<?xml version="1.0" encoding="UTF-8"?>
<!DOCTYPE article PUBLIC "-//NLM//DTD Journal Publishing with OASIS Tables v3.0 20080202//EN" "journalpub-oasis3.dtd">
<article xmlns:xlink="http://www.w3.org/1999/xlink" xmlns:mml="http://www.w3.org/1998/Math/MathML" xmlns:oasis="http://docs.oasis-open.org/ns/oasis-exchange/table" dtd-version="3.0"><?xmltex \makeatother\@nolinetrue\makeatletter?>
  <front>
    <journal-meta>
<journal-id journal-id-type="publisher">BG</journal-id>
<journal-title-group>
<journal-title>Biogeosciences</journal-title>
<abbrev-journal-title abbrev-type="publisher">BG</abbrev-journal-title>
<abbrev-journal-title abbrev-type="nlm-ta">Biogeosciences</abbrev-journal-title>
</journal-title-group>
<issn pub-type="epub">1726-4189</issn>
<publisher><publisher-name>Copernicus Publications</publisher-name>
<publisher-loc>Göttingen, Germany</publisher-loc>
</publisher>
</journal-meta>

    <article-meta>
      <article-id pub-id-type="doi">10.5194/bg-13-3225-2016</article-id><title-group><article-title>Reconstructions of biomass burning from sediment-charcoal <?xmltex \hack{\newline}?>records to improve
data–model comparisons</article-title>
      </title-group><?xmltex \runningtitle{Reconstructions of biomass burning}?><?xmltex \runningauthor{J. R. Marlon et al.}?>
      <contrib-group>
        <contrib contrib-type="author" corresp="yes" rid="aff1">
          <name><surname>Marlon</surname><given-names>Jennifer R.</given-names></name>
          <email>jennmarlon@gmail.com</email>
        <ext-link>https://orcid.org/0000-0002-8299-9609</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff2">
          <name><surname>Kelly</surname><given-names>Ryan</given-names></name>
          
        </contrib>
        <contrib contrib-type="author" corresp="no" rid="aff3">
          <name><surname>Daniau</surname><given-names>Anne-Laure</given-names></name>
          
        </contrib>
        <contrib contrib-type="author" corresp="no" rid="aff4">
          <name><surname>Vannière</surname><given-names>Boris</given-names></name>
          
        </contrib>
        <contrib contrib-type="author" corresp="no" rid="aff5">
          <name><surname>Power</surname><given-names>Mitchell J.</given-names></name>
          
        </contrib>
        <contrib contrib-type="author" corresp="no" rid="aff6">
          <name><surname>Bartlein</surname><given-names>Patrick</given-names></name>
          
        </contrib>
        <contrib contrib-type="author" corresp="no" rid="aff7">
          <name><surname>Higuera</surname><given-names>Philip</given-names></name>
          
        </contrib>
        <contrib contrib-type="author" corresp="no" rid="aff8">
          <name><surname>Blarquez</surname><given-names>Olivier</given-names></name>
          
        </contrib>
        <contrib contrib-type="author" corresp="no" rid="aff5">
          <name><surname>Brewer</surname><given-names>Simon</given-names></name>
          
        <ext-link>https://orcid.org/0000-0002-6810-1911</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff9">
          <name><surname>Brücher</surname><given-names>Tim</given-names></name>
          
        <ext-link>https://orcid.org/0000-0003-3345-3252</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff10 aff11">
          <name><surname>Feurdean</surname><given-names>Angelica</given-names></name>
          
        <ext-link>https://orcid.org/0000-0002-2497-3005</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff12">
          <name><surname>Romera</surname><given-names>Graciela Gil</given-names></name>
          
        <ext-link>https://orcid.org/0000-0001-5726-2536</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff4">
          <name><surname>Iglesias</surname><given-names>Virginia</given-names></name>
          
        </contrib>
        <contrib contrib-type="author" corresp="no" rid="aff13">
          <name><surname>Maezumi</surname><given-names>S. Yoshi</given-names></name>
          
        </contrib>
        <contrib contrib-type="author" corresp="no" rid="aff14">
          <name><surname>Magi</surname><given-names>Brian</given-names></name>
          
        <ext-link>https://orcid.org/0000-0001-8131-0083</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff15">
          <name><surname>Courtney Mustaphi</surname><given-names>Colin J.</given-names></name>
          
        </contrib>
        <contrib contrib-type="author" corresp="no" rid="aff16">
          <name><surname>Zhihai</surname><given-names>Tonishtan</given-names></name>
          
        </contrib>
        <aff id="aff1"><label>1</label><institution>School of Forestry and Environmental Studies, Yale University, New Haven, USA</institution>
        </aff>
        <aff id="aff2"><label>2</label><institution>Neptune and Company, Inc., Lakewood, CO, USA</institution>
        </aff>
        <aff id="aff3"><label>3</label><institution>Centre National de la Recherche Scientifique (CNRS), Environnements et Paléoenvironnements Océaniques et Continentaux (EPOC),
Unité Mixte de Recherche (UMR) 5805, Université de Bordeaux, 33615 Pessac cedex, France</institution>
        </aff>
        <aff id="aff4"><label>4</label><institution>Chrono-environnement UMR6249 and MSHE USR3124, CNRS, Univ. Bourgogne
Franche-Comté, <?xmltex \hack{\newline}?>25000 Besançon, France</institution>
        </aff>
        <aff id="aff5"><label>5</label><institution>Department of Geography, Natural History Museum of Utah, University of Utah, Salt Lake City, USA</institution>
        </aff>
        <aff id="aff6"><label>6</label><institution>Department of Geography, University of Oregon, Eugene, USA</institution>
        </aff>
        <aff id="aff7"><label>7</label><institution>College of Forestry and Conservation, University of Montana, Missoula, USA</institution>
        </aff>
        <aff id="aff8"><label>8</label><institution>Département de Géographie, Université de Montréal, Montréal, Québec</institution>
        </aff>
        <aff id="aff9"><label>9</label><institution>GEOMAR Helmholtz Centre for Ocean Research Kiel, Düsternbrooker Weg
20, Kiel, Germany</institution>
        </aff>
        <aff id="aff10"><label>10</label><institution>Senckenberg Biodiversity and Climate Research Centre (BiK-F), Frankfurt
am Main, Germany</institution>
        </aff>
        <aff id="aff11"><label>11</label><institution>Department of Geology, Babeş-Bolyai University,
Cluj-Napoca, Romania</institution>
        </aff>
        <aff id="aff12"><label>12</label><institution>IPE-CSIC, Zaragoza, Spain</institution>
        </aff>
        <aff id="aff13"><label>13</label><institution>Department of Archaeology, University of Exeter, Exeter, UK</institution>
        </aff>
        <aff id="aff14"><label>14</label><institution>Department of Geography and Earth Sciences, University of North Carolina at Charlotte, Charlotte, USA</institution>
        </aff>
        <aff id="aff15"><label>15</label><institution>Environment Department, University of York, York, UK</institution>
        </aff>
        <aff id="aff16"><label>16</label><institution>State Key Laboratory of Loess and Quaternary Geology, Key Laboratory of Aerosol Chemistry and Physics,
Institute of Earth Environment, Chinese Academy of Sciences, Xi'an, Shaanxi 710075, PR China</institution>
        </aff>
      </contrib-group>
      <author-notes><corresp id="corr1">Jennifer R. Marlon (jennmarlon@gmail.com)</corresp></author-notes><pub-date><day>3</day><month>June</month><year>2016</year></pub-date>
      
