we
calculate the radiative forcing that occurred between 1985 and 1997. Our
results show that the net RF of forest expansion ranges from <inline-formula><mml:math display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>24 W m<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:msup></mml:math></inline-formula>
at low elevations of the northern Prealps to 2 W m<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:msup></mml:math></inline-formula> at high elevations
of the Central Alps. The albedo RF increases with increasing altitude, which
offsets the CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> RF at high elevations with long snow-covered periods,
high global radiation and low carbon sequestration. Albedo RF is
particularly relevant during transitions from open land to open forest but
not in later stages of forest development. Between 1985 and 1997, when
overall forest expansion in Switzerland was approximately 4 %, the albedo
RF offset the CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> RF by an average of 40 %. We conclude that the
albedo RF should be considered at an appropriately high resolution when
estimating the climatic effect of forestation in temperate mountainous
regions.</p>
  </abstract>
    </article-meta>
  </front>
<body>
      

<sec id="Ch1.S1" sec-type="intro">
  <title>Introduction</title>
      <p>The United Nations Framework Convention on Climate Change (UNFCC) declared
in the Kyoto Protocol (Decision 11/CP.7) that changes in the carbon stocks
of ecosystems, induced by LULUCF (land use, land-use change and forestry)
activities can be included in the greenhouse gas emission budget of the
signatory nations  (UNFCC, 2001). Beside biogeochemical processes,
LULUCF also influences biogeophysical processes  (Betts, 2011; Bonan,
2008), but these effects are not yet considered in current climate policies.</p>
      <p>Global climate models suggest that biogeochemical and biogeophysical effects
vary greatly with latitude  (Schaeffer et al., 2006; Bala et al., 2007;
Bathiany et al., 2010; Davin and de Noblet-Ducoudre, 2010; Arora and
Montenegro, 2011). In the tropics, biogeochemical and biogeophysical effects
tend to act in the same direction, since tropical forests cool climate
through both evaporative cooling and carbon sequestration  (Costa and
Foley, 2000; Gibbard et al., 2005). However, at middle and high latitudes,
biogeophysical processes tend to counter the biogeochemical effect
(Gibbard et al., 2005; Betts, 2000; Govindasamy et al., 2001), thus
making the net LULUCF effect more challenging to assess. Indeed, forestation
in boreal regions lowers the albedo and thus counterbalances the cooling
effect of carbon storage.</p>
      <p>Global climate models are important for understanding the climatic processes
related to LULUCF and for quantifying the impacts on climate of forestation
or deforestation over large areas. However, in highly heterogeneous
landscapes, such as mid-latitude mountain ranges, global climate models are
limited by their relatively coarse resolution, and the concept of radiative forcing (RF) (Myhre et al., 2013) can provide a useful
alternative. The RF concept has already been employed to investigate the
balance between biogeophysical (mainly albedo) and biogeochemical effects
following forestation or deforestation. While some studies suggest that
albedo RF can completely offset CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> RF  (Betts, 2000; Bernier et al.,
2011; de Wit et al., 2013), others have found that the offset is rather
small (e.g.  Montenegro et al., 2009; Kirschbaum et al., 2011). The
offset seems to vary widely depending on the regional characteristics of the
determining variables: global radiation, snow cover, albedo change and
carbon sequestration. These factors vary greatly, but little research has
focused on how each of them influences RF  (Kirschbaum et al., 2011), and
only a few attempts have been made to quantify RF in a spatially explicit way
(e.g. Betts, 2000; Montenegro et al., 2009). Betts (2000) estimated
spatially explicit RF data on a resolution of 3.75<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula> longitude by
2.5<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula> latitude. Montenegro et al. (2009) performed their analysis on
a resolution of 5 to 25 km. However, RF varies on much smaller scales.
Moreover, decisions in regional planning are usually based on very local and
regional information; therefore, it is crucial to quantify RF at finer
resolutions.</p>
      <p>Forest cover has increased in many temperate mountainous regions  (Alewell
and Bebi, 2011; MacDonald et al., 2000; Ramankutty et al., 2010; Kozak,
2003; Hagedorn et al., 2014) and analysis of LULUCF change in the Alps
suggests that changes in forest cover near the treeline will further
increase (Gehrig-Fasel et al., 2007). Some of the effects of
this increase on various ecosystem services are already relatively well
known  (MacDonald et al., 2000; Laiolo et al., 2004; Bolliger et al.,
2008) and have increasingly been considered in management strategies and
subsidizing systems for agriculture and forestry  (e.g. Gret-Regamey et
al., 2013). However, few attempts to quantify RF in such regions have been
made, even though snow cover and hence albedo in temperate mountains varies
greatly. To optimize the effects of future land-use decisions, further
research on the climatic impacts of forests is essential and should be
included in spatially explicit valuation methods (Bebi et al., 2012).</p>
      <p>Switzerland is a particular suitable study area for researching the effects
of changes in forest cover, since many spatially explicit high-quality
data sets are available and forest expansion is an ongoing dominant process
of land-use change. The spatially explicit data series available are on land use/land cover (LULC) (1 ha raster), snow cover (1 km), global radiation (2.2 km),
albedo change and carbon sequestration (both explicit for biogeographical
regions) for the whole area of Switzerland. Forest cover is expanding by
4 % per decade at the country scale and by 8 % per decade in alpine
areas (National Forest Inventory, NFI).</p>
      <p>Our study design is twofold: first, we use the spatially explicit data sets
to show the pattern of RF assuming that each location in Switzerland is
facing a transition from agriculturally used open land to forest. This is
not related to any particular or realistic scenario; however, the spatial
pattern of RF can be of high interest for any land-use policies steering
forest-cover change towards desired futures. In Switzerland agricultural
subsidies directly influence farmers decisions on whether to keep managing
or abandon their land. The latter will usually result in forest expansion.
Second, we include the type and location of five different land-use transitions
to calculate RF in Switzerland between 1985 and 1997. In summary, we estimate
(i) to which extent albedo RF offsets CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> RF in different parts of temperate
mountainous regions, (ii) how each input parameter influences RF and (iii)
what the inclusion of albedo change implies for the greenhouse gas inventory
in Switzerland.</p>
</sec>
<sec id="Ch1.S2">
  <title>Data and methods</title>
<sec id="Ch1.S2.SS1">
  <title>Study area</title>
      <p>Switzerland covers an area of 41 284 km<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msup></mml:math></inline-formula> and can be divided into five
biogeographical regions (Fig. 1). Each region has
biogeographical features that can be found globally at temperate latitudes: (1) Jura – oceanic low mountain range with elevations averaging 800 m a.s.l.;
(2) Plateau – oceanic lowlands with elevations averaging 550 m a.s.l.; (3)
northern Prealps – oceanic subalpine mountain range with elevations averaging
1400 m a.s.l.; (4) Central Alps – continental alpine mountain range with
elevations averaging 2150 m a.s.l.; (5) southern Prealps –
Mediterranean/Insubric line subalpine mountain range with elevations averaging
1500 m a.s.l. In each region, deciduous forests and mixed deciduous forest
dominate at low elevations (mostly <italic>Fagus sylvatica</italic>), while
coniferous forests are dominant at higher elevations (mostly <italic>Picea abies</italic>).</p>
      <p>The Swiss landscape has been strongly affected by several centuries of intensive
human land-use (Bürgi and Schuler, 2003; Schneeberger et al., 2007; Gimmi
et al., 2009) followed by the widespread abandonment of marginal agricultural
land and the subsequent expansion of forest cover since the end of the 19th
century (Baur, 2006). Due to unfavorable pedologic and climatic conditions
and high slope angles, marginal land and forest expansion are mainly found at
higher elevations (Baur, 2006). Land abandonment was the most dominant driver
for the establishment of new forest areas; however, a small fraction of
forest expansion at the treeline can be attributed to recent climate
warming (Gehrig-Fasel et al., 2007). The treeline is not only often the result of former
land use but also depends on various climatic factors and is generally
higher in the Central Swiss Alps (approx. 2100–2300 m) than in the northern
and southern Prealps (approx. 1800–2000 m) (Fig. 3f; Körner, 2012).</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F1"><caption><p>Digital elevation model and biogeographical regions of Switzerland.</p></caption>
          <?xmltex \igopts{width=241.848425pt}?><graphic xlink:href="https://www.biogeosciences.net/12/467/2015/bg-12-467-2015-f01.png"/>