      <volume>13</volume>
      <issue>11</issue>
      <fpage>3225</fpage><lpage>3244</lpage>
      <history>
        <date date-type="received"><day>25</day><month>September</month><year>2015</year></date>
           <date date-type="rev-request"><day>18</day><month>November</month><year>2015</year></date>
           <date date-type="rev-recd"><day>17</day><month>April</month><year>2016</year></date>
           <date date-type="accepted"><day>2</day><month>May</month><year>2016</year></date>
      </history>
      <permissions>
<license license-type="open-access">
<license-p>This work is licensed under a Creative Commons Attribution 3.0 Unported License. To view a copy of this license, visit <ext-link ext-link-type="uri" xlink:href="http://creativecommons.org/licenses/by/3.0/">http://creativecommons.org/licenses/by/3.0/</ext-link></license-p>
</license>
</permissions><self-uri xlink:href="https://bg.copernicus.org/articles/13/3225/2016/bg-13-3225-2016.html">This article is available from https://bg.copernicus.org/articles/13/3225/2016/bg-13-3225-2016.html</self-uri>
<self-uri xlink:href="https://bg.copernicus.org/articles/13/3225/2016/bg-13-3225-2016.pdf">The full text article is available as a PDF file from https://bg.copernicus.org/articles/13/3225/2016/bg-13-3225-2016.pdf</self-uri>


      <abstract>
    <p>The location, timing, spatial extent, and frequency of wildfires are
changing rapidly in many parts of the world, producing substantial impacts
on ecosystems, people, and potentially climate. Paleofire records based on
charcoal accumulation in sediments enable modern changes in biomass burning
to be considered in their long-term context. Paleofire records also provide
insights into the causes and impacts of past wildfires and emissions when
analyzed in conjunction with other paleoenvironmental data and with fire
models. Here we present new 1000-year and 22 000-year trends and gridded
biomass burning reconstructions based on the Global Charcoal Database
version 3 (GCDv3), which includes 736 charcoal records (57 more than in
version 2). The new gridded reconstructions reveal the spatial patterns
underlying the temporal trends in the data, allowing insights into likely
controls on biomass burning at regional to global scales. In the most recent
few decades, biomass burning has sharply increased in both hemispheres but
especially in the north, where charcoal fluxes are now higher than at any
other time during the past 22 000 years. We also discuss methodological
issues relevant to data–model comparisons and identify areas for future
research. Spatially gridded versions of the global data set from GCDv3 are
provided to facilitate comparison with and validation of global fire
simulations.</p>
  </abstract>
    </article-meta>
  </front>
<body>
      

<sec id="Ch1.S1" sec-type="intro">
  <title>Introduction</title>
      <p>Fire has long been recognized as an important ecological process because of
its influence on species distributions and role in shaping other key
ecosystem properties (Bond and Keeley, 2005). Fire also affects
regional and global biogeochemical and hydrologic cycles
(Shakesby and Doerr, 2006; van der Werf et al., 2006),
geophysical processes (Morris and Moses, 1987; DeBano, 2000), and
the climate system (Randerson et al., 2006; Ward et al., 2012; Saleh et al.,
2014).
Nevertheless, large gaps remain in our understanding of the interactions
between fire and climate, despite an increasing need to manage fire and its
emissions (Keywood et al., 2013).</p>
      <p>Fire activity has been characterized at a wide range of spatial and temporal
scales using field observations and historical data
(e.g., Mouillot and Field, 2005; Gavin et al.,
2007), dendrochronological data (e.g., Falk et al., 2011),
satellites (e.g., Mouillot et al., 2014), ice cores
(e.g., McConnell et al., 2007), and charcoal deposits
in sediments, peat bogs, swamps, soils, and other environments
(e.g., Whitlock and Bartlein, 2004). Sedimentary records are
unique among these data sources because of the broad temporal and spatial
coverage they provide, which includes reconstructions of fire history at
local to global spatial scales and decadal to millennial temporal scales
(e.g., Carcaillet et al., 2002; Brown, 2005; Marlon et al., 2008; Iglesias
and Whitlock, 2014).</p>
      <p>Results from paleofire research have helped lay a foundation for
understanding the linkages among fire, climate, vegetation change, and human
activities across a broad range of temporal and spatial scales. Fire-history
data from sediment records highlight the importance of fire as a force of
long-term global environmental change. Syntheses of data in the Global
Charcoal Database (GCD), for example, reveal important variations in biomass
burning during the last glacial period (Daniau et al., 2010), the
last 21 000 years (Power et al., 2008; Daniau et al., 2012), and the last 2000 years (Marlon et al., 2008). With
the increasing number of sites in the GCD, regional syntheses became
possible, including long-term analyses of climate and human influences on
burning in Australasia (Mooney et al., 2011; Williams et al., 2015), the
Mediterranean (Colombaroli et al., 2009; Vanniere et al., 2011), the
western USA (Marlon et al., 2012), and the Americas more
broadly (Whitlock et al., 2007; Power et al., 2012).</p>
      <p>Here we briefly review the history of biomass burning reconstructions based
on charcoal data, and we introduce version 3 of the GCD (GCDv3, <inline-formula><mml:math display="inline"><mml:mrow><mml:mi>n</mml:mi><mml:mo>=</mml:mo></mml:mrow></mml:math></inline-formula> 736), which
improves on GCDv1 (Power et al., 2008) and GCDv2 (Daniau et al., 2012) by
adding 57 records. We also present the GCDv3 in a new globally gridded
format along with several broad-scale syntheses created using the
open-source paleofire R package (Blarquez et al., 2014). The new
gridded maps illustrate the spatial and temporal variability in fire
activity over the past 22 000 years, highlighting recent departures from the
long-term trends. The maps should be useful for modelers as well as others
in the Earth sciences, particularly given the wide-ranging impacts of fire.
Finally, we review several important limitations to charcoal-based records
and identify promising future directions for the field.</p>
</sec>
<sec id="Ch1.S2">
  <title>Reconstructing fire history with sediment-charcoal data</title>
      <p>Fire-history research based on sediment-charcoal data has advanced rapidly
in recent decades. Early analyses of sedimentary charcoal were typically
conducted to support studies focused primarily on reconstructing past
vegetation changes (Heusser, 1995; Fuller et al., 1998; Haberle, 1998;
Behling, 2001). A few early studies focused more directly on fire (Swain,
1973; Burney, 1987; Delcourt et al., 1998). In many cases, microscopic (&lt; 100 <inline-formula><mml:math display="inline"><mml:mi mathvariant="normal">µ</mml:mi></mml:math></inline-formula>m) charcoal
particles were tallied alongside pollen grains. Pollen and charcoal
particles were converted to concentrations using the abundance of exotic
markers of a known quantity added to each sample, and charcoal data were
presented as ratios of the relative abundance of charcoal to pollen. Records
were usually sampled at low temporal resolution due to the intensive labor
and time required to analyze pollen. Samples represented broad spatial areas
because microscopic charcoal can travel hundreds of kilometers (Clark,
1988; Conedera and Tinner, 2010). Variations in both pollen and charcoal
abundances can influence the ratios, however, and so changes in pollen
productivity could produce apparent changes in fire activity when none
occurred. The differential production of charcoal from grass versus wood
species could also alter charcoal / pollen ratios. Thus, early reconstructions
based on microscopic charcoal-to-pollen ratios provided new and often useful
insights, but the information was relatively coarse and potentially
unreliable for inferring past regional fire activity.</p>
      <p>Currently, most paleofire researchers analyze macroscopic charcoal particles
(&gt; 100 <inline-formula><mml:math display="inline"><mml:mi mathvariant="normal">µ</mml:mi></mml:math></inline-formula>m) sampled contiguously from sediment cores to
produce fire-history records that are more spatially and temporally precise
(e.g., “local” histories at decadal timescales) compared to earlier
methods. Macroscopic charcoal is typically quantified by simple particle
counts or area measurements made using image analysis (Carcaillet et al., 2001b). However, particles can also be characterized using morphotypes.
Two primary particle forms for non-arboreal charcoal exist: (1) cellular
“graminoid” (thin rectangular pieces; one cell layer thick with pores and
visible vessels and cell wall separations) and (2) fibrous (collections or
bundles of this filamentous charcoal clumped together). Arboreal charcoal
can be characterized by three morphotypes: (1) dark (opaque, thick, solid,
geometric in shape, some luster, and straight edges), (2) lattice
(cross-hatched forming rectangular ladder-like structure with spaces
between), and (3) branched (dendroidal, generally cylindrical with
successively smaller jutting arms; Jensen et al., 2007;
Tweiten et al., 2009).</p>
      <p>The analysis of high-resolution macroscopic charcoal records focuses on
decomposing temporal variations in particle measurements into low- and high-frequency signals. The low-frequency signals were originally termed
“background” (Clark and Patterson, 1997) and were thought
to primarily reflect non-fire processes within and around a site unrelated
to fire occurrence, largely due to sediment redeposition. The background
component was therefore explicitly filtered out and disregarded in analyses.
Subsequent research, however, demonstrated that background charcoal contains
important information about the relative amount of biomass burned through
time (Haberle and Ledru, 2001; Carcaillet et al., 2002), particularly when
combined across multiple records (Clark and Royall, 1996; Carcaillet et al.,
2002; Marlon et al., 2006). Thus, the collection of many records into a
single repository, including those of insufficient resolution for local fire-history reconstructions, became an important prerequisite for reconstructing
variations in biomass burning at regional to global spatial scales.</p>
      <p>Recent studies demonstrate that background charcoal corresponds well with
independent evidence of area and/or biomass burned both at landscape scales
(Higuera et al., 2011; Kelly et al., 2013) and regionally
(Marlon et al., 2012). However, it is not possible to
quantify absolute area burned in the absence of a calibration data set, and
the influences of non-fire-related processes such as erosion or vegetation
change on biomass burning reconstructions remain poorly understood
(Aleman et al., 2013). These limitations highlight a need for
more calibration studies to understand how charcoal production and taphonomy
relates to the area and amount of biomass burned across a range of
vegetation types and climate conditions.</p>
      <p>Another recent advance in fire research is the reconstruction of fire
frequency based on peaks in sedimentary charcoal records. Fire frequency is
an important component of the fire regime, but such analyses require
data sets with decadal resolution that are relatively uncommon in the GCD. In
order to reconstruct fire frequency, records must be sampled contiguously,
have high temporal resolution relative to the expected mean fire return
intervals, and have sufficient particle counts in each sample to separate
peaks from “background” (Higuera et al., 2007, 2010). In addition, relatively stable sediment accumulation rates are
ideal because peak frequencies will vary with changes in sedimentation rates
(Carcaillet et al., 2001a; Higuera et al., 2010). For these
reasons, our analyses of the GCDv3, which are focused on broad-scale changes
in fire, are limited to the reconstruction of fire activity or biomass
burning rather than to changes in fire frequencies.</p>
      <p>Many other methodological approaches to long-term fire-history
reconstruction are developing from a variety of combustion products in ice
cores (Kehrwald et al., 2013), including the analysis of ammonium
(NH<inline-formula><mml:math display="inline"><mml:mrow><mml:msubsup><mml:mi/><mml:mn mathvariant="normal">4</mml:mn><mml:mo>+</mml:mo></mml:msubsup></mml:mrow></mml:math></inline-formula>; Savarino and Legrand, 1998), methane (Fischer et al., 2008), carbon monoxide (CO; Wang et al., 2010), black carbon
(Han et al., 2012; Lehndorff et al., 2015), vanillic acid
(McConnell et al., 2007), and levoglucosan (Zennaro et al., 2014), as indicators of past fire activity. Laboratory and analytical
methods are also advancing through the use of image analysis for counting
charcoal and charcoal morphotypes (Enache and Cumming, 2006; Jensen et al., 2007; Thevenon and Anselmetti, 2007; Gu et al., 2008; Moos and
Cumming, 2012).</p>
      <p>Overall, the wealth of methods and approaches to fire research are providing
a broad range of insights into fire, both as an ecological process and as an
integrated component of the Earth system. However, much work remains to
understand the impact of wildfires and biomass burning emissions on climate,
and vice versa (Keywood et al., 2013). Research on human–fire
interactions using paleorecords is developing rapidly (Colombaroli et al.,
2008; Perry et al., 2012; McLauchlan et al., 2014; Munoz et al., 2014), but
applying insights from paleofire research to fire management and emissions
reduction plans remains comparatively limited (Whitlock et al., 2003; Cyr et al., 2009;
Coughlan and Petty, 2012;
Munoz et al., 2014). By compiling diverse types of paleofire data in a
central location and developing open-source analysis tools to explore those
data, research can advance more quickly on these topics.</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F1" specific-use="star"><caption><p>Location of paleofire sites and sampling density in the GCDv3.</p></caption>
        <?xmltex \igopts{width=426.791339pt}?><graphic xlink:href="https://bg.copernicus.org/articles/13/3225/2016/bg-13-3225-2016-f01.pdf"/>