        </fig>

</sec>
<sec id="Ch1.S2.SS2">
  <title>Spatial variability of RF and RF of Swiss forest expansion</title>
      <p>We calculated the net RF and the offset of CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> RF through albedo RF
(<inline-formula><mml:math display="inline"><mml:mi mathvariant="normal">Δ</mml:mi></mml:math></inline-formula>RF<inline-formula><mml:math display="inline"><mml:mrow><mml:msub><mml:mi/><mml:mrow><mml:mi>C</mml:mi><mml:msub><mml:mi>O</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:mrow></mml:msub><mml:mo>/</mml:mo><mml:mi mathvariant="normal">Δ</mml:mi></mml:mrow></mml:math></inline-formula>RF<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mi mathvariant="normal">albedo</mml:mi></mml:msub></mml:math></inline-formula>) to show the
spatial variability of RF in Switzerland and to calculate RF of Swiss
forest-cover expansion between 1985 and 1997. To illustrate the pattern of RF
in Switzerland we calculated a value of RF for every location in Switzerland,
excluding non-vegetated land, water, settlement and areas that lie above the
treeline (Fig. 3). These are hypothetical values because we calculated RF
for the change from open land to forest for all vegetated areas, and not only
for the ones where forest expansion was actually observed. At lower
elevations, transitions from intensively used open land to forest are
frequent, while in higher elevations transitions from extensively used open
land to forest are most likely (Table A4). We considered this by
separating our estimation of the hypothetical RF in transitions from
intensively used open land to forest below 1000 m and transitions from
extensively used open land to forest above 1000 m. The results of the
spatial variability of RF are shown in maps of the study area (Fig. 4c ,d).</p>
      <p>To obtain results for RF of forest-cover expansion between 1985 and 1997 in
Switzerland we calculated net radiative forcing as the sum of RF for all
pixels where forests expanded. This meant including information on the type
of forest expansion and on the location of forest expansion:
            <disp-formula id="Ch1.E1" content-type="numbered"><mml:math display="block"><mml:mrow><mml:mi mathvariant="normal">Δ</mml:mi><mml:msub><mml:mi mathvariant="normal">RF</mml:mi><mml:mi mathvariant="normal">ges</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:mstyle displaystyle="true"><mml:mfrac style="display"><mml:mrow><mml:msubsup><mml:mo>∑</mml:mo><mml:mrow><mml:mi>x</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow><mml:mi>n</mml:mi></mml:msubsup><mml:msub><mml:mi mathvariant="normal">RF</mml:mi><mml:mrow><mml:mi>x</mml:mi><mml:mo>,</mml:mo><mml:mi>T</mml:mi></mml:mrow></mml:msub></mml:mrow><mml:mrow><mml:msub><mml:mi>A</mml:mi><mml:mi mathvariant="normal">E</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>,</mml:mo></mml:mrow></mml:math></disp-formula>
          where <inline-formula><mml:math display="inline"><mml:mi mathvariant="normal">Δ</mml:mi></mml:math></inline-formula>RF<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mi mathvariant="normal">ges</mml:mi></mml:msub></mml:math></inline-formula> is the net RF, <inline-formula><mml:math display="inline"><mml:mi>n</mml:mi></mml:math></inline-formula>
the number of pixels where forests expanded and RF the radiative forcing,
which depends on the location <inline-formula><mml:math display="inline"><mml:mi>x</mml:mi></mml:math></inline-formula> and the type of transition <inline-formula><mml:math display="inline"><mml:mi>T</mml:mi></mml:math></inline-formula>. The sum
over RF is divided by the the earth's surface <inline-formula><mml:math display="inline"><mml:mrow><mml:msub><mml:mi>A</mml:mi><mml:mi mathvariant="normal">E</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> to convert local
RF into a global average RF (de Wit et al., 2014).</p>
</sec>
<sec id="Ch1.S2.SS3">
  <title>Swiss forest expansion between 1985 and 1997</title>
      <p>We use aerial photographs processed by Swiss Statistics at a spatial
resolution of 100 m to derive changes in LULC. These
aerial photographs are from the Swiss Federal Office of Topography and are
fully available for the two inventory periods: 1979–1985 and 1992–1997
(Humbel et al., 2010). We reclassified the data of the different
inventory periods into five aggregated classes (Rutherford et al., 2008).
While 18 classes were aggregated into four classes: closed forest, open
forest, extensively and intensively used open land (Fig. 2), the remaining 56 were classified as other and
consisted mainly of settlements, water and non-vegetated land (Table A5). The
aggregation of the original land-use classes results in a sufficiently large
sample of reliable albedo values and carbon stocks in each of the five
biogeographical regions and three elevational strata for five relevant and
well-established land-use classes.</p>
      <p>To calculate RF of land-use change between 1985 and 1997, we included five
transitions: (1) intensively used open land <inline-formula><mml:math display="inline"><mml:mo>→</mml:mo></mml:math></inline-formula> closed forest, (2)
extensively used open land <inline-formula><mml:math display="inline"><mml:mo>→</mml:mo></mml:math></inline-formula> closed forest, (3) intensively used
open land <inline-formula><mml:math display="inline"><mml:mo>→</mml:mo></mml:math></inline-formula> open forest, (4) extensively used open land
<inline-formula><mml:math display="inline"><mml:mo>→</mml:mo></mml:math></inline-formula> open forest and (5) open forest <inline-formula><mml:math display="inline"><mml:mo>→</mml:mo></mml:math></inline-formula> closed forest.
Forest expansion mainly took place in elevations above 1200 m in the Prealps
and the Central Alps (Fig. A1). We focused on transitions where forest cover
and carbon stocks increase because these transitions highly exceeded
transitions with forest decrease in Switzerland. In fact, Swiss law
strongly protects forests, so there have been only a few changes from
forest to agriculturally used land during the last 30 years (Bloetzer, 2004,
Rutherford et al., 2008).</p>
</sec>
<sec id="Ch1.S2.SS4">
  <?xmltex \opttitle{CO${}_{{2}}$ RF}?><title>CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> RF</title>
      <p>An increase in carbon stocks in terrestrial ecosystems is related to a sink
of atmospheric CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula>, followed by a change in the earth's radiation
balance. Myhre et al. (1998) developed a parameterization to derive RF,
related to a change in the CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> concentration in the atmosphere, based on
radiative transfer schemes:
            <disp-formula id="Ch1.E2" content-type="numbered"><mml:math display="block"><mml:mrow><mml:mi mathvariant="normal">Δ</mml:mi><mml:mi mathvariant="normal">RF</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo><mml:mo>=</mml:mo><mml:mn>5.35</mml:mn><mml:mi>ln⁡</mml:mi><mml:mstyle displaystyle="true"><mml:mfrac style="display"><mml:mrow><mml:mi>C</mml:mi><mml:mspace width="0.125em" linebreak="nobreak"/><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow><mml:mrow><mml:msub><mml:mi>C</mml:mi><mml:mn mathvariant="normal">0</mml:mn></mml:msub></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>,</mml:mo></mml:mrow></mml:math></disp-formula>
          where <inline-formula><mml:math display="inline"><mml:mi mathvariant="normal">Δ</mml:mi></mml:math></inline-formula>RF(t) is the Radiative Forcing, <inline-formula><mml:math display="inline"><mml:mrow><mml:mi>C</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> is the atmospheric
CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> concentration after perturbation and <inline-formula><mml:math display="inline"><mml:mrow><mml:msub><mml:mi>C</mml:mi><mml:mn mathvariant="normal">0</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> is the unperturbed
atmospheric CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> concentration. Equation (2) can be solved if the
CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> concentration in the atmosphere after perturbation is known.</p>
      <p>Following Switzerland's Greenhouse Gas Inventory 1990–2010 (Heldstab et al.,
2012), which is based on the Good Practice Guidance for Land Use, Land Use
Change and Forestry (IPCC, 2003), we calculated the carbon stock changes
resulting from land-use changes as the differences between the carbon stocks
of the land-use categories before and after a transition. This takes into
account changes in living plant biomass, dead wood and soil carbon stocks.</p>
      <p>Data on carbon stocks in the living biomass and dead wood of closed forests
and open forests were derived from the third NFI
(Brändli, 2010). The data are based on 6608 field plots measured on a
regular sampling 1.4 km <inline-formula><mml:math display="inline"><mml:mo>×</mml:mo></mml:math></inline-formula> 1.4 km grid from 2004 to 2006. Two 200 m<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msup></mml:math></inline-formula> concentric
circles are used for trees with diameter at breast height (DBH) <inline-formula><mml:math display="inline"><mml:mo>≥</mml:mo></mml:math></inline-formula> 12 cm but <inline-formula><mml:math display="inline"><mml:mo>&lt;</mml:mo></mml:math></inline-formula> 36 cm, and two 500 m<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msup></mml:math></inline-formula> concentric
circles for
trees with DBH <inline-formula><mml:math display="inline"><mml:mo>≥</mml:mo></mml:math></inline-formula> 36 cm. This results in DBH measurements of
approximately 11 trees per plot. On a sub-sample of approximately two trees
per plot, the diameter at 7 m tree height and the tree height are measured.
The biomass of all single trees is estimated according to allometric
functions. The assessment of stem wood over bark, including stock, coarse
branches (<inline-formula><mml:math display="inline"><mml:mo>≥</mml:mo></mml:math></inline-formula>7 cm) and small branches (<inline-formula><mml:math display="inline"><mml:mo>&lt;</mml:mo></mml:math></inline-formula> 7 cm) is based on
Kaufmann (2001), and that of needles/leaves on Perruchoud et al. (1999). Roots
are estimated with equations from Wirth et al. (2004) for coniferous trees
and Wutzler et al. (2008) for deciduous trees. Estimates for branches,
foliage and roots were derived from the DBH only, while for stem wood over
bark including stock, the diameter at 7 m tree height and the total tree
height were also required (Kaufmann, 2001). For this study, only living trees
were considered. The NFI data were stratified for each biogeographical region
of Switzerland and three elevation strata (Table A2). Open forest is
represented by forest plots permanently reduced in stocking, and closed
forest by all forest plots minus plots permanently and temporarily reduced in
stocking.</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F2"><caption><p>Examples of <bold>(a)</bold> intensively used open land <bold>(b)</bold>
extensively used open land <bold>(c)</bold> open forest <bold>(d)</bold> closed
forest.</p></caption>
          <?xmltex \igopts{width=241.848425pt}?><graphic xlink:href="https://www.biogeosciences.net/12/467/2015/bg-12-467-2015-f02.pdf"/>