      </fig>

</sec>
<sec id="Ch1.S3">
  <title>The Global Charcoal Database (version 3)</title>
      <p>The structure and contents of earlier versions of the GCD are outlined in
Power et al. (2010). Here we review the database design and focus
primarily on detailing new entries in GCDv3. Version 3 extends the total
number of sites in the GCD to 736. It includes 679 sites from version 2
(Daniau et al., 2012) as well as new sites from recent regional
syntheses from Australasia (Mooney et al.,
2011), the Americas (Marlon et al., 2009, 2012; Power et al., 2012),
and Europe (Vanniere et al., 2011).</p>
<sec id="Ch1.S3.SS1">
  <title>Geographical distribution</title>
      <p>Sites in GCDv3 come from five continents and exhibit a wide variety of
temporal resolutions (Fig. 1). Most of the sites (436) are located in the
Northern Hemisphere, which is due partly to its larger land area and partly
to sampling bias; 300 sites come from the Southern Hemisphere. About 20 %
of southern hemispheric sites (178) are located in the tropics; most are
located in forested regions, although sites increasingly come from
grasslands, shrublands, and woodlands as well. The geographical distribution
of the data reflects locations where fire research has traditionally focused
and the presence of suitable locations for paleoenvironmental indicators.
Sites are distributed between elevations of <inline-formula><mml:math display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>9 to 4060 m a.s.l., with more than
half (58 %) below 500 m a.s.l.; some records come from marine cores. Previous
analyses of the distribution of GCD sites in climate space showed that the
data set has relatively broad coverage with respect to global biomes and
climate gradients (Daniau et al., 2012). Many newly published fire-history records exist that can potentially be incorporated into subsequent
versions of the GCD (Brown, 2005; Han et al., 2012; Harley et al., 2012;
Iglesias et al., 2012; Daniau et al., 2013; Kelly et al., 2013;
Quintana-Krupinski et al., 2013; Tan and Huang, 2013; Cordeiro et al., 2014;
Courtney Mustaphi and Pisaric, 2014; Dunnette et al., 2014; Higuera et al.,
2014; Iglesias and Whitlock, 2014; Neumann et al., 2014; Walsh et al., 2015)
and many more are in development that will fill important spatial gaps where
fire is key, including Africa and the tropics.</p>
</sec>
<sec id="Ch1.S3.SS2">
  <title>Type of records, data entry, and database structure</title>
      <p>The majority of sites in the database are associated with a single record in
which charcoal was quantified using a single method. However, 96 sites in GCDv3
have more than one charcoal record, typically because charcoal was
quantified using multiple metrics or laboratory techniques.</p>
      <p>The charcoal data and metadata from the GCDv3 are stored in several formats.
The primary complete data set is stored in a Microsoft Access relational
database with four main and 23 supporting tables. The four main tables hold
(1) site metadata such as site name and type, geographical coordinates,
elevation, catchment size, data source, and dating type; (2) sample data,
including depths, volume, and estimated ages; (3) charcoal data, including
quantity, units, and quantification method; and (4) date information,
including depth and type of dates, laboratory identification numbers,
material dated, and associated errors. Additional tables include information
such as the contact (i.e., the corresponding data contributor) and
publications associated with each record, index tables (e.g., linking sites
to publications and contacts), and full descriptions of codes used in the
main tables. The original database was not designed to be a long-term
archival repository but rather a research database, and it is therefore
currently being replaced with a new structure. A significant percentage of
the site metadata, such as geographic characteristics and methodological
details, remains undocumented, however, and requires completion if
scientific questions that draw on such data are to be addressed.</p>
      <p>In addition to the database format, the GCDv3 data set is now available as
part of the paleofire R package (Blarquez et al., 2014) for use with
the R computer programming environment (R Development Core Team, 2013).
The R package currently lacks some of the metadata that are contained in the
full database, but the site metadata, charcoal data, and modeled ages are
available. The complete GCD in the form of a relational database can be
downloaded from <uri>paleofire.org</uri> and from the National Oceanic and Atmospheric
Administration's (NOAA) National Centers for Environmental Information
(NCEI; Power et al., 2008) website (<uri>http://www.ncdc.noaa.gov/paleo/impd/gcd.html</uri>).</p>

<?xmltex \floatpos{t}?><table-wrap id="Ch1.T1" specific-use="star"><caption><p>Total number of sites by sediment and measurement type. The
sediment types are lacustrine (LACU), bog (BOGM), unknown (NOTK), soil
(SOIL), coastal (COAS), and marine (MARI). The measurement types stored in
the database are concentration (CONC), influx, (INFL), proportions (C0P0; e.g.,
ratio of charcoal particles to pollen grains), and other (OTHE).</p></caption><oasis:table frame="topbot"><oasis:tgroup cols="7">
     <oasis:colspec colnum="1" colname="col1" align="left"/>
     <oasis:colspec colnum="2" colname="col2" align="right"/>
     <oasis:colspec colnum="3" colname="col3" align="right"/>
     <oasis:colspec colnum="4" colname="col4" align="right"/>
     <oasis:colspec colnum="5" colname="col5" align="right"/>
     <oasis:colspec colnum="6" colname="col6" align="right"/>
     <oasis:colspec colnum="7" colname="col7" align="right"/>
     <oasis:thead>
       <oasis:row rowsep="1">  
         <oasis:entry colname="col1"/>  
         <oasis:entry colname="col2">LACU</oasis:entry>  
         <oasis:entry colname="col3">BOGM</oasis:entry>  
         <oasis:entry colname="col4">NOTK</oasis:entry>  
         <oasis:entry colname="col5">SOIL</oasis:entry>  
         <oasis:entry colname="col6">COAS</oasis:entry>  
         <oasis:entry colname="col7">MARI</oasis:entry>
       </oasis:row>
     </oasis:thead>
     <oasis:tbody>
       <oasis:row>  
         <oasis:entry colname="col1">CONC</oasis:entry>  
         <oasis:entry colname="col2">178</oasis:entry>  
         <oasis:entry colname="col3">120</oasis:entry>  
         <oasis:entry colname="col4">33</oasis:entry>  
         <oasis:entry colname="col5">43</oasis:entry>  
         <oasis:entry colname="col6">22</oasis:entry>  
         <oasis:entry colname="col7">8</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">INFL</oasis:entry>  
         <oasis:entry colname="col2">157</oasis:entry>  
         <oasis:entry colname="col3">37</oasis:entry>  
         <oasis:entry colname="col4">9</oasis:entry>  
         <oasis:entry colname="col5">3</oasis:entry>  
         <oasis:entry colname="col6">4</oasis:entry>  
         <oasis:entry colname="col7">2</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">C0P0</oasis:entry>  
         <oasis:entry colname="col2">45</oasis:entry>  
         <oasis:entry colname="col3">37</oasis:entry>  
         <oasis:entry colname="col4">8</oasis:entry>  
         <oasis:entry colname="col5">4</oasis:entry>  
         <oasis:entry colname="col6">9</oasis:entry>  
         <oasis:entry colname="col7">2</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">OTHE</oasis:entry>  
         <oasis:entry colname="col2">10</oasis:entry>  
         <oasis:entry colname="col3">3</oasis:entry>  
         <oasis:entry colname="col4">2</oasis:entry>  
         <oasis:entry colname="col5">2</oasis:entry>  
         <oasis:entry colname="col6">0</oasis:entry>  
         <oasis:entry colname="col7">0</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Total</oasis:entry>  
         <oasis:entry colname="col2">390</oasis:entry>  
         <oasis:entry colname="col3">197</oasis:entry>  
         <oasis:entry colname="col4">52</oasis:entry>  
         <oasis:entry colname="col5">52</oasis:entry>  
         <oasis:entry colname="col6">35</oasis:entry>  
         <oasis:entry colname="col7">12</oasis:entry>
       </oasis:row>
     </oasis:tbody>
   </oasis:tgroup></oasis:table></table-wrap>

<?xmltex \floatpos{t}?><table-wrap id="Ch1.T2" specific-use="star"><caption><p>Total number of sites by sediment type and catchment size. The
sediment types are lacustrine (LACU), bog (BOGM), unknown (NOTK), soil
(SOIL), coastal (COAS), and marine (MARI).
Catchment sizes are small (SMAL; &lt; 10 km<inline-formula><mml:math display="inline"><mml:mrow><mml:msup><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msup><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula>, medium (MEDI; &gt; 10.1 and &lt; 500 km<inline-formula><mml:math display="inline"><mml:mrow><mml:msup><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msup><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula>, large (LARG; &gt; 500 km<inline-formula><mml:math display="inline"><mml:mrow><mml:msup><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msup><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula>,
and unknown (NOTK).</p></caption><oasis:table frame="topbot"><oasis:tgroup cols="7">
     <oasis:colspec colnum="1" colname="col1" align="left"/>
     <oasis:colspec colnum="2" colname="col2" align="right"/>
     <oasis:colspec colnum="3" colname="col3" align="right"/>
     <oasis:colspec colnum="4" colname="col4" align="right"/>
     <oasis:colspec colnum="5" colname="col5" align="right"/>
     <oasis:colspec colnum="6" colname="col6" align="right"/>
     <oasis:colspec colnum="7" colname="col7" align="right"/>
     <oasis:thead>
       <oasis:row rowsep="1">  
         <oasis:entry colname="col1"/>  
         <oasis:entry colname="col2">LACU</oasis:entry>  
         <oasis:entry colname="col3">BOGM</oasis:entry>  
         <oasis:entry colname="col4">NOTK</oasis:entry>  
         <oasis:entry colname="col5">SOIL</oasis:entry>  
         <oasis:entry colname="col6">COAS</oasis:entry>  
         <oasis:entry colname="col7">MARI</oasis:entry>
       </oasis:row>
     </oasis:thead>
     <oasis:tbody>
       <oasis:row>  
         <oasis:entry colname="col1">SMAL</oasis:entry>  
         <oasis:entry colname="col2">194</oasis:entry>  
         <oasis:entry colname="col3">100</oasis:entry>  
         <oasis:entry colname="col4">4</oasis:entry>  
         <oasis:entry colname="col5">13</oasis:entry>  
         <oasis:entry colname="col6">15</oasis:entry>  
         <oasis:entry colname="col7">0</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">MEDI</oasis:entry>  
         <oasis:entry colname="col2">33</oasis:entry>  
         <oasis:entry colname="col3">22</oasis:entry>  
         <oasis:entry colname="col4">2</oasis:entry>  
         <oasis:entry colname="col5">9</oasis:entry>  
         <oasis:entry colname="col6">7</oasis:entry>  
         <oasis:entry colname="col7">0</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">LARG</oasis:entry>  
         <oasis:entry colname="col2">14</oasis:entry>  
         <oasis:entry colname="col3">2</oasis:entry>  
         <oasis:entry colname="col4">7</oasis:entry>  
         <oasis:entry colname="col5">1</oasis:entry>  
         <oasis:entry colname="col6">0</oasis:entry>  
         <oasis:entry colname="col7">12</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">NOTK</oasis:entry>  
         <oasis:entry colname="col2">149</oasis:entry>  
         <oasis:entry colname="col3">73</oasis:entry>  
         <oasis:entry colname="col4">39</oasis:entry>  
         <oasis:entry colname="col5">29</oasis:entry>  
         <oasis:entry colname="col6">13</oasis:entry>  
         <oasis:entry colname="col7">0</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Total</oasis:entry>  
         <oasis:entry colname="col2">390</oasis:entry>  
         <oasis:entry colname="col3">197</oasis:entry>  
         <oasis:entry colname="col4">52</oasis:entry>  
         <oasis:entry colname="col5">52</oasis:entry>  
         <oasis:entry colname="col6">35</oasis:entry>  
         <oasis:entry colname="col7">12</oasis:entry>
       </oasis:row>
     </oasis:tbody>
   </oasis:tgroup></oasis:table></table-wrap>

      <p>Records in the GCD come from diverse environments (Tables 1, 2).
Most of the sites in the database (<inline-formula><mml:math display="inline"><mml:mrow><mml:mi>n</mml:mi><mml:mo>=</mml:mo></mml:mrow></mml:math></inline-formula> 390) are lacustrine, which are
primarily natural lakes that are often of glacial origin, but may also be of
tectonic, volcanic, or thermokarst origin. Other records (<inline-formula><mml:math display="inline"><mml:mrow><mml:mi>n</mml:mi><mml:mo>=</mml:mo></mml:mrow></mml:math></inline-formula> 197) are from
terrestrial environments, such as bogs, marshes, mires, and fens. A smaller
number of records were obtained from soils (<inline-formula><mml:math display="inline"><mml:mrow><mml:mi>n</mml:mi><mml:mo>=</mml:mo></mml:mrow></mml:math></inline-formula> 52) and from
coastal/fluvial (<inline-formula><mml:math display="inline"><mml:mrow><mml:mi>n</mml:mi><mml:mo>=</mml:mo></mml:mrow></mml:math></inline-formula> 35) or marine environments (<inline-formula><mml:math display="inline"><mml:mrow><mml:mi>n</mml:mi><mml:mo>=</mml:mo></mml:mrow></mml:math></inline-formula> 12). Depending on the
objective of a particular study, some site types will be more suitable than
others. Marine records, for example, are among the longest in the database,
making them suitable for analyses of biomass burning during the last glacial
cycle (Daniau et al., 2010). However, marine sites have
large catchment areas, making them suitable for regional but unsuitable for
fine-scale analyses of fire activity.</p>