        </fig>

      <p>The estimates of soil carbon stocks were taken from previous assessments in
approximately 1000 soil profiles in forests (Hagedorn et al., 2010; Nussbaum
et al., 2012) and 500 soil profiles in open land across Switzerland (Leifeld
et al., 2005; Bolliger et al., 2008), where each soil profile in the forest
had been sampled according to horizons analyzed for their C content using a
C / N analyzer. The bulk density and volumetric stone content measured were
used to calibrate pedotransfer functions. The carbon stock changes in soils
were calculated as the differences between the stocks of different LULC
classes. Since soil carbon stocks in agriculturally used open land and forest
correlate fairly well across different altitudes (Sjörgersten-Turner et
al., 2011), we decided to use the difference between mean soil carbon stocks
in open land and forest for the whole of Switzerland.</p>
      <p>Changes in carbon stocks in ecosystems can be related to changes in
atmospheric CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> concentrations by including the molecular masses of
carbon, dry air and the mass of the atmosphere (Schwaiger and Bird, 2010;
O'Halloran et al., 2012):
            <disp-formula id="Ch1.E3" content-type="numbered"><mml:math display="block"><mml:mrow><mml:mi mathvariant="normal">Δ</mml:mi><mml:msub><mml:mi>C</mml:mi><mml:mi mathvariant="normal">A</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:mstyle displaystyle="true"><mml:mfrac style="display"><mml:mrow><mml:mi mathvariant="normal">Δ</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msub><mml:mi>m</mml:mi><mml:mi mathvariant="normal">C</mml:mi></mml:msub><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msub><mml:mi>M</mml:mi><mml:mi mathvariant="normal">a</mml:mi></mml:msub></mml:mrow><mml:mrow><mml:msub><mml:mi>M</mml:mi><mml:mi mathvariant="normal">C</mml:mi></mml:msub><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msub><mml:mi>m</mml:mi><mml:mi mathvariant="normal">a</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>,</mml:mo></mml:mrow></mml:math></disp-formula>
          where <inline-formula><mml:math display="inline"><mml:mrow><mml:mi mathvariant="normal">Δ</mml:mi><mml:msub><mml:mi>C</mml:mi><mml:mi mathvariant="normal">A</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> is the change in atmospheric CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula>
concentration, <inline-formula><mml:math display="inline"><mml:mrow><mml:mi mathvariant="normal">Δ</mml:mi><mml:msub><mml:mi>m</mml:mi><mml:mi mathvariant="normal">C</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> the difference between carbon stocks of
two LULC classes, <inline-formula><mml:math display="inline"><mml:mrow><mml:msub><mml:mi>M</mml:mi><mml:mi mathvariant="normal">C</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> is the molecular mass of carbon,
<inline-formula><mml:math display="inline"><mml:mrow><mml:msub><mml:mi>m</mml:mi><mml:mi mathvariant="normal">a</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> is the mass of the atmosphere and <inline-formula><mml:math display="inline"><mml:mrow><mml:msub><mml:mi>M</mml:mi><mml:mi mathvariant="normal">a</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> is the
molecular mass of dry air. Before using this CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> value (<inline-formula><mml:math display="inline"><mml:mrow><mml:mi mathvariant="normal">Δ</mml:mi><mml:msub><mml:mi>C</mml:mi><mml:mi mathvariant="normal">A</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>) to calculate <inline-formula><mml:math display="inline"><mml:mi mathvariant="normal">Δ</mml:mi></mml:math></inline-formula>RF (Eq. 2), we took into
consideration the fact that each CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> pulse emitted to the atmosphere
disappears partly in sinks of the global carbon cycle (e.g. oceans). A carbon-pulse response function can be used to describe such fluxes (Forster et al.,
2007):
            <disp-formula id="Ch1.E4" content-type="numbered"><mml:math display="block"><mml:mrow><mml:mi mathvariant="normal">fr</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo><mml:mo>=</mml:mo><mml:msub><mml:mi>a</mml:mi><mml:mn mathvariant="normal">0</mml:mn></mml:msub><mml:mo>+</mml:mo><mml:munderover><mml:mo movablelimits="false">∑</mml:mo><mml:mrow><mml:mi>j</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow><mml:mn mathvariant="normal">3</mml:mn></mml:munderover><mml:msub><mml:mi>a</mml:mi><mml:mi>i</mml:mi></mml:msub><mml:msup><mml:mi>e</mml:mi><mml:mstyle scriptlevel="+1"><mml:mfrac><mml:mrow><mml:mo>-</mml:mo><mml:mi>t</mml:mi></mml:mrow><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mi>j</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle></mml:msup><mml:mo>,</mml:mo></mml:mrow></mml:math></disp-formula>
          where fr(<inline-formula><mml:math display="inline"><mml:mi>t</mml:mi></mml:math></inline-formula>) is the fraction of a CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> pulse, which can still be found
in the atmosphere after time <inline-formula><mml:math display="inline"><mml:mi>t</mml:mi></mml:math></inline-formula>. Generally, the coefficients <inline-formula><mml:math display="inline"><mml:mrow><mml:msub><mml:mi>a</mml:mi><mml:mn mathvariant="normal">0</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> to
<inline-formula><mml:math display="inline"><mml:mrow><mml:msub><mml:mi>a</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> and <inline-formula><mml:math display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mn mathvariant="normal">1</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> to <inline-formula><mml:math display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">τ</mml:mi><mml:mn mathvariant="normal">3</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> have no direct process-based meaning,
but are fitting parameters chosen to represent a given model-based
carbon-pulse response function (Joos et al., 2013). CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> sequestration
equals negative CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> pulses (CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> removal from the atmosphere). The
carbon-pulse response function can be applied to negative pulses because a
reduced CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> concentration in the atmosphere will reduce the amount of
CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> sequestrated by the terrestrial biosphere and oceans. To estimate
how much carbon dioxide from continuous CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> pulses after time <inline-formula><mml:math display="inline"><mml:mi>t</mml:mi></mml:math></inline-formula> stays
airborne, a widely used convolution function can be applied (Siegenthaler and
Oeschger, 1978; Cherubini et al., 2011):
            <disp-formula id="Ch1.E5" content-type="numbered"><mml:math display="block"><mml:mrow><mml:mi>C</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo><mml:mo>=</mml:mo><mml:munderover><mml:mo movablelimits="false">∫</mml:mo><mml:mn mathvariant="normal">0</mml:mn><mml:mi>t</mml:mi></mml:munderover><mml:mi>g</mml:mi><mml:mo>(</mml:mo><mml:msup><mml:mi>t</mml:mi><mml:mo>′</mml:mo></mml:msup><mml:mo>)</mml:mo><mml:mi>y</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>-</mml:mo><mml:msup><mml:mi>t</mml:mi><mml:mo>′</mml:mo></mml:msup><mml:mo>)</mml:mo><mml:mi mathvariant="normal">d</mml:mi><mml:msup><mml:mi>t</mml:mi><mml:mo>′</mml:mo></mml:msup><mml:mo>,</mml:mo></mml:mrow></mml:math></disp-formula>
          where <inline-formula><mml:math display="inline"><mml:mrow><mml:mi>C</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> stands for the yearly change in carbon dioxide in the
atmosphere that can be exclusively related to carbon sequestration during
forest expansion, <inline-formula><mml:math display="inline"><mml:mrow><mml:mi>g</mml:mi><mml:mo>(</mml:mo><mml:msup><mml:mi>t</mml:mi><mml:mo>′</mml:mo></mml:msup><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> characterizes the carbon sequestration due to
forest expansion, depending on the gradient of CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> uptake during
succession and <inline-formula><mml:math display="inline"><mml:mrow><mml:mi>y</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>-</mml:mo><mml:msup><mml:mi>t</mml:mi><mml:mo>′</mml:mo></mml:msup><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> accounts for the reduced uptake of carbon dioxide in
the carbon cycle. We estimate the integral using a simple numerical
approximation and time intervals of 1 year:
            <disp-formula id="Ch1.E6" content-type="numbered"><mml:math display="block"><mml:mrow><mml:msub><mml:mi>C</mml:mi><mml:mi>t</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:munderover><mml:mo movablelimits="false">∑</mml:mo><mml:mrow><mml:mi>i</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow><mml:mi>t</mml:mi></mml:munderover><mml:msub><mml:mi>g</mml:mi><mml:mi>i</mml:mi></mml:msub><mml:msub><mml:mi>y</mml:mi><mml:mrow><mml:mi>t</mml:mi><mml:mo>-</mml:mo><mml:mi>i</mml:mi></mml:mrow></mml:msub><mml:mo>.</mml:mo></mml:mrow></mml:math></disp-formula>
          We kept the background CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> concentration fixed to solve the radiative
transfer parameterization (Eq. 2) and accordingly used parameters that
describe carbon sequestration in the carbon cycle for a fixed CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula>
background concentration. The CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> concentration and parameters for the
carbon cycle will, however, change (IPCC, 2001) and it is not clear whether
it is necessary to take these changes into account. Joos et al. (2013) showed
that radiative forcing was more or less constant when CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> pulses were
emitted to atmospheres with different CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> background concentrations.
They suggest that the carbon uptake per unit atmospheric
CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> decreases not only with a high background concentration of CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula>, but
also with the RF per unit change in atmospheric CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula>. They both decrease
in such a way that RF of a CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> emission is almost identical at
preindustrial and present-day conditions. Keeping background CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula>
concentrations and carbon cycle parameters constant thus seems to be a reasonable
approximation.</p>
</sec>
<sec id="Ch1.S2.SS5">
  <title>Albedo RF</title>
      <p>The yearly radiative forcing <inline-formula><mml:math display="inline"><mml:mi mathvariant="normal">Δ</mml:mi></mml:math></inline-formula>RF(t) at the top of the
atmosphere of an albedo change is calculated (modified from Montenegro et
al., 2009) as the seasonal average:
            <disp-formula id="Ch1.E7" content-type="numbered"><mml:math display="block"><mml:mrow><mml:mi mathvariant="normal">Δ</mml:mi><mml:mi mathvariant="normal">RF</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo><mml:mo>=</mml:mo><mml:mstyle displaystyle="true"><mml:mfrac style="display"><mml:mn mathvariant="normal">1</mml:mn><mml:mn>12</mml:mn></mml:mfrac></mml:mstyle><mml:munderover><mml:mo movablelimits="false">∑</mml:mo><mml:mrow><mml:mi>t</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow><mml:mn>12</mml:mn></mml:munderover><mml:msub><mml:mi>I</mml:mi><mml:mi>t</mml:mi></mml:msub><mml:mspace linebreak="nobreak" width="0.125em"/><mml:mi mathvariant="normal">Δ</mml:mi><mml:msub><mml:mi mathvariant="italic">α</mml:mi><mml:mi>t</mml:mi></mml:msub><mml:mo>-</mml:mo><mml:mn>0.23</mml:mn><mml:mspace width="0.25em" linebreak="nobreak"/><mml:msub><mml:mi>I</mml:mi><mml:mi>t</mml:mi></mml:msub><mml:mspace linebreak="nobreak" width="0.125em"/><mml:mi mathvariant="normal">Δ</mml:mi><mml:msub><mml:mi mathvariant="italic">α</mml:mi><mml:mi>t</mml:mi></mml:msub><mml:mo>,</mml:mo></mml:mrow></mml:math></disp-formula>
          where <inline-formula><mml:math display="inline"><mml:mrow><mml:msub><mml:mi>I</mml:mi><mml:mi>t</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> is the monthly global radiation at the earth surface, <inline-formula><mml:math display="inline"><mml:mrow><mml:mi mathvariant="normal">Δ</mml:mi><mml:msub><mml:mi mathvariant="italic">α</mml:mi><mml:mi>t</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> is the albedo difference between two LULC classes depending on
the monthly fraction of snow cover and <inline-formula><mml:math display="inline"><mml:mn>0.23</mml:mn></mml:math></inline-formula> is a factor to account for the
absorption of the reflected radiation in the atmosphere. The first part of
Eq. (7) describes which part of the global radiation is absorbed at the
surface. The second part describes which part of the reflected short-wave
radiation is absorbed in the atmosphere. The factor of 0.23 is that used by
Montenegro et al. (2009) who modified the value 0.3 given by Weaver et
al. (2001).</p>
      <p>We used gridded global radiation (i.e. surface short-wave irradiance) data
from MeteoSwiss (MeteoSwiss, 2012) in monthly data sets averaged over the
period from 2004 to 2010 to eliminate inter-annual variability. The spatial
resolution of the global radiation data set is 2.2 km. The derivation of the
global radiation data was based on the Heliosat method (Cano et al., 1986;
Beyer et al., 1996; Hammer et al., 2003), applied to Meteosat SEVIRI data. It
was verified using high-quality surface measurements and sensitivity runs for
key input parameters (Durr et al., 2010). The albedo was estimated using the
following equation (modified from Barnes and Roy, 2010; Roesch et al., 2002):
            <disp-formula id="Ch1.E8" content-type="numbered"><mml:math display="block"><mml:mrow><mml:mi mathvariant="normal">Δ</mml:mi><mml:mi mathvariant="italic">α</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo><mml:mo>=</mml:mo><mml:mi>f</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo><mml:mspace width="0.125em" linebreak="nobreak"/><mml:mi mathvariant="normal">Δ</mml:mi><mml:msub><mml:mi mathvariant="italic">α</mml:mi><mml:mi mathvariant="normal">s</mml:mi></mml:msub><mml:mo>+</mml:mo><mml:mo>(</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>-</mml:mo><mml:mi>f</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi><mml:mo>)</mml:mo><mml:mo>)</mml:mo><mml:mspace linebreak="nobreak" width="0.125em"/><mml:mi mathvariant="normal">Δ</mml:mi><mml:msub><mml:mi mathvariant="italic">α</mml:mi><mml:mi>v</mml:mi></mml:msub><mml:mo>,</mml:mo></mml:mrow></mml:math></disp-formula>
          where <inline-formula><mml:math display="inline"><mml:mrow><mml:mi mathvariant="normal">Δ</mml:mi><mml:mi mathvariant="italic">α</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi></mml:mrow></mml:math></inline-formula>) is the monthly albedo difference between two LULC
classes, <inline-formula><mml:math display="inline"><mml:mrow><mml:mi mathvariant="normal">Δ</mml:mi><mml:msub><mml:mi mathvariant="italic">α</mml:mi><mml:mi mathvariant="normal">s</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> the average albedo difference between two
LULC classes when snow covered, <inline-formula><mml:math display="inline"><mml:mrow><mml:mi mathvariant="normal">Δ</mml:mi><mml:msub><mml:mi mathvariant="italic">α</mml:mi><mml:mi>v</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> the average albedo
difference between two LULC classes when snow free and <inline-formula><mml:math display="inline"><mml:mrow><mml:mi>f</mml:mi><mml:mo>(</mml:mo><mml:mi>t</mml:mi></mml:mrow></mml:math></inline-formula>) the fraction of
snow cover per month. We used average albedo differences of snow-free and
snow-covered albedo differences and not monthly differences for two
reasons. First, by far the strongest seasonal trend is related to the
presence of snow, which we explicitly included (Zhou et al., 2003). Second,
in some months reliable albedo data was missing and we considered the average
to be a robust estimate. Since we found that the seasonal variation of the
albedos of different LULC classes is similar, the averaging of snow-covered
and snow-free albedo differences results in a fairly good approximation
(Fig. A2 and A3).</p>
      <p>Monthly data sets on snow cover were provided by the Remote Sensing Research
Group at the University of Bern. Raw 1 km data from AVHRR (Advanced Very
High-Resolution Radiometer) were processed using an algorithm to estimate the
snow cover (Separation of Pixels using Aggregated Rating Over
Canada, SPARC). The algorithm was adapted to the mountainous region of the European
Alps and verified (Huesler et al., 2012). Again we calculated an average
value for each month using data of the years 2002–2009 to eliminate
inter-annual variability.</p>
      <p>For assigning albedos to different LULC classes, we overlaid NASA's
Moderate-resolution Imaging Spectroradiometer (MODIS) 0.5 km bidirectional reflectance distribution function (BRDF)/albedo
product (MCD43A) and gridded 0.1 km LULC data from the Swiss area
statistics aggregated into five categories (see Sect. 2.3). We used MODIS data between 2004 and 2009 and the area
statistics from the third inventory period 2004–2009 to ensure a temporal
overlap between the albedo and the LULC records. Since not all the LULC data
for south-eastern Switzerland are available yet, we complemented the area
statistics (2004–2009) with data from the second inventory period
(1992–1997) and accepted a temporal displacement for this region.</p>
      <p>We applied several different methods for calculating land-use-specific
albedos. First, we retained albedo pixels of the highest quality (full BRDF
inversion) for our analysis. To reduce the error caused by assigning albedo
values to a mixed pixel (several 0.1 km LULC classes in one 0.5 km albedo
pixel), we only assigned albedo values to a specific LULC class if at least
92 % of the albedo pixel were covered by just one LULC class (similar to
Kvalevag et al., 2010). The threshold of 92 % (23 out of 25 pixels) is a
trade-off between using as many albedo pixels in the study area as possible
and at the same time reducing the error due to 8 % random land cover (2
out of 25 pixels). If not enough pixels covered by at least 92 % of one
LULC class were available, we would apply additional methods to calculate
specific albedos, accepting the trade-off of using more pixels, but with less
quality. For the first step, we included albedo values of lower quality
produced with magnitude inversion (Liu et al., 2009). If still not enough
pixels were available, we would change the cell size of the area statistics to
0.5 km (according to the most frequent LULC class within a pixel) and
assign albedo values to this new LULC data set. To estimate closed forest
albedos, we only used pixels of at least 92 % land cover and the best quality
(full BRDF inversion). Open forest values were mainly calculated using lower
quality values (magnitude inversion) and the resampled pixels (Table A1). For
extensively and intensively used open land, we used the average albedo values
for the whole study area because it was not possible to derive specific
values for each region (e.g. extensively used open land hardly occurs
below 600 m a.s.l. and intensively used open land hardly occurs above
1200 m a.s.l.). We accepted a bias of open land albedos in these regions
since they were usually not important for LULC change. For instance, LULC
change below 600 m involves almost exclusively intensively used open land.</p>
      <p>The MCD43A product is available for free online. It provides atmospherically
corrected gridded albedo data for a variety of spectrums. We used broadband
white-sky albedo (0.3–5.0 <inline-formula><mml:math display="inline"><mml:mi mathvariant="normal">µ</mml:mi></mml:math></inline-formula>m). In order to distinguish between
snow-covered and snow-free areas, we applied the quality flags of the MODIS
product MCD43A2. The albedo product MCD43A3 has been produced applying the
MODIS BRDF/albedo algorithm (Strahler et al., 1999). This algorithm makes use
of 16-days worth of multi-date data from both the Terra and Aqua platforms
and a semi-empirical kernel-driven bidirectional reflectance model to
determine a global set of parameters describing the BRDF of the land surface (Schaaf, 2010).</p>
</sec>
<sec id="Ch1.S2.SS6">
  <title>Temporal signature of RF</title>
      <p>Both albedo RF and CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> RF are a function of time. The annual variation
of CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> RF depends on the carbon-pulse response function and the yearly
carbon sequestration in biomass and soil during succession. The annual
variation of albedo RF depends on the albedo change during succession. For
both carbon sequestration and albedo change we had to rely on the static
difference between land-use classes, e.g. open land (as starting point) and
closed forests (end of succession). We assumed that carbon sequestration as
well as albedo change follow linear trends until they reach an approximately
steady state. Since a detailed description of the temporal evolution of
albedo change and carbon sequestration is complex and varies with location,
we used a simplified scheme in which we assumed that albedo change is complete
after 30 years, carbon sequestration in biomass after 50 years and carbon
sequestration in soils after 100 years. This seems to be a reasonable
approximation since albedo change is likely to end before carbon accumulation
in biomass does (Kirschbaum et al., 2011, de Wit et al., 2013) and carbon
sequestration in soils will most likely not end in less than 100 years (Poeplau et
al., 2011). We assumed that albedo change and carbon sequestration stop after
a certain time; however, interactions with the global carbon cycle will still
cause changes in the atmospheric CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> concentration. Thus, we compared
the temporal mean of radiative forcing for two different time horizons, 100
and 1000 years. For the representation of our results, we chose the mean RF
in 100 years. The temporal average of RF is useful not only when representing
spatial variability but also for a comparison with earlier studies on
spatial variability of RF, which did not explicitly include temporal
variation (Betts, 2000, Montenegro et al., 2009).</p>
</sec>
<sec id="Ch1.S2.SS7">
  <title>Sensitivity analysis</title>
      <p>In a sensitivity analysis we tested how the spatial variability and
uncertainty of each input factor influence our results. We based the
sensitivity analysis on FAST (Fourier amplitude sensitivity test), developed
by Saltelli et al. (1999), and used an implementation (fast99) provided in
the R package sensitivity (Pujol et al., 2012). Applying a sensitivity
analysis with FAST allowed us to show how varying input factors influenced
the variance of the output including first-order effects and interactions for
each parameter. We approximated input as either uniform or normal
distributed, according to the distribution of each input factor in the study
area. We assumed input factors were uncorrelated, which only holds to a
certain degree since, for example, all factors either increase or decrease with
elevation. We separated our sensitivity analysis into two parts. First, we
applied data on spatial variability. Each of the factors, carbon
sequestration, snow cover, global radiation, albedo difference (snow covered)
and albedo difference (snow free) vary spatially and temporally. We averaged
every input factor temporally and applied the spatial minimum and maximum of
each factor to the function fast99 (Table A3). The minimum and maximum of
snow-covered albedo differences are, e.g. 0.208 (Alps 600–1200) and 0.375
(Jura <inline-formula><mml:math display="inline"><mml:mo>&gt;</mml:mo></mml:math></inline-formula> 1200). This analysis showed which factor had the greatest influence
on the variation in RF caused by change in forest cover at a specific
location. In a second sensitivity analysis, we applied the uncertainty values
of each factor (Table A3). The sensitivity analysis of uncertainties,
represented by random sampling and measurement errors, indicates which
parameter causes high or low uncertainty in the output.</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F3"><caption><p><bold>(a)</bold> Mean yearly global radiation. <bold>(b)</bold> Days with
snow cover per year. <bold>(c)</bold> Albedo difference without snow (difference
between intensively and extensively used open land and closed forest).
<bold>(d)</bold> Albedo difference with snow (difference between
intensively and extensively used open land and closed forest). <bold>(e)</bold>
Carbon sequestration (difference between intensively and extensively used open land and closed
forest). <bold>(f)</bold> Treeline elevation calculated using the Swiss area
statistics of 1997 and applying the method of Paulsen and Körner (2001).</p></caption>
          <?xmltex \igopts{width=241.848425pt}?><graphic xlink:href="https://www.biogeosciences.net/12/467/2015/bg-12-467-2015-f03.pdf"/>