<?xmltex \floatpos{t}?><table-wrap id="Ch1.T3" specific-use="star"><caption><p>Total number of sites by quantification type and laboratory
analysis method.</p></caption><oasis:table frame="topbot"><oasis:tgroup cols="5">
     <oasis:colspec colnum="1" colname="col1" align="left"/>
     <oasis:colspec colnum="2" colname="col2" align="right"/>
     <oasis:colspec colnum="3" colname="col3" align="right"/>
     <oasis:colspec colnum="4" colname="col4" align="right"/>
     <oasis:colspec colnum="5" colname="col5" align="right"/>
     <oasis:thead>
       <oasis:row>  
         <oasis:entry colname="col1"/>  
         <oasis:entry colname="col2">Proportion</oasis:entry>  
         <oasis:entry colname="col3">Concentration</oasis:entry>  
         <oasis:entry colname="col4">Influx</oasis:entry>  
         <oasis:entry colname="col5">Soil</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">  
         <oasis:entry colname="col1"/>  
         <oasis:entry colname="col2">(C0P0)</oasis:entry>  
         <oasis:entry colname="col3">(CONC)</oasis:entry>  
         <oasis:entry colname="col4">(INFL)</oasis:entry>  
         <oasis:entry colname="col5">(SOIL)</oasis:entry>
       </oasis:row>
     </oasis:thead>
     <oasis:tbody>
       <oasis:row>  
         <oasis:entry colname="col1">Soil charcoal (CPRO)</oasis:entry>  
         <oasis:entry colname="col2">0</oasis:entry>  
         <oasis:entry colname="col3">0</oasis:entry>  
         <oasis:entry colname="col4">0</oasis:entry>  
         <oasis:entry colname="col5">1</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Gravimetric (GRAV)</oasis:entry>  
         <oasis:entry colname="col2">1</oasis:entry>  
         <oasis:entry colname="col3">1</oasis:entry>  
         <oasis:entry colname="col4">0</oasis:entry>  
         <oasis:entry colname="col5">0</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Hand picked (HNPK)</oasis:entry>  
         <oasis:entry colname="col2">0</oasis:entry>  
         <oasis:entry colname="col3">7</oasis:entry>  
         <oasis:entry colname="col4">0</oasis:entry>  
         <oasis:entry colname="col5">0</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Heavy liquid preparation (HVLQ)</oasis:entry>  
         <oasis:entry colname="col2">0</oasis:entry>  
         <oasis:entry colname="col3">4</oasis:entry>  
         <oasis:entry colname="col4">0</oasis:entry>  
         <oasis:entry colname="col5">0</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Imaging analysis (IMAG)</oasis:entry>  
         <oasis:entry colname="col2">0</oasis:entry>  
         <oasis:entry colname="col3">12</oasis:entry>  
         <oasis:entry colname="col4">2</oasis:entry>  
         <oasis:entry colname="col5">0</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Oxidation resistant elemental</oasis:entry>  
         <oasis:entry colname="col2">0</oasis:entry>  
         <oasis:entry colname="col3">1</oasis:entry>  
         <oasis:entry colname="col4">0</oasis:entry>  
         <oasis:entry colname="col5">0</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Carbon OREC  % of dry weight (OREC)</oasis:entry>  
         <oasis:entry colname="col2"/>  
         <oasis:entry colname="col3"/>  
         <oasis:entry colname="col4"/>  
         <oasis:entry colname="col5"/>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Pollen slide (POLS)</oasis:entry>  
         <oasis:entry colname="col2">81</oasis:entry>  
         <oasis:entry colname="col3">259</oasis:entry>  
         <oasis:entry colname="col4">98</oasis:entry>  
         <oasis:entry colname="col5">0</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Sieved (SIEV)</oasis:entry>  
         <oasis:entry colname="col2">4</oasis:entry>  
         <oasis:entry colname="col3">151</oasis:entry>  
         <oasis:entry colname="col4">118</oasis:entry>  
         <oasis:entry colname="col5">0</oasis:entry>
       </oasis:row>
       <oasis:row>  
         <oasis:entry colname="col1">Total</oasis:entry>  
         <oasis:entry colname="col2">86</oasis:entry>  
         <oasis:entry colname="col3">435</oasis:entry>  
         <oasis:entry colname="col4">218</oasis:entry>  
         <oasis:entry colname="col5">1</oasis:entry>
       </oasis:row>
     </oasis:tbody>
   </oasis:tgroup></oasis:table></table-wrap>

      <?xmltex \floatpos{t}?><fig id="Ch1.F2" specific-use="star"><caption><p>Temporal and latitudinal distribution of dates used to develop
chronologies for records in the GCDv3 over the past 22 000 years.</p></caption>
          <?xmltex \igopts{width=398.338583pt}?><graphic xlink:href="https://bg.copernicus.org/articles/13/3225/2016/bg-13-3225-2016-f02.pdf"/>

        </fig>

</sec>
<sec id="Ch1.S3.SS3">
  <title>Charcoal quantification methods</title>
      <p>Important differences exist in the types of quantification methods within
the database. Taken together, the 736 sites in GCDv3 have 134 269 charcoal
samples with estimated ages. For most of the sites, charcoal is quantified
as concentration (<inline-formula><mml:math display="inline"><mml:mrow><mml:mi>n</mml:mi><mml:mo>=</mml:mo></mml:mrow></mml:math></inline-formula> 402) or influx (<inline-formula><mml:math display="inline"><mml:mrow><mml:mi>n</mml:mi><mml:mo>=</mml:mo></mml:mrow></mml:math></inline-formula> 212); 105 are expressed in terms
of charcoal to pollen ratios or similar measures of relative abundance; and
the remaining 17 sites have uncommon units, such as cumulative probabilities
or presence/absence of charcoal. Influx is the preferred unit of measurement
for most biomass burning reconstructions because it accounts for variations
in sedimentation rates over time, which can vary widely. If concentrations,
depths, and ages exist, then influx can be calculated prior to analyses.
Charcoal-to-pollen ratios, which were common in early analyses, are now
relatively rare due to the ambiguities inherent in their interpretation
(Conedera et al., 2009).</p>
      <p>Different laboratory methods are used to quantify charcoal (Table 3). The
majority of charcoal records included in the database (436 sites) are
quantified using the pollen-slide method (POLS); 271 sites by sieving method
(SIEV); 14 sites using image analysis (IMAG); and 15 sites were quantified
using other methods such as hand picking charcoal from soil samples,
gravimetric chemical assay (Winkler, 1985), and charcoal separation by
heavy liquid preparation. Several records included were based on the
cumulative probability of charcoal in alluvial fan deposits (Pierce et al., 2004), and several records employed other chemical, thermal, or optical
treatments or some combination of these methods to quantify black or
elemental carbon (Verardo et al., 1990).</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F3" specific-use="star"><caption><p>Example of untransformed and transformed charcoal influx (using
the Box–Cox transformation) from Lago de Acessa, Tuscany, Italy
(Vanniere et al., 2008). Number of particles per influx
class is shown (left panels).</p></caption>
          <?xmltex \igopts{width=398.338583pt}?><graphic xlink:href="https://bg.copernicus.org/articles/13/3225/2016/bg-13-3225-2016-f03.pdf"/>