        </fig>

      <?xmltex \floatpos{t}?><fig id="Ch1.F4"><caption><p><bold>(a)</bold> Albedo RF, <bold>(b)</bold> CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> RF, <bold>(c)</bold> offset:
albedo RF/CO2 RF and <bold>(d)</bold> net RF: albedo RF plus CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> RF. All data sets
were derived for transitions from intensively used open land (<inline-formula><mml:math display="inline"><mml:mrow><mml:mo>&lt;</mml:mo><mml:mn>1000</mml:mn></mml:mrow></mml:math></inline-formula> m) and
extensively used open land (<inline-formula><mml:math display="inline"><mml:mrow><mml:mo>&gt;</mml:mo><mml:mn>1000</mml:mn></mml:mrow></mml:math></inline-formula>) to closed forest.</p></caption>
          <?xmltex \igopts{width=241.848425pt}?><graphic xlink:href="https://www.biogeosciences.net/12/467/2015/bg-12-467-2015-f04.pdf"/>

        </fig>

      <?xmltex \floatpos{t}?><fig id="Ch1.F5"><caption><p>Radiative forcings of the forest expansion between 1985 and 1997 for
three elevations in the biogeographical regions Jura, Plateau, northern
Prealps, Central Alps and southern Prealps.</p></caption>
          <?xmltex \igopts{width=241.848425pt}?><graphic xlink:href="https://www.biogeosciences.net/12/467/2015/bg-12-467-2015-f05.pdf"/>

        </fig>

      <?xmltex \floatpos{t}?><fig id="Ch1.F6" specific-use="star"><caption><p>Global sensitivity analysis FAST: <bold>(a)</bold> sensitivity analysis
of spatial variability of <inline-formula><mml:math display="inline"><mml:mrow><mml:msub><mml:mi>c</mml:mi><mml:mi mathvariant="normal">seq</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> (carbon sequestration),
<inline-formula><mml:math display="inline"><mml:mrow><mml:msub><mml:mi>s</mml:mi><mml:mi mathvariant="normal">c</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> (snow cover), glob (global radiation), <inline-formula><mml:math display="inline"><mml:mrow><mml:msub><mml:mi>a</mml:mi><mml:mi mathvariant="normal">ns</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> (albedo
difference no snow cover) and <inline-formula><mml:math display="inline"><mml:mrow><mml:msub><mml:mi>a</mml:mi><mml:mi mathvariant="normal">s</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> (albedo difference
snow covered), <bold>(b)</bold> sensitivity analysis of uncertainty for each
pixel: <inline-formula><mml:math display="inline"><mml:mrow><mml:msub><mml:mi>c</mml:mi><mml:mi mathvariant="normal">seq</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math display="inline"><mml:mrow><mml:mi>s</mml:mi><mml:mi>c</mml:mi></mml:mrow></mml:math></inline-formula>, glob, <inline-formula><mml:math display="inline"><mml:mrow><mml:msub><mml:mi>a</mml:mi><mml:mi mathvariant="normal">ns</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math display="inline"><mml:mrow><mml:msub><mml:mi>a</mml:mi><mml:mi mathvariant="normal">s</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>,
<inline-formula><mml:math display="inline"><mml:mrow><mml:msub><mml:mi>c</mml:mi><mml:mi mathvariant="normal">cycle</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> (carbon cycle), <inline-formula><mml:math display="inline"><mml:mrow><mml:msub><mml:mi>r</mml:mi><mml:mi mathvariant="normal">t</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> (radiative transfer) and
<inline-formula><mml:math display="inline"><mml:mrow><mml:mi>a</mml:mi><mml:mi>t</mml:mi><mml:msub><mml:mi>m</mml:mi><mml:mi mathvariant="normal">absorb</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> (atmospheric absorption).</p></caption>
          <?xmltex \igopts{width=398.338583pt}?><graphic xlink:href="https://www.biogeosciences.net/12/467/2015/bg-12-467-2015-f06.pdf"/>

        </fig>

<?xmltex \hack{\newpage}?>
</sec>
</sec>
<sec id="Ch1.S3">
  <title>Results</title>
      <p>In the forest, average C stocks in biomass ranged from 95 to 170 tC ha<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:math></inline-formula>
(NFI). In the soil (mineral soil 0–100 cm <inline-formula><mml:math display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula> organic layer) the average
across Switzerland was 143 tc ha<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:math></inline-formula> (Nussbaum et al., 2012). In the
intensively used open land, the average value for biomass was
4.34 tc ha<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:math></inline-formula> (Heldstab et al., 2012) and 91 tc ha<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:math></inline-formula> in the
soils. In comparison, the C stocks of the extensively used open land amounted
to 7 tc ha<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:math></inline-formula> in the biomass and 63 tc ha<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:math></inline-formula> in the soil. Minimum
and maximum carbon sequestration in Switzerland of transitions from
intensively or extensively used open land to closed forest thus ranged from
143 to 241 tc ha<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:math></inline-formula> (Fig. 3e). This corresponded to a CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> RF of
<inline-formula><mml:math display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>16 to <inline-formula><mml:math display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>27 W m<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:msup></mml:math></inline-formula> (Fig. 4b).</p>
      <p>The albedos of snow-covered closed forests ranged from 0.168 to 0.267, while
those of snow-free closed forests ranged from 0.101 to 0.139. Similarly, the
albedos of snow-covered open forests ranged from 0.217 to 0.307 and those of
snow-free open forests from 0.117 to 0.141 (Fig. 5). The average albedo of
intensively used open land was 0.170/0.475 (snow free/snow covered) while
that of extensively used open land was 0.154/0.549 (snow free/snow covered).
The albedo differences between transitions from intensively or extensively
used open land to closed forest thus ranged from 0.208 to 0.375
(snow covered) and from 0.023 to 0.066 (snow free) (Fig. 3c, d). Albedo
change in Switzerland caused albedo RF ranging from 2 to 21 W m<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:msup></mml:math></inline-formula>
(Fig. 4a).</p>
      <p>The net RF at different locations in the study area ranged from <inline-formula><mml:math display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>24 to
2 W m<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:msup></mml:math></inline-formula> and the offset of CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> RF caused by albedo RF differed
between 11 and 109 %. The differences were particularly marked on an
elevational gradient and between the five biogeographical regions. Below an elevation of
600 m a.s.l., the albedo RF offset CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> RF by 15 % on average.
Between 600 and 1200 m, the offset was 22 % and above 1200 m 54 %.
The highest RF was observed in high snow-rich alpine regions. In the Central
Alps, 13 % of all possible LULC areas had an offset of more than
80 %. In the southern Prealps 7 % of all possible LULC areas had an
offset higher than 80 %. In the Central Alps, as well as in the southern
Prealps, we found areas where net RF was positive. However, these areas
amounted to less than 0.2 % of both regions. We only found positive RF at
elevations above 1850 m. The albedo RF was the lowest in the Plateau region and
at low elevations of the Jura. Albedo RF accounted, on average, for less than
14 % of the CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> RF in these regions. The lowest net RF (average
<inline-formula><mml:math display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>20 W m<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:msup></mml:math></inline-formula>) was found in the northern Prealps below 1200 m. Above
1200 m the net RF in the northern Prealps was lower than in southern Prealps
and Central Alps, although the snow cover in the northern Prealps was
persistent. However, the effect of a persistent snow cover in the northern
Prealps was outweighed by the low global radiation, the low treeline and the
high carbon sequestration in this alpine region (see Fig. 3).</p>
      <p>The net RF forcing of forests across the mountainous terrain in Switzerland
strongly depended on the elevation (Fig. 5). The albedo RF increased with
altitude, with several factors contributing to this increase (Fig. 3). First,
global radiation increased with altitude, reaching maxima in continental
parts of the Alps. Second, albedo (snow free, snow covered) of forests was in
general lower above 1200m a.s.l. (where coniferous species dominate). Third,
both snow cover and snow-cover period strongly increased with altitude. The
carbon sequestration potential of forests decreased with altitude (due to
unfavorable climatic conditions towards treeline).</p>
      <p>Radiative forcing depended not only on location but also on the type of LULC
transition. LULC transitions from open land to open forest had a higher
offset (approx. 80 %) than transitions from already established open
forest to closed forest (approx. 40 %). Transitions from open forest to
closed forest were generally associated with relatively high change in carbon
stocks (around 70 tc ha<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:math></inline-formula>), and relatively small albedo change (for
snow-covered albedo <inline-formula><mml:math display="inline"><mml:mo>&lt;</mml:mo></mml:math></inline-formula> 0.06 and for snow-free albedo <inline-formula><mml:math display="inline"><mml:mo>&lt;</mml:mo></mml:math></inline-formula> 0.01).</p>
      <p>Between 1985 and 1997, all five types of LULC transitions (Table A4) took
place on an area of 24 000 ha. More than 70 % of them were situated above
1000 m. Above 1000 m, the most frequent transition was the one from open to
closed forest (50 %), followed by transitions from extensively used open
land to open forest (20 %) and from extensively used open land to closed
forest (13 %). Land-use change occurred especially in high altitude
regions where albedo RF strongly offsets CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> RF (Fig. 5). At
the same time, the most frequent transition was the one from open forest to
closed forest, where albedo RF had the least influence on net RF. In summary,
the CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> RF for all land-use transitions that were part of the forest
expansion and succession between 1985 and 1997 in Switzerland was reduced by
approximately 40 %, if the albedo RF is taken into account.</p>
      <p>The net RF varied greatly on small spatial scales. In our study area, the
parameter that had the most significant influence on the spatial variability
of RF was snow cover, followed by carbon sequestration, difference in
snow-covered albedo, global radiation and difference in snow-free albedo
(Fig. 6a). Thus, if the average snow cover (in days year<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:math></inline-formula>) and carbon
sequestration at a certain location were known, the net RF and the offset of
CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> RF through albedo RF could be estimated well without including all
factors in an explicit calculation. We found that the offset of CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> RF
can only be higher than 50 % if the snow cover lasted over 120 days
year<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:math></inline-formula>. An increase in snow cover caused an exponential increase in
albedo RF. Because, first, persistent snow cover was found in regions with
high global radiation and second because the longer snow covered the surface,
the longer it would be present during the days with high global radiation in
March, April and May.</p>
      <p>The uncertainty of net radiative forcing was mainly attributed to
uncertainties in carbon sequestration, followed by snow-covered albedo
difference, global radiation and snow-free albedo difference. The main
effects of uncertainty related to each input factor were much more relevant
than the contribution of interactions between all input factors (Fig. 6a).</p>
</sec>
<sec id="Ch1.S4">
  <title>Discussion</title>
      <p>Our detailed assessment of radiative forcing across Switzerland shows that
the albedo RF increases with increasing altitude, which offsets the CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> RF
at high elevations with long snow-covered periods, high global radiation and
low carbon sequestration. The altitudinal RF gradient in Switzerland is very
strong in comparison to the latitudinal gradient in boreal regions (Betts,
2000; Montenegro et al., 2009). The persistence of snow cover increases with
increasing elevation and increasing latitude. However, while persistence of
snow cover and global radiation are usually positively correlated in
mountainous regions, causing high RF, they are negatively correlated in
boreal regions. The strong altitudinal RF gradient found in this study is
likely to be even more pronounced if altitudinal changes in forest structure
and its influence on albedo and carbon sequestration are included on higher
resolution because forests with very low carbon stocks can also have a low
albedo (de Wit et al., 2014).</p>
      <p>However, despite the general increase of RF with elevation, each
biogeographical region has its specific characteristics. While the Jura and
Plateau (under 600 m a.s.l.) are characterized, respectively, by albedos of 0.136 and 0.139
(snow free) and 0.272 and 0.276 (snow covered), the albedo in the southern
Prealps under 600 m is comparably low (0.112 and 0.185, respectively). This
regional difference may be related to the different forest types and soil
characteristics. Whereas beech dominates in the Jura and the Plateau, many
stands in the southern Prealps are dominated by chestnut. Moreover, forest
soils in the lowlands of the southern Prealps contain particularly high
fractions of black fire derived carbon (Eckmeier et al., 2010). The
darker soil colour may lower the albedo in addition to differences in the
canopy.</p>
      <p>Our spatially highly resolved estimates of RF are in agreement with the
results of Betts (2000) and Montenegro et al. (2009) which are based on much
coarser resolutions. In Betts (2000), the two pixels encompassing Switzerland
show a net carbon sequestration of 100–150 and 150–200 tc ha<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:math></inline-formula>
associated with reforestation of pasture, while in Montenegro et al. (2009),
the pixels indicate a drawdown ranging between 100–150 tc ha<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:math></inline-formula>
(maximum scenario) and 0–20 tc ha<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:math></inline-formula> (minimum scenario). However, many
pixels are missing in alpine regions. In our study, we found similar values
for the drawdown with values ranging from <inline-formula><mml:math display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>10 to 160 tc ha<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:math></inline-formula> (minus 10
indicates emission instead of drawdown).</p>
      <p>Small-scale variability, especially in topographically complex areas, was not
captured in these former studies. Here, we used data on global radiation and
snow cover on a resolution of 2.2 km (MeteoSwiss) and 1 km (Huesler et al.,
2012). To calculate differences in carbon sequestration and albedo, we relied
on a biogeographical categorization and altitudinal stratification, which are
based on major differences in vegetation and other ecological factors
(Gutersohn, 1973; Wohlgemuth, 1996). Although the spatial resolution was high
in our study, it should be refined further to allow, for instance, a comparison
of RF on northern and southern slopes and a better capture of forest types
and structures near the treeline.</p>
      <p>The global sensitivity analysis with FAST showed that interactions between
the input parameters were small in comparison to the main effects of each
parameter. Thus, we verified the results of our global sensitivity analysis
in a local analysis by estimating the partial derivates for each factor. Both
analyses showed good agreement since not only the interactions but also
non-linear effects had little influence. The amount of RF at a specific
location is essentially influenced by carbon sequestration and snow cover
(Fig. 6a). These two factors are good indicators for estimating the amount of
RF. Global radiation and snow-covered albedo are also important, but they
influence RF five times less than carbon sequestration and even seven times
less than snow cover. The factors with the most potential for improving our
results are better estimates of carbon sequestration (Fig. 6b), followed by
reducing albedo uncertainty and the global radiation. The average parameter
for atmospheric absorption (0.23) could be replaced by a spatially explicit
parameter. Including a spatially explicit parameter for atmospheric
absorption would probably increase the elevation gradient of RF, because
atmospheric absorption should be higher in low elevations than in high
elevations. According to our sensitivity analysis and Bright and
Kvalevag (2013) improving data on atmospheric absorption will have a
relatively small influence on the results.</p>
      <p>The sensitivity analysis with FAST is based on uncertainty estimates because
exact values are not available. For example, MODIS values can be assigned to
an average uncertainty of 10 % (O'Halloran et al., 2012, Strahler et al.,
1999). This uncertainty is likely to be much higher in topographically
complex areas than in even terrain as the algorithm used to produce albedo
values only indirectly accounts for topography. Moreover, the uncertainties
associated with the different input parameters do not always refer to the
same statistical measures and are thus not completely consistent (references
of uncertainties are listed in the Table A3).</p>
      <p>Regarding the different time horizons, RF increased by 17 % for a time
horizon of 1000 years compared with a <?xmltex \hack{\mbox\bgroup}?>horizon<?xmltex \hack{\egroup}?> of 100 years. Larger time
horizons increased RF since CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> RF constantly decreased due to
interactions of atmospheric CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> with the carbon cycle, while albedo RF
became constant after forests reached a steady state. Since we used a time
horizon of 100 years, we rather underestimated albedo RF. This apparently
goes against the findings of Schaeffer et al. (2006) and Kirschbaum et
al. (2011) who both argue that CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> RF becomes more dominant for larger
time horizons. However, they consider relatively short time periods
(including rotations) where carbon sequestration does not end before the
forests are removed. We think it is also necessary to include large time
horizons when estimating RF, but it is of course an oversimplification to
assume a fixed state after transition, since forests are frequently disturbed
(e.g. O'Halloran et al., 2012). Over longer periods snow cover will be less
persistent in the Alps, which will potentially decrease the albedo RF (de Wit
et al., 2013; Pitman et al., 2011).</p>
      <p>The maps of the possible carbon sequestration and albedo of forests in each
biogeographical region (Fig. 3) reflect mainly ecosystem characteristics.
However, forests in Switzerland have long been under permanent anthropogenic
influence and are thus not in a naturally balanced equilibrium, i.e. at the
end of succession. For example, forests at high elevations in the Jura are
often used for pasturing. Hence, they are less dense and sequester less
carbon than they would if left to develop as undisturbed forests. The
relationship between albedo RF, CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> RF and net RF will thus always
depend on the actual and previous forest and land management. Our analysis of
transitions from extensively used open land to closed forest and open forest
corroborates this conclusion. The results indicated that the climatic benefit
will be smaller if forests are kept in an open structure (e.g. due to
pasturing) than when the canopy closes during succession. These findings are
in line with former studies that estimate the effects of succession and
forest structure on RF and show that changes in carbon stocks and changes in
albedo are not linearly related (Kirschbaum et al., 2011; Bernier et al.,
2011).</p>
      <p>Our results should be valuable for future studies on the climatic impacts of
LULUCF, especially for comparing and evaluating the results from climate
models since our results are mainly based on satellite and field data. They
should also be valuable for assessing the ecosystem (climate) service of
forests in temperate mountains, i.e. for addressing the question of whether
forest expansion in these regions is beneficial for climate or not. However,
the interpretation of RF values has to be done carefully. First, the concept
of radiative forcing has been developed to compare the impact of different
forcing agents on the global mean temperature (Hansen et al., 2005). When
applied at the regional and local scales one should keep in mind that the
comparison of different forcing agents is far from being straightforward. For
instance, the impact of albedo will remain mostly local while those from
CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> will be globally distributed and therefore diluted. Furthermore, the
climate sensitivities of CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> RF and albedo RF may differ (Davin et al.,
2007).</p>
      <p>Finally, it is important to note that our study does not account for all
possible effects of forests on climate such as changes in
evapotranspiration, surface roughness, and impacts on aerosols and other
gases than CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula>. The uptake of CH<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">4</mml:mn></mml:msub></mml:math></inline-formula> during forest expansion in the
Alps increased according to Hiltbrunner et al. (2012), which adds to a
negative CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> RF. This effect is, however, one magnitude smaller than the
effect of CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> sequestration. The effect of a changing aerosol
concentration, providing cooling (through cloud formation), may be very
important (Spracklen et al., 2008). Changes in evapotranspiration and surface
roughness, due to forest expansion, have a cooling impact in many
geographical contexts (Bonan, 2008; Luyssaert et al., 2014), but we did not
include the impacts of these changes in our study. In general, the influence
of evapotranspiration and surface roughness will be low in those areas where
snow cover plays an important role (Bonan, 2008; Bathiany et al., 2010;
Gibbard et al., 2005; Lee et al., 2011). Hence, these effects will be more
important at low elevations (e.g. on the Swiss Plateau) than at high
elevations in the Alps. The gap between the benefits of forest expansion at
low and high elevations may thus become even wider if evapotranspiration and
surface roughness are also integrated.</p>
</sec>
<sec id="Ch1.S5" sec-type="conclusions">
  <title>Conclusions</title>
      <p>In the temperate mountainous regions of Switzerland, the net RF associated
with changes in forest cover varies greatly on small spatial scales. At low
elevations, with low to moderate snow cover, RF is strongly negative due to
a dominance of CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> RF. At high elevations in continental regions with
persistent snow cover, a very high global radiation, low carbon
sequestration and low albedos of mostly evergreen tree species, RF can be
positive. As a consequence, both clearly negative and positive values of RF
can be found within a horizontal distance of 5 km in alpine valleys.
Therefore, the climatic benefits of changes in forest cover can only be
properly assessed using data at a high spatial resolution.</p>
      <p>Our results indicate that it is very important to include albedo RF when
estimating the impact on climate of changes in forest cover. Maps of RF,
such as those produced in this study, indicate where climatic benefits from
changes in forest cover can be expected and where not. In the Swiss Alps,
the relevance of albedo RF is especially high because most transitions from
open land to forest occur in regions where albedo RF causes a strong offset
of CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> RF. Practitioners and politicians who need information about
ecosystem services on local and regional scales should take into account
that RF in the Swiss Alps mainly depends on the persistence of snow cover
and the potential for carbon sequestration. Moreover, late successional
forest-cover changes from open forests to closed forests are more beneficial
for climate than early successional changes.</p>
      <p>Our results could be improved if changes in evapotranspiration, surface
roughness, aerosols and gases other than CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> were included. To
determine the impacts of RF better, however, further advances in climate
modelling are necessary. A promising approach could thus be the coupling of
regional climate models with global climate models. Regional models are able
to simulate all the effects of changes in land use on climate (including
evapotranspiration, surface roughness and so on) on a small scale. Coupling
regional models with global models allows the integration of feedbacks with
the global circulation. This could help to close the gap between RF and
temperature changes, and thus answer the question about where temperature
changes caused by RF can be expected and how much change is likely.</p><?xmltex \hack{\clearpage}?>
</sec>