        </fig>

</sec>
<sec id="Ch1.S3.SS4">
  <title>Chronology</title>
      <p>Accurate chronological dating of sediments is essential to paleo research.
The quantity and quality of dating controls in GCDv3 records vary widely
(Fig. 2). Some records have numerous, high-precision AMS radiocarbon dates,
while others have few dates and poorly constrained chronologies with high or
unknown uncertainties. Five common types of dates exist in the GCD,
including AMS <inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mn>14</mml:mn></mml:msup></mml:math></inline-formula>C, conventional <inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mn>14</mml:mn></mml:msup></mml:math></inline-formula>C, <inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mn>210</mml:mn></mml:msup></mml:math></inline-formula>Pb, pollen-based
correlations, and stratigraphy markers (e.g., tephras). Methods used to
develop long-record stratigraphies are based on <inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mn>234</mml:mn></mml:msup></mml:math></inline-formula>U <inline-formula><mml:math display="inline"><mml:mo>/</mml:mo></mml:math></inline-formula> <inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mn>230</mml:mn></mml:msup></mml:math></inline-formula>Th ratios
or orbital tie points. There are no major spatial patterns in the type of
dating methods used, aside from the terrestrial/marine distinction, and the
use of tephras in areas with volcanic activity (e.g., western coasts of the
Americas). Differences in tephra dates among several records in the Pacific
Northwest that use an ash layer associated with the eruption of Mount Mazama
around 7700 years before present (yr BP, where present is 1950 CE)
(Bacon, 1983) as a date in their age–depth models appear to need
revision, as the eruption date was subsequently dated in multiple studies to
7627 <inline-formula><mml:math display="inline"><mml:mo>±</mml:mo></mml:math></inline-formula> 150 cal yr BP (Hallett et al., 1997; Zdanowicz
et al., 1999). In general, radiocarbon dates (AMS or conventional) are the
most common dating method reported in the GCD. The <inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mn>210</mml:mn></mml:msup></mml:math></inline-formula>Pb dating is used
for dating uppermost sediments (i.e., spanning the past 150 years) because
<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mn>210</mml:mn></mml:msup></mml:math></inline-formula>Pb has the shortest half-life of the radioisotopes. When the
sediment–water interface is retrieved during coring and is undisturbed, that
core top sample is typically assigned the year in which the core was
obtained; this sample is marked as “stratigraphic” in the legend of Fig. 2 and accounts for the stack of orange-colored dots around 0 cal yr BP
(i.e., 1950 CE).</p>
</sec>
</sec>
<sec id="Ch1.S4">
  <title>Charcoal data standardization and compositing</title>
<sec id="Ch1.S4.SS1">
  <title>From raw data to standardized accumulation rates</title>
      <p>Charcoal measurements can be obtained in a variety of ways, but the most
common techniques employ particle counts, area measurements, or relative
abundances (Power et al., 2010). The effects of local site
characteristics such as lake size, watershed topography, and vegetation type
on absolute charcoal influx values (Marlon et al., 2006), along with
the diversity of quantification methods in common use (Conedera
et al., 2009), results in values that vary over 13 orders of magnitude (Power
et al., 2010), making it impossible at this time to directly compare metrics
of biomass burned among sites. Charcoal records therefore must be
standardized in order to examine relative changes in charcoal influx over
time (Power et al., 2010). Once standardized, charcoal influx
anomalies can be averaged from multiple records, even if the records are
based on different methods, creating a composite series in which maxima,
minima, trends, and other features can be identified and interpreted.</p>
      <p>The charcoal syntheses presented here were standardized using a protocol
(Marlon et al., 2008; Power et al., 2010) that
includes (1) transforming non-influx values (e.g., concentration expressed
as particles cm<inline-formula><mml:math display="inline"><mml:mrow><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">3</mml:mn></mml:mrow></mml:msup><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> to influx values (e.g., particles cm<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:msup></mml:math></inline-formula> yr<inline-formula><mml:math display="inline"><mml:mrow><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula> by dividing the concentration values by sample deposition times
(yr cm<inline-formula><mml:math display="inline"><mml:mrow><mml:msup><mml:mi/><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup><mml:mo>)</mml:mo></mml:mrow></mml:math></inline-formula>, (2) homogenizing the variance using the Box–Cox
transformation, (3) rescaling the values using a minimax transformation to
allow comparisons among sites, and (4)  rescaling values once more to
<inline-formula><mml:math display="inline"><mml:mi>z</mml:mi></mml:math></inline-formula> scores using a base period of 21 000 to 200 yrs BP. The base period ends
at 200 yrs BP because of the large human impacts on ignitions and
suppression during the 19th and 20th centuries, which if included
would obscure variability in charcoal accumulation rates prior to this
period. However, the transformed records do extend into the 20th
century (<inline-formula><mml:math display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>50 yr BP, where CE 1950 <inline-formula><mml:math display="inline"><mml:mo>=</mml:mo></mml:math></inline-formula> 0 BP). The most important step of the
transformation is the homogenization of the variance (Fig. 3), which serves
to make small-scale variations visible while also reducing the importance of
high-value outliers.</p>
</sec>
<sec id="Ch1.S4.SS2">
  <title>Compositing multiple standardized time series</title>
      <p>The purpose of compositing multiple charcoal records is to identify shared
features and trends in fire history that may exist in a given spatial or
temporal domain (e.g., North America during the Holocene). Given that
individual charcoal time series are typically highly variable, averaging
multiple records can provide insights into changes in fire history that only
manifest at broad spatial scales (e.g., the impact of a changing climate
within a given region). The variability in a record comes from a variety of
factors, including the stochastic nature of lightning-caused fires
(Bartlein et al., 2008), site-specific factors such as
topography, soils, and local vegetation that influence fire history, the
complexities of charcoal production, transportation, and deposition,
sediment sampling, and processing methods (Gavin et al., 2006). As
a result, composite curves that are based on few records also tend to show
relatively high variability (Fig. 4, top panel). As more records are
included in the composite curve, the curve becomes smoother and the
confidence intervals around the mean narrow (Fig. 4, middle and bottom
panels), because averaging among many sites necessarily reduces peaks and
other variations evident in individual sites.</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F4"><caption><p>Three 3000-year biomass burning curves from eastern North America
based on sites from an increasing number of adjacent grid cells show how the
reconstructions become smoother and confidence intervals narrow as the
number of sites and the spatial area included expand. Biomass burning
reconstruction based on two adjacent grid cells containing a total of 19
records (top panel); three adjacent grid cells containing 40 records (middle
panel), including the 19 from the top panel; and four adjacent grid cells
representing a total of 59 records (bottom panel), including all previous.
In all panels, red lines are based on 400-year smoothing windows, black
lines based on 200-year windows, and bootstrap 95 % confidence intervals
from resampling by site are shown as gray bands.</p></caption>
          <?xmltex \igopts{width=213.395669pt}?><graphic xlink:href="https://bg.copernicus.org/articles/13/3225/2016/bg-13-3225-2016-f04.pdf"/>

        </fig>

      <p>Although charcoal records are typically composited to examine trends in fire
history in a given geographic domain, composites can also be used to explore
additional research questions. For example, combining all available records
in the GCD from islands might yield insights into patterns of fire use
associated with human colonization
(McWethy et al., 2013). Alternatively,
contrasting fire history from lakes versus peat bogs or marine records might
yield insights into methodological questions about charcoal transportation
and deposition. Compositing all records available during a particular time
period may also offer insights into globally influential events like
potential comet impacts (or lack thereof)
(Marlon et al., 2009), volcanic
events (Marlon et al., 2012), or into the effects of abrupt
climate changes on fire (Daniau et al., 2010).</p>
      <p>Irrespective of the research question, the process for compositing records
is the same in each case. Each record is standardized as described above,
but only after it is resampled to a common temporal resolution
(“presampled”) in order to standardize the influence of each record on the
final composite curve. Presampling can be done using simple binning
techniques, but a preferred method is to fit a loess curve to the series at
regularly spaced target points (e.g., at 20-year intervals); the latter
smooths over uncertainties in the sediment data as well as in the age model,
whereas binning creates artificial cutoff points between samples that are
in reality uncertain. After presampling, the records are standardized using
a common base period, and a loess curve is again fitted to the pooled,
transformed data using a fixed window width (e.g., 1000 years to generate a
record of nominally “millennial-scale” variability). Composite curves in
this paper were produced following these methods as implemented in the R
paleofire package (Blarquez et al., 2014).</p>
      <p>Two issues that are not addressed by the above standardization and
compositing approach relate to age uncertainties and spatial
representativeness. While compositing many records can highlight regional
trends in biomass burning, the different temporal uncertainty in individual
records can make it difficult to accurately determine the precise timing of
changes or to explore questions about synchroneity, for example. The number
of radiocarbon dates or other chronological constraints in a record provide
information about age uncertainties, and these dates are available in the
GCD. However, formally assessing every age–depth model for the records in
the GCD is a non-trivial task and should ideally be undertaken with the
researchers who produced each record. Smoothing and gridding data accounts
for age uncertainty in the records informally because the process only
reveals trends and shifts in biomass burning that are robust across multiple
records. More detailed analysis will always be needed, however, to address
research questions about the sequence of particular changes or the precise
timing of specific events. Similarly, the varying spatial representativeness
of individual records are not accounted for in the compositing method
described here. The myriad factors that affect charcoal production,
transportation, and deposition in sediments means that there is no universal
relationship between charcoal quantities and area burned that can be applied
to all records. The conversion of all units to <inline-formula><mml:math display="inline"><mml:mi>z</mml:mi></mml:math></inline-formula> scores therefore allows the
detection of trends in biomass burning over time but removes any information
that may exist about the specific magnitude of area burned recorded by
different records that make up a composite curve.</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F5" specific-use="star"><caption><p>Trends in biomass burning (left panel) for the Northern
Hemisphere, globe, and Southern Hemisphere for the past 1000 years and
spatially gridded charcoal influx <inline-formula><mml:math display="inline"><mml:mi>z</mml:mi></mml:math></inline-formula> scores reflecting  biomass burning (right panel) for the period 1950–2010,
1850–1950, and 950–1050 CE. Vertical gray bars through the time series on
the left panel correspond to the time intervals shown in the gridded dot
maps on the right panel. The charcoal influx anomaly base period for all
panels is 1000–1800 CE. The smoothing window widths for the time series
(left panel) are 40 years (red line) and 20 years (black line).
Bootstrap-by-site confidence intervals (95 %) are filled in gray.</p></caption>
          <?xmltex \igopts{width=426.791339pt}?><graphic xlink:href="https://bg.copernicus.org/articles/13/3225/2016/bg-13-3225-2016-f05.pdf"/>