      
      </body>
    <back><app-group><app id="App1.Ch1.S1">
  <title/>

<?xmltex \floatpos{h!}?><table-wrap id="App1.Ch1.T1" position="anchor"><?xmltex \hack{\hsize\textwidth}?><caption><p>Albedo of forest (snow, no snow) and open
forest (snow, no snow) and corresponding standard errors (SE) of each
biogeographical region. Values marked with <inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula> were derived using majority
pixels and/or magnitude inversion (see chapter albedo RF), all other values
have been derived using 92 % pixel cover and full inversion. Numbers 1–3 behind the biogeographical regions
indicate the elevation (1 is below 600 m, 2 is 600–1200 m, 3 is above
1200 m).</p></caption><oasis:table frame="bottom"><oasis:tgroup cols="9">
     <oasis:colspec colnum="1" colname="col1" align="left"/>
     <oasis:colspec colnum="2" colname="col2" align="right"/>
     <oasis:colspec colnum="3" colname="col3" align="right"/>
     <oasis:colspec colnum="4" colname="col4" align="right"/>
     <oasis:colspec colnum="5" colname="col5" align="right"/>
     <oasis:colspec colnum="6" colname="col6" align="right"/>
     <oasis:colspec colnum="7" colname="col7" align="right"/>
     <oasis:colspec colnum="8" colname="col8" align="right"/>
     <oasis:colspec colnum="9" colname="col9" align="right"/>
     <oasis:thead>
       <oasis:row rowsep="1">  
         <oasis:entry colname="col1">Region</oasis:entry>  
         <oasis:entry namest="col2" nameend="col5" align="center">Forest </oasis:entry>  
         <oasis:entry namest="col6" nameend="col9" align="center">Open forest </oasis:entry>
       </oasis:row>
     </oasis:thead>
     <oasis:tbody>
       <oasis:row rowsep="1">  
         <oasis:entry colname="col1"/>  
         <oasis:entry colname="col2">Snow</oasis:entry>  
         <oasis:entry colname="col3">SE(%)</oasis:entry>  
         <oasis:entry colname="col4">Not Snow</oasis:entry>  
         <oasis:entry colname="col5">SE(%)</oasis:entry>  
         <oasis:entry colname="col6">Snow</oasis:entry>  
         <oasis:entry colname="col7">SE(%)</oasis:entry>  
         <oasis:entry colname="col8">Not Snow</oasis:entry>  
         <oasis:entry colname="col9">SE(%)</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Jura 1</oasis:entry>  
         <oasis:entry colname="col2">0.263</oasis:entry>  
         <oasis:entry colname="col3">1.30</oasis:entry>  
         <oasis:entry colname="col4">0.139</oasis:entry>  
         <oasis:entry colname="col5">0.08</oasis:entry>  
         <oasis:entry colname="col6">NA</oasis:entry>  
         <oasis:entry colname="col7">NA</oasis:entry>  
         <oasis:entry colname="col8">NA</oasis:entry>  
         <oasis:entry colname="col9">NA</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Jura 2</oasis:entry>  
         <oasis:entry colname="col2">0.221</oasis:entry>  
         <oasis:entry colname="col3">0.50</oasis:entry>  
         <oasis:entry colname="col4">0.133</oasis:entry>  
         <oasis:entry colname="col5">0.04</oasis:entry>  
         <oasis:entry colname="col6">0.303<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>  
         <oasis:entry colname="col7">1.80<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>  
         <oasis:entry colname="col8">0.141<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>  
         <oasis:entry colname="col9">0.23<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Jura 3</oasis:entry>  
         <oasis:entry colname="col2">0.175</oasis:entry>  
         <oasis:entry colname="col3">0.64</oasis:entry>  
         <oasis:entry colname="col4">0.109</oasis:entry>  
         <oasis:entry colname="col5">0.12</oasis:entry>  
         <oasis:entry colname="col6">0.232<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>  
         <oasis:entry colname="col7">0.67<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>  
         <oasis:entry colname="col8">0.117<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>  
         <oasis:entry colname="col9">0.13<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Plateau 1</oasis:entry>  
         <oasis:entry colname="col2">0.266</oasis:entry>  
         <oasis:entry colname="col3">0.61</oasis:entry>  
         <oasis:entry colname="col4">0.136</oasis:entry>  
         <oasis:entry colname="col5">0.03</oasis:entry>  
         <oasis:entry colname="col6">NA</oasis:entry>  
         <oasis:entry colname="col7">NA</oasis:entry>  
         <oasis:entry colname="col8">NA</oasis:entry>  
         <oasis:entry colname="col9">NA</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Plateau 2</oasis:entry>  
         <oasis:entry colname="col2">0.251</oasis:entry>  
         <oasis:entry colname="col3">0.53</oasis:entry>  
         <oasis:entry colname="col4">0.134</oasis:entry>  
         <oasis:entry colname="col5">0.05</oasis:entry>  
         <oasis:entry colname="col6">NA</oasis:entry>  
         <oasis:entry colname="col7">NA</oasis:entry>  
         <oasis:entry colname="col8">NA</oasis:entry>  
         <oasis:entry colname="col9">NA</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Plateau 3</oasis:entry>  
         <oasis:entry colname="col2">NA</oasis:entry>  
         <oasis:entry colname="col3">NA</oasis:entry>  
         <oasis:entry colname="col4">0</oasis:entry>  
         <oasis:entry colname="col5">NA</oasis:entry>  
         <oasis:entry colname="col6">NA</oasis:entry>  
         <oasis:entry colname="col7">NA</oasis:entry>  
         <oasis:entry colname="col8">NA</oasis:entry>  
         <oasis:entry colname="col9">NA</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Northern Prealps 1</oasis:entry>  
         <oasis:entry colname="col2">0.240</oasis:entry>  
         <oasis:entry colname="col3">3.16</oasis:entry>  
         <oasis:entry colname="col4">0.127</oasis:entry>  
         <oasis:entry colname="col5">0.45</oasis:entry>  
         <oasis:entry colname="col6">NA</oasis:entry>  
         <oasis:entry colname="col7">NA</oasis:entry>  
         <oasis:entry colname="col8">NA</oasis:entry>  
         <oasis:entry colname="col9">NA</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Northern Prealps 2</oasis:entry>  
         <oasis:entry colname="col2">0.194</oasis:entry>  
         <oasis:entry colname="col3">0.38</oasis:entry>  
         <oasis:entry colname="col4">0.118</oasis:entry>  
         <oasis:entry colname="col5">0.06</oasis:entry>  
         <oasis:entry colname="col6">0.217<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>  
         <oasis:entry colname="col7">2.36<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>  
         <oasis:entry colname="col8">0.114<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>  
         <oasis:entry colname="col9">0.49<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Northern Prealps 3</oasis:entry>  
         <oasis:entry colname="col2">0.203</oasis:entry>  
         <oasis:entry colname="col3">0.40</oasis:entry>  
         <oasis:entry colname="col4">0.112</oasis:entry>  
         <oasis:entry colname="col5">0.11</oasis:entry>  
         <oasis:entry colname="col6">0.275<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>  
         <oasis:entry colname="col7">0.42<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>  
         <oasis:entry colname="col8">0.121</oasis:entry>  
         <oasis:entry colname="col9">0.095</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Central Alps 1</oasis:entry>  
         <oasis:entry colname="col2">0.168<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>  
         <oasis:entry colname="col3">2.1<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>  
         <oasis:entry colname="col4">0.107</oasis:entry>  
         <oasis:entry colname="col5">1.31</oasis:entry>  
         <oasis:entry colname="col6">0.353<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>  
         <oasis:entry colname="col7">15.7<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>  
         <oasis:entry colname="col8">0.141<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>  
         <oasis:entry colname="col9">0.58<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Central Alps 2</oasis:entry>  
         <oasis:entry colname="col2">0.174</oasis:entry>  
         <oasis:entry colname="col3">0.61</oasis:entry>  
         <oasis:entry colname="col4">0.104</oasis:entry>  
         <oasis:entry colname="col5">0.15</oasis:entry>  
         <oasis:entry colname="col6">0.285<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>  
         <oasis:entry colname="col7">3.09<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>  
         <oasis:entry colname="col8">0.134<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>  
         <oasis:entry colname="col9">0.18<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Central Alps 3</oasis:entry>  
         <oasis:entry colname="col2">0.192</oasis:entry>  
         <oasis:entry colname="col3">0.23</oasis:entry>  
         <oasis:entry colname="col4">0.101</oasis:entry>  
         <oasis:entry colname="col5">0.08</oasis:entry>  
         <oasis:entry colname="col6">0.274<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>  
         <oasis:entry colname="col7">0.33<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>  
         <oasis:entry colname="col8">0.118<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>  
         <oasis:entry colname="col9">0.068<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Southern Prealps 1</oasis:entry>  
         <oasis:entry colname="col2">0.185<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>  
         <oasis:entry colname="col3">4.9<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>  
         <oasis:entry colname="col4">0.111</oasis:entry>  
         <oasis:entry colname="col5">0.23</oasis:entry>  
         <oasis:entry colname="col6">NA</oasis:entry>  
         <oasis:entry colname="col7">NA</oasis:entry>  
         <oasis:entry colname="col8">0.127<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>  
         <oasis:entry colname="col9">0.78<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Southern Prealps 2</oasis:entry>  
         <oasis:entry colname="col2">0.190</oasis:entry>  
         <oasis:entry colname="col3">3.13</oasis:entry>  
         <oasis:entry colname="col4">0.121</oasis:entry>  
         <oasis:entry colname="col5">0.07</oasis:entry>  
         <oasis:entry colname="col6">0.258<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>  
         <oasis:entry colname="col7">3.28<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>  
         <oasis:entry colname="col8">0.126<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>  
         <oasis:entry colname="col9">0.27<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Southern Prealps 3</oasis:entry>  
         <oasis:entry colname="col2">0.194</oasis:entry>  
         <oasis:entry colname="col3">0.78</oasis:entry>  
         <oasis:entry colname="col4">0.107</oasis:entry>  
         <oasis:entry colname="col5">0.12</oasis:entry>  
         <oasis:entry colname="col6">0.307<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>  
         <oasis:entry colname="col7">0.50<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>  
         <oasis:entry colname="col8">0.119</oasis:entry>  
         <oasis:entry colname="col9">0.089</oasis:entry>
       </oasis:row>
     </oasis:tbody>
   </oasis:tgroup></oasis:table></table-wrap>

<?xmltex \floatpos{h!}?><table-wrap id="App1.Ch1.T2" position="anchor"><?xmltex \hack{\hsize\textwidth}?><caption><p>Carbon stocks of closed forests and open
forests in biomass and soils. The standard errors (SEs) refer to the
deviation from the sample mean in every biogeographical region. Not included
are errors of the model parameters which have been used to derive carbon
stocks from tree measurements.</p></caption><oasis:table frame="topbot"><oasis:tgroup cols="5">
     <oasis:colspec colnum="1" colname="col1" align="left"/>
     <oasis:colspec colnum="2" colname="col2" align="right"/>
     <oasis:colspec colnum="3" colname="col3" align="right"/>
     <oasis:colspec colnum="4" colname="col4" align="right"/>
     <oasis:colspec colnum="5" colname="col5" align="right"/>
     <oasis:thead>
       <oasis:row rowsep="1">  
         <oasis:entry colname="col1">Region</oasis:entry>  
         <oasis:entry namest="col2" nameend="col3" align="center">Forest </oasis:entry>  
         <oasis:entry namest="col4" nameend="col5" align="center">Open forest </oasis:entry>
       </oasis:row>
     </oasis:thead>
     <oasis:tbody>
       <oasis:row>  
         <oasis:entry colname="col1"/>  
         <oasis:entry colname="col2">biomass</oasis:entry>  
         <oasis:entry colname="col3">SE</oasis:entry>  
         <oasis:entry colname="col4">biomass</oasis:entry>  
         <oasis:entry colname="col5">SE</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">  
         <oasis:entry colname="col1"/>  
         <oasis:entry colname="col2">(tC ha<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:math></inline-formula>)</oasis:entry>  
         <oasis:entry colname="col3">(%)</oasis:entry>  
         <oasis:entry colname="col4">(tC ha<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:math></inline-formula>)</oasis:entry>  
         <oasis:entry colname="col5">(%)</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Jura 1</oasis:entry>  
         <oasis:entry colname="col2">134.1</oasis:entry>  
         <oasis:entry colname="col3">4.6</oasis:entry>  
         <oasis:entry colname="col4">NA</oasis:entry>  
         <oasis:entry colname="col5">NA</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Jura 2</oasis:entry>  
         <oasis:entry colname="col2">144.3</oasis:entry>  
         <oasis:entry colname="col3">2.3</oasis:entry>  
         <oasis:entry colname="col4">101.27</oasis:entry>  
         <oasis:entry colname="col5">27.2</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Jura 3</oasis:entry>  
         <oasis:entry colname="col2">103.7</oasis:entry>  
         <oasis:entry colname="col3">5.5</oasis:entry>  
         <oasis:entry colname="col4">68.44</oasis:entry>  
         <oasis:entry colname="col5">13.4</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Plateau 1</oasis:entry>  
         <oasis:entry colname="col2">140.4</oasis:entry>  
         <oasis:entry colname="col3">2.7</oasis:entry>  
         <oasis:entry colname="col4">21.08</oasis:entry>  
         <oasis:entry colname="col5">51.0</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Plateau 2</oasis:entry>  
         <oasis:entry colname="col2">159.7</oasis:entry>  
         <oasis:entry colname="col3">3.2</oasis:entry>  
         <oasis:entry colname="col4">92.80</oasis:entry>  
         <oasis:entry colname="col5">46.2</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Plateau 3</oasis:entry>  
         <oasis:entry colname="col2">137.58</oasis:entry>  
         <oasis:entry colname="col3">14.1</oasis:entry>  
         <oasis:entry colname="col4">71.26</oasis:entry>  
         <oasis:entry colname="col5">33.6</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Northern Prealps 1</oasis:entry>  
         <oasis:entry colname="col2">169.8</oasis:entry>  
         <oasis:entry colname="col3">9.5</oasis:entry>  
         <oasis:entry colname="col4">NA</oasis:entry>  
         <oasis:entry colname="col5">NA</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Northern Prealps 2</oasis:entry>  
         <oasis:entry colname="col2">166.8</oasis:entry>  
         <oasis:entry colname="col3">2.7</oasis:entry>  
         <oasis:entry colname="col4">85.60</oasis:entry>  
         <oasis:entry colname="col5">22.7</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Northern Prealps 3</oasis:entry>  
         <oasis:entry colname="col2">152.5</oasis:entry>  
         <oasis:entry colname="col3">4.5</oasis:entry>  
         <oasis:entry colname="col4">72.21</oasis:entry>  
         <oasis:entry colname="col5">11.0</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Central Alps 1</oasis:entry>  
         <oasis:entry colname="col2">122</oasis:entry>  
         <oasis:entry colname="col3">11.2</oasis:entry>  
         <oasis:entry colname="col4">NA</oasis:entry>  
         <oasis:entry colname="col5">NA</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Central Alps 2</oasis:entry>  
         <oasis:entry colname="col2">124.7</oasis:entry>  
         <oasis:entry colname="col3">4.0</oasis:entry>  
         <oasis:entry colname="col4">70.41</oasis:entry>  
         <oasis:entry colname="col5">29.8</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Central Alps 3</oasis:entry>  
         <oasis:entry colname="col2">115.5</oasis:entry>  
         <oasis:entry colname="col3">2.3</oasis:entry>  
         <oasis:entry colname="col4">57.90</oasis:entry>  
         <oasis:entry colname="col5">7.9</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Southern Prealps 1</oasis:entry>  
         <oasis:entry colname="col2">95.2</oasis:entry>  
         <oasis:entry colname="col3">6.8</oasis:entry>  
         <oasis:entry colname="col4">NA</oasis:entry>  
         <oasis:entry colname="col5">NA</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Southern Prealps 2</oasis:entry>  
         <oasis:entry colname="col2">94.7</oasis:entry>  
         <oasis:entry colname="col3">4.7</oasis:entry>  
         <oasis:entry colname="col4">37.40</oasis:entry>  
         <oasis:entry colname="col5">39.3</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Southern Prealps 3</oasis:entry>  
         <oasis:entry colname="col2">100.6</oasis:entry>  
         <oasis:entry colname="col3">4.5</oasis:entry>  
         <oasis:entry colname="col4">41.60</oasis:entry>  
         <oasis:entry colname="col5">11.6</oasis:entry>
       </oasis:row>
     </oasis:tbody>
   </oasis:tgroup></oasis:table></table-wrap>

<?xmltex \hack{\clearpage}?><?xmltex \floatpos{h!}?><table-wrap id="App1.Ch1.T3" position="anchor"><?xmltex \hack{\hsize\textwidth}?><caption><p>Spatial variability and uncertainties
applied in the sensitivity analysis.</p></caption><oasis:table frame="topbot"><oasis:tgroup cols="3">
     <oasis:colspec colnum="1" colname="col1" align="left"/>
     <oasis:colspec colnum="2" colname="col2" align="left"/>
     <oasis:colspec colnum="3" colname="col3" align="justify" colwidth="85.358268pt"/>
     <oasis:thead>
       <oasis:row rowsep="1">  
         <oasis:entry colname="col1"/>  
         <oasis:entry colname="col2">Spatial variability</oasis:entry>  
         <oasis:entry colname="col3">Uncertainty <?xmltex \hack{\hfill\break}?>(standard deviation)</oasis:entry>
       </oasis:row>
     </oasis:thead>
     <oasis:tbody>
       <oasis:row>  
         <oasis:entry colname="col1">Carbon sequestration [tC ha<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:math></inline-formula>]</oasis:entry>  
         <oasis:entry colname="col2">143–241</oasis:entry>  
         <oasis:entry colname="col3"><inline-formula><mml:math display="inline"><mml:mrow><mml:mo>+</mml:mo><mml:mo>/</mml:mo><mml:mo>-</mml:mo><mml:mn>35</mml:mn><mml:mi mathvariant="italic">%</mml:mi></mml:mrow></mml:math></inline-formula> <?xmltex \hack{\hfill\break}?>(Hagedorn et al., 2010; Heldstab et al., 2012)</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Snow cover [days year<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:math></inline-formula>]</oasis:entry>  
         <oasis:entry colname="col2">10–240</oasis:entry>  
         <oasis:entry colname="col3"><inline-formula><mml:math display="inline"><mml:mrow><mml:mo>+</mml:mo><mml:mo>/</mml:mo><mml:mo>-</mml:mo><mml:mn>10</mml:mn><mml:mi mathvariant="italic">%</mml:mi></mml:mrow></mml:math></inline-formula> <?xmltex \hack{\hfill\break}?>(Husler et al., 2012)</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Global radiation [W m<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:msup></mml:math></inline-formula>]</oasis:entry>  
         <oasis:entry colname="col2">117–180</oasis:entry>  
         <oasis:entry colname="col3"><inline-formula><mml:math display="inline"><mml:mrow><mml:mo>+</mml:mo><mml:mo>/</mml:mo><mml:mo>-</mml:mo><mml:mn>28</mml:mn><mml:mi mathvariant="italic">%</mml:mi></mml:mrow></mml:math></inline-formula> <?xmltex \hack{\hfill\break}?>(Durr et al., 2010)</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Albedo snow covered</oasis:entry>  
         <oasis:entry colname="col2">0.208–0.375</oasis:entry>  
         <oasis:entry colname="col3"><inline-formula><mml:math display="inline"><mml:mrow><mml:mo>+</mml:mo><mml:mo>/</mml:mo><mml:mo>-</mml:mo><mml:mn>36</mml:mn><mml:mi mathvariant="italic">%</mml:mi></mml:mrow></mml:math></inline-formula> <?xmltex \hack{\hfill\break}?>(Combined error of MODIS data and methods<?xmltex \hack{\hfill\break}?>for assigning LULC specific albedos)</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Albedo snow free</oasis:entry>  
         <oasis:entry colname="col2">0.025–0.066</oasis:entry>  
         <oasis:entry colname="col3"><inline-formula><mml:math display="inline"><mml:mrow><mml:mo>+</mml:mo><mml:mo>/</mml:mo><mml:mo>-</mml:mo><mml:mn>28</mml:mn><mml:mi mathvariant="italic">%</mml:mi></mml:mrow></mml:math></inline-formula> <?xmltex \hack{\hfill\break}?>(Combined error of MODIS data and methods<?xmltex \hack{\hfill\break}?>for assigning LULC specific albedos)</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Carbon cycle</oasis:entry>  
         <oasis:entry colname="col2">–</oasis:entry>  
         <oasis:entry colname="col3"><inline-formula><mml:math display="inline"><mml:mrow><mml:mo>+</mml:mo><mml:mo>/</mml:mo><mml:mo>-</mml:mo><mml:mn>15</mml:mn><mml:mi mathvariant="italic">%</mml:mi></mml:mrow></mml:math></inline-formula> <?xmltex \hack{\hfill\break}?>(Joos et al., 2013)</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Radiative transfer</oasis:entry>  
         <oasis:entry colname="col2">–</oasis:entry>  
         <oasis:entry colname="col3"><inline-formula><mml:math display="inline"><mml:mrow><mml:mo>+</mml:mo><mml:mo>/</mml:mo><mml:mo>-</mml:mo><mml:mn>10</mml:mn><mml:mi mathvariant="italic">%</mml:mi></mml:mrow></mml:math></inline-formula> <?xmltex \hack{\hfill\break}?>(Myhre et al., 1998)</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Atmospheric absorption</oasis:entry>  
         <oasis:entry colname="col2">–</oasis:entry>  
         <oasis:entry colname="col3"><inline-formula><mml:math display="inline"><mml:mrow><mml:mo>+</mml:mo><mml:mo>/</mml:mo><mml:mo>-</mml:mo><mml:mn>30</mml:mn><mml:mi mathvariant="italic">%</mml:mi></mml:mrow></mml:math></inline-formula> <?xmltex \hack{\hfill\break}?>radiative transfer tests<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula></oasis:entry>
       </oasis:row>
     </oasis:tbody>
   </oasis:tgroup></oasis:table><table-wrap-foot><p><inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>*</mml:mo></mml:msup></mml:math></inline-formula> For the uncertainty in atmospheric absorption, we relied on
experiments with the Fu and Liou online model (Fu and Liou, 2005). We tested
different scenarios of cloudiness, aerosol concentration and elevation to
determine how much atmospheric absorption could vary over Switzerland.</p></table-wrap-foot></table-wrap>

<?xmltex \hack{\clearpage}?><?xmltex \floatpos{h!}?><table-wrap id="App1.Ch1.T4" position="anchor"><?xmltex \hack{\hsize\textwidth}?><caption><p>Area affected by each type of transition
between 1985 and 1997. Numbers 1–3 behind the biogeographical regions
indicate the elevation (1 is below 600 m, 2 is 600–1200 m, 3 is above
1200 m).</p></caption><oasis:table frame="topbot"><oasis:tgroup cols="7">
     <oasis:colspec colnum="1" colname="col1" align="left"/>
     <oasis:colspec colnum="2" colname="col2" align="right"/>
     <oasis:colspec colnum="3" colname="col3" align="right"/>
     <oasis:colspec colnum="4" colname="col4" align="right"/>
     <oasis:colspec colnum="5" colname="col5" align="right"/>
     <oasis:colspec colnum="6" colname="col6" align="right"/>
     <oasis:colspec colnum="7" colname="col7" align="right"/>
     <oasis:thead>
       <oasis:row rowsep="1">  
         <oasis:entry colname="col1">Region</oasis:entry>  
         <oasis:entry namest="col2" nameend="col7" align="center">Area affected by each transition between 1985 and 1997 [in ha] </oasis:entry>
       </oasis:row>
     </oasis:thead>
     <oasis:tbody>
       <oasis:row>  
         <oasis:entry colname="col1">Biogeo-</oasis:entry>  
         <oasis:entry colname="col2">Intensively</oasis:entry>  
         <oasis:entry colname="col3">Extensively</oasis:entry>  
         <oasis:entry colname="col4">Intensively</oasis:entry>  
         <oasis:entry colname="col5">Extensively</oasis:entry>  
         <oasis:entry colname="col6">Open</oasis:entry>  
         <oasis:entry colname="col7">Forest</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">graphical</oasis:entry>  
         <oasis:entry colname="col2">used open</oasis:entry>  
         <oasis:entry colname="col3">used open</oasis:entry>  
         <oasis:entry colname="col4">used open</oasis:entry>  
         <oasis:entry colname="col5">used open</oasis:entry>  
         <oasis:entry colname="col6">forest</oasis:entry>  
         <oasis:entry colname="col7">expansion</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">region</oasis:entry>  
         <oasis:entry colname="col2">land to</oasis:entry>  
         <oasis:entry colname="col3">land to</oasis:entry>  
         <oasis:entry colname="col4">land to</oasis:entry>  
         <oasis:entry colname="col5">land to</oasis:entry>  
         <oasis:entry colname="col6">to closed</oasis:entry>  
         <oasis:entry colname="col7">(sum of all</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">  
         <oasis:entry colname="col1"/>  
         <oasis:entry colname="col2">closed forest</oasis:entry>  
         <oasis:entry colname="col3">closed forest</oasis:entry>  
         <oasis:entry colname="col4">open forest</oasis:entry>  
         <oasis:entry colname="col5">open forest</oasis:entry>  
         <oasis:entry colname="col6">forest</oasis:entry>  
         <oasis:entry colname="col7">transitions)</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Jura 1</oasis:entry>  
         <oasis:entry colname="col2">116</oasis:entry>  
         <oasis:entry colname="col3">31</oasis:entry>  
         <oasis:entry colname="col4">98</oasis:entry>  
         <oasis:entry colname="col5">35</oasis:entry>  
         <oasis:entry colname="col6">106</oasis:entry>  
         <oasis:entry colname="col7">386</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Jura 2</oasis:entry>  
         <oasis:entry colname="col2">113</oasis:entry>  
         <oasis:entry colname="col3">238</oasis:entry>  
         <oasis:entry colname="col4">73</oasis:entry>  
         <oasis:entry colname="col5">330</oasis:entry>  
         <oasis:entry colname="col6">522</oasis:entry>  
         <oasis:entry colname="col7">1276</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Jura 3</oasis:entry>  
         <oasis:entry colname="col2">1</oasis:entry>  
         <oasis:entry colname="col3">46</oasis:entry>  
         <oasis:entry colname="col4">1</oasis:entry>  
         <oasis:entry colname="col5">155</oasis:entry>  
         <oasis:entry colname="col6">490</oasis:entry>  
         <oasis:entry colname="col7">693</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Plateau 1</oasis:entry>  
         <oasis:entry colname="col2">613</oasis:entry>  
         <oasis:entry colname="col3">87</oasis:entry>  
         <oasis:entry colname="col4">379</oasis:entry>  
         <oasis:entry colname="col5">52</oasis:entry>  
         <oasis:entry colname="col6">264</oasis:entry>  
         <oasis:entry colname="col7">1395</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Plateau 2</oasis:entry>  
         <oasis:entry colname="col2">232</oasis:entry>  
         <oasis:entry colname="col3">60</oasis:entry>  
         <oasis:entry colname="col4">110</oasis:entry>  
         <oasis:entry colname="col5">44</oasis:entry>  
         <oasis:entry colname="col6">85</oasis:entry>  
         <oasis:entry colname="col7">531</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Plateau 3</oasis:entry>  
         <oasis:entry colname="col2">NA</oasis:entry>  
         <oasis:entry colname="col3">NA</oasis:entry>  
         <oasis:entry colname="col4">NA</oasis:entry>  
         <oasis:entry colname="col5">NA</oasis:entry>  
         <oasis:entry colname="col6">1</oasis:entry>  
         <oasis:entry colname="col7">1</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Northern Prealps 1</oasis:entry>  
         <oasis:entry colname="col2">109</oasis:entry>  
         <oasis:entry colname="col3">21</oasis:entry>  
         <oasis:entry colname="col4">78</oasis:entry>  
         <oasis:entry colname="col5">13</oasis:entry>  
         <oasis:entry colname="col6">53</oasis:entry>  
         <oasis:entry colname="col7">274</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Northern Prealps 2</oasis:entry>  
         <oasis:entry colname="col2">321</oasis:entry>  
         <oasis:entry colname="col3">497</oasis:entry>  
         <oasis:entry colname="col4">295</oasis:entry>  
         <oasis:entry colname="col5">401</oasis:entry>  
         <oasis:entry colname="col6">959</oasis:entry>  
         <oasis:entry colname="col7">2473</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Northern Prealps 3</oasis:entry>  
         <oasis:entry colname="col2">34</oasis:entry>  
         <oasis:entry colname="col3">955</oasis:entry>  
         <oasis:entry colname="col4">77</oasis:entry>  
         <oasis:entry colname="col5">1180</oasis:entry>  
         <oasis:entry colname="col6">2476</oasis:entry>  
         <oasis:entry colname="col7">4722</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Central Alps 1</oasis:entry>  
         <oasis:entry colname="col2">4</oasis:entry>  
         <oasis:entry colname="col3">4</oasis:entry>  
         <oasis:entry colname="col4">6</oasis:entry>  
         <oasis:entry colname="col5">11</oasis:entry>  
         <oasis:entry colname="col6">29</oasis:entry>  
         <oasis:entry colname="col7">54</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Central Alps 2</oasis:entry>  
         <oasis:entry colname="col2">93</oasis:entry>  
         <oasis:entry colname="col3">101</oasis:entry>  
         <oasis:entry colname="col4">267</oasis:entry>  
         <oasis:entry colname="col5">154</oasis:entry>  
         <oasis:entry colname="col6">679</oasis:entry>  
         <oasis:entry colname="col7">1294</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Central Alps 3</oasis:entry>  
         <oasis:entry colname="col2">102</oasis:entry>  
         <oasis:entry colname="col3">739</oasis:entry>  
         <oasis:entry colname="col4">291</oasis:entry>  
         <oasis:entry colname="col5">1700</oasis:entry>  
         <oasis:entry colname="col6">3687</oasis:entry>  
         <oasis:entry colname="col7">6519</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Southern Prealps 1</oasis:entry>  
         <oasis:entry colname="col2">61</oasis:entry>  
         <oasis:entry colname="col3">25</oasis:entry>  
         <oasis:entry colname="col4">145</oasis:entry>  
         <oasis:entry colname="col5">22</oasis:entry>  
         <oasis:entry colname="col6">196</oasis:entry>  
         <oasis:entry colname="col7">449</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Southern Prealps 2</oasis:entry>  
         <oasis:entry colname="col2">76</oasis:entry>  
         <oasis:entry colname="col3">77</oasis:entry>  
         <oasis:entry colname="col4">170</oasis:entry>  
         <oasis:entry colname="col5">135</oasis:entry>  
         <oasis:entry colname="col6">731</oasis:entry>  
         <oasis:entry colname="col7">1189</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Southern Prealps 3</oasis:entry>  
         <oasis:entry colname="col2">23</oasis:entry>  
         <oasis:entry colname="col3">274</oasis:entry>  
         <oasis:entry colname="col4">71</oasis:entry>  
         <oasis:entry colname="col5">604</oasis:entry>  
         <oasis:entry colname="col6">1541</oasis:entry>  
         <oasis:entry colname="col7">2513</oasis:entry>
       </oasis:row>
     </oasis:tbody>
   </oasis:tgroup></oasis:table></table-wrap>