        </fig>

      <?xmltex \floatpos{t}?><fig id="Ch1.F6" specific-use="star"><caption><p>Trends in biomass burning (left panel) from 22 to 0 ka from the
GCDv3 (red) and GCDv2 (gray; Daniau et al., 2012) for the entire globe,
northern extratropics (&gt; 30<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula> N latitude), tropics
(&gt; 30<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula> N latitude and &lt; 30<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula> S
latitude), and the southern extratropics (&lt; 30<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula> S
latitude), along with spatially gridded charcoal influx <inline-formula><mml:math display="inline"><mml:mi>z</mml:mi></mml:math></inline-formula> scores reflecting biomass burning (right panel) for
the periods 0–1, 5.5–6.5, and 20.5–21.5 ka. Vertical gray bars on the
left panel correspond to the intervals shown in the maps (right panel). The
charcoal influx anomaly base period for all panels is 21 ka–200 cal yr BP;
the smoothing window width is 1000 years. Bootstrap-by-site confidence
intervals (95 %) are filled in gray.</p></caption>
          <?xmltex \igopts{width=426.791339pt}?><graphic xlink:href="https://bg.copernicus.org/articles/13/3225/2016/bg-13-3225-2016-f06.pdf"/>

        </fig>

</sec>
</sec>
<sec id="Ch1.S5">
  <title>The gridded charcoal data set</title>
      <p>To efficiently visualize GCDv3 and facilitate comparisons with model output,
we present a spatially gridded version of GCDv3 using dot maps (Figs. 5, 6)
alongside composite time-series curves (Figs. 5, 6). Vertical gray bars on
the composite graphs indicate the time periods reflected in the maps. Each
dot on the map represents a composite charcoal series constructed from all
records within a fixed distance of the dot, such that the area represented
by each dot is the same. However, the dots are positioned on a regular
latitude/longitude grid, and the area of each grid cell varies by latitude
(i.e., cells near the Equator cover larger areas than those near the poles);
spacing dots in this way maximizes the compatibility of the gridded charcoal
data set with other global data products. On such a grid, the absolute
distance between dots (or nodes) decreases with distance from the Equator.
We defined the radius used to identify sites contributing to a dot as half
the distance between diagonally adjacent dots at the Equator (e.g.,
<inline-formula><mml:math display="inline"><mml:mo>∼</mml:mo></mml:math></inline-formula> 395 km for a 5<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula> <inline-formula><mml:math display="inline"><mml:mo>×</mml:mo></mml:math></inline-formula> 5<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula> grid). This radius
ensures that all GCD sites contribute to at least one dot but also causes
sites to influence multiple dots, especially at high latitudes where dots
are relatively close together in terms of absolute distance (Fig. 7).
Finally, our gridding approach prevents interpolation into areas that are
not represented in the GCD, which is desirable given the great spatial
heterogeneity of fire regimes.</p>
      <p>Anomaly maps illustrate the gridding approach at six discrete intervals
during the past 1000 years (Fig. 5, left panel) and 22 000 years (Figs. 6,
left panel). Maps from each 100-year period during the past millennium and
each 1000-year interval since the Last Glacial Maximum (LGM) are provided in
the Supplement. The charcoal values are plotted on a
5<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula> grid, and the dots are colored and sized to reflect the value
and statistical significance, respectively, of the biomass burning anomalies
(Fig. 5 and 6, right panels). The maps include data from three 100-year
intervals (Fig. 5) and three 1000-year intervals (Fig. 6). Red dots on the
maps indicate positive mean <inline-formula><mml:math display="inline"><mml:mi>z</mml:mi></mml:math></inline-formula> scores for sites in that location relative to
their own long-term mean, which was calculated using a base period between
1000–200 years (Fig. 5) and 21 000–200 cal yr BP (Fig. 6). Blue dots on the
map indicate negative mean <inline-formula><mml:math display="inline"><mml:mi>z</mml:mi></mml:math></inline-formula> scores. Because each dot shows changes in
biomass burning relative to its own long-term average for that location, comparisons among
dot colors on a single map (i.e., for a specific time) cannot be used to
infer geographic patterns in biomass burning. For example, it is possible
(or very likely, in fact) that for a given time period, a blue dot in Africa
represents more biomass burning than a red dot in the Arctic. By contrast,
changes in the color of a dot over time indicate meaningful temporal
variability in the relative rate of biomass burning. A red dot in one time
period that changes to a blue dot in the same location at another time
period, for example, reflects an actual decrease in biomass burning over
time at that location. One point of note is that it is possible in some
cases for a recent time period to have less data than an older time period
because samples from sediment cores are not regularly spaced in time, and
core sections or tops are sometimes lost or destroyed in the field or during
extraction. Most lake sediments provide continuous records, but soil and bog
profiles often have hiatuses when sites dry out or peat is burned, and
occasionally this happens in lake and marine sediments as well. Another
reason that a site may have less data closer to present than in the distant
past is when sedimentation rates decline over time. In this case, a section
of the core the represents the most recent past may only have one or two
samples, whereas sections of the same size further down core may contain
many samples.</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F7"><caption><p>Diagnostic maps for the globally gridded data showing the number
of sites per grid cell at <bold>(a)</bold> 0–1, <bold>(b)</bold> 5.5–6.5, and <bold>(c)</bold> 20.5–21.5 ka.</p></caption>
        <?xmltex \igopts{width=199.169291pt}?><graphic xlink:href="https://bg.copernicus.org/articles/13/3225/2016/bg-13-3225-2016-f07.pdf"/>

      </fig>

      <p>A diagnostic map of the gridded charcoal data shows the effects of
summarizing all data within a constant specified distance from each dot
(Fig. 7). Effectively, the gridding approach allows each site to influence
an equivalent spatial area on the map. However, it is helpful to keep in
mind that given the same number of sites at high latitudes and at the
Equator, the high-latitude sites will be more smoothed relative to those at
the Equator, which is evident in the diagnostic maps from different time
periods. Another effect of using equal-area circles to construct the dot
maps is that a circle can be centered quite far from shore but still
encompass