<?xmltex \hack{\clearpage}?><?xmltex \floatpos{h!}?><table-wrap id="App1.Ch1.T5" position="anchor"><?xmltex \hack{\hsize\textwidth}?><caption><p>Aggregation of land-use classes from Swiss
Arealstatistik (ASCH85, ASCH97 and ASCH04) adapted from Rutherford et
al. (2008).</p></caption><oasis:table frame="topbot"><oasis:tgroup cols="5">
     <oasis:colspec colnum="1" colname="col1" align="justify" colwidth="56.905512pt"/>
     <oasis:colspec colnum="2" colname="col2" align="left"/>
     <oasis:colspec colnum="3" colname="col3" align="justify" colwidth="113.811024pt"/>
     <oasis:colspec colnum="4" colname="col4" align="justify" colwidth="56.905512pt"/>
     <oasis:colspec colnum="5" colname="col5" align="justify" colwidth="113.811024pt"/>
     <oasis:thead>
       <oasis:row rowsep="1">  
         <oasis:entry colname="col1">Aggregated class</oasis:entry>  
         <oasis:entry colname="col2">Area <?xmltex \hack{\hfill\break}?>[ha]</oasis:entry>  
         <oasis:entry colname="col3">Classes from Swiss land-use statistics</oasis:entry>  
         <oasis:entry colname="col4">Area <?xmltex \hack{\hfill\break}?>[ha]</oasis:entry>  
         <oasis:entry colname="col5">Broad definition</oasis:entry>
       </oasis:row>
     </oasis:thead>
     <oasis:tbody>
       <oasis:row rowsep="1">  
         <oasis:entry colname="col1">Closed<?xmltex \hack{\hfill\break}?>forest</oasis:entry>  
         <oasis:entry colname="col2">1 121 544 <?xmltex \hack{\hfill\break}?></oasis:entry>  
         <oasis:entry colname="col3">Afforestation, 52 <?xmltex \hack{\hfill\break}?>Forest dieback, 54 <?xmltex \hack{\hfill\break}?>Normal forest, 50 <?xmltex \hack{\hfill\break}?>Slender forest, 14 <?xmltex \hack{\hfill\break}?>Bushes, 57 <?xmltex \hack{\hfill\break}?>Groves and hedges, 58</oasis:entry>  
         <oasis:entry colname="col4">3349 <?xmltex \hack{\hfill\break}?>14 851 <?xmltex \hack{\hfill\break}?>962 312 <?xmltex \hack{\hfill\break}?>44 711 <?xmltex \hack{\hfill\break}?>60 514 <?xmltex \hack{\hfill\break}?>35 807</oasis:entry>  
         <oasis:entry colname="col5">Vegetation height <inline-formula><mml:math display="inline"><mml:mo>&gt;</mml:mo></mml:math></inline-formula> 3 m, cover density &gt;60 %, composed of tree species</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">  
         <oasis:entry colname="col1">Open<?xmltex \hack{\hfill\break}?>forest</oasis:entry>  
         <oasis:entry colname="col2">150101</oasis:entry>  
         <oasis:entry colname="col3">On non-agriculturally used land, 56 <?xmltex \hack{\hfill\break}?>On agriculturally used land, 55 <?xmltex \hack{\hfill\break}?>Groups of trees on agriculturally used land, 59 <?xmltex \hack{\hfill\break}?>Groups of trees on non-agriculturally used land, 60</oasis:entry>  
         <oasis:entry colname="col4">52 825 <?xmltex \hack{\hfill\break}?>24 108 <?xmltex \hack{\hfill\break}?>38 157 <?xmltex \hack{\hfill\break}?>35 011</oasis:entry>  
         <oasis:entry colname="col5">Vegetation height &gt;3m, cover density 20–60 %, composed of tree species</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">  
         <oasis:entry colname="col1">Extensively<?xmltex \hack{\hfill\break}?>used open land</oasis:entry>  
         <oasis:entry colname="col2">767842</oasis:entry>  
         <oasis:entry colname="col3">Pasture in the vicinity of settlements, 43 <?xmltex \hack{\hfill\break}?>Alpine meadows, 45 <?xmltex \hack{\hfill\break}?>Sheep alps, 49 <?xmltex \hack{\hfill\break}?>Favourable to pasturing, 46 <?xmltex \hack{\hfill\break}?>Stony alpine pasture, 48 <?xmltex \hack{\hfill\break}?>Grass and herb vegetation, 65</oasis:entry>  
         <oasis:entry colname="col4">87 303 <?xmltex \hack{\hfill\break}?>32 316 <?xmltex \hack{\hfill\break}?>51 124 <?xmltex \hack{\hfill\break}?>368 691 <?xmltex \hack{\hfill\break}?>46 024 <?xmltex \hack{\hfill\break}?>182 384</oasis:entry>  
         <oasis:entry colname="col5">Used for grazing, use not year-round, not machine accessible</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">  
         <oasis:entry colname="col1">Intensively<?xmltex \hack{\hfill\break}?>used open land</oasis:entry>  
         <oasis:entry colname="col2">837128</oasis:entry>  
         <oasis:entry colname="col3">Arable land, 41 <?xmltex \hack{\hfill\break}?>Natural meadows, 42</oasis:entry>  
         <oasis:entry colname="col4">547 754 <?xmltex \hack{\hfill\break}?>289 374</oasis:entry>  
         <oasis:entry colname="col5">Year-round use, in the vicinity of settlements, Mown</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Other</oasis:entry>  
         <oasis:entry colname="col2">125186</oasis:entry>  
         <oasis:entry colname="col3">1–40, 44, 47, 61–64, 66–72</oasis:entry>  
         <oasis:entry colname="col4">125 186</oasis:entry>  
         <oasis:entry colname="col5"/>
       </oasis:row>
     </oasis:tbody>
   </oasis:tgroup></oasis:table><table-wrap-foot><p>Numbers in column 3 represent the official ASCH classes of the
nomenclature 2004 (Humbel et al., 2010). The aggregation in Rutherford et
al. (2008) was adapted to the new nomenclature.</p></table-wrap-foot></table-wrap>

<?xmltex \hack{\clearpage}?><?xmltex \floatpos{h!}?><fig id="App1.Ch1.F1" position="anchor"><caption><p>Spatial pattern of forest expansion. The pattern illustrates the
density of forest expansion in Switzerland. The density was calculated
including the area of all five transitions we used for calculating RF (see
Sect. 2.3) and a
kernel-density function in ArcGis 10.1 (ESRI).</p></caption>
        <?xmltex \hack{\hsize\textwidth}?>
        <?xmltex \igopts{width=341.433071pt}?><graphic xlink:href="https://www.biogeosciences.net/12/467/2015/bg-12-467-2015-f07.png"/>

      </fig>

<?xmltex \hack{\clearpage}?><?xmltex \floatpos{h!}?><fig id="App1.Ch1.F2" position="anchor"><caption><p>Seasonal variation of albedo values of the four snow-free LULC
classes: closed forest, intensively used open land, extensively used open land
and open forest (only full BRDF inversion albedo values).</p></caption>
        <?xmltex \hack{\hsize\textwidth}?>
        <?xmltex \igopts{width=341.433071pt}?><graphic xlink:href="https://www.biogeosciences.net/12/467/2015/bg-12-467-2015-f08.png"/>

      </fig>

<?xmltex \hack{\clearpage}?><?xmltex \floatpos{h!}?><fig id="App1.Ch1.F3" position="anchor"><caption><p>Seasonal variation of the albedo values of the four snow-covered
LULC classes: closed forest, intensively used open land, extensively used open
land and open forest (full BRDF albedo values and magnitude inversion albedo
values for open forest).</p></caption>
        <?xmltex \hack{\hsize\textwidth}?>
        <?xmltex \igopts{width=341.433071pt}?><graphic xlink:href="https://www.biogeosciences.net/12/467/2015/bg-12-467-2015-f09.png"/>

      </fig>

<?xmltex \hack{\clearpage}?>
</app>
  </app-group><ack><title>Acknowledgements</title><p>This work was part of the Swiss research programme “Forests and Climate
change” and supported by the Swiss Federal Office for Environment (BAFU) and
the Federal Institute for Forest, Snow and Landscape Research (WSL). We acknowledge the financial support by the
NRP 68 programme of the Swiss National Science foundation and are grateful to
Fortunat Joos, Stefan Wunderle, Janine Bolliger, Tobias Jonas,
Henning Löwe and Michaela Teich for helpful discussions and
Silvia Dingwall for language revision.
<?xmltex \hack{\newline}?><?xmltex \hack{\newline}?>
Edited by: C. A. Williams</p></ack><ref-list>
    <title>References</title>

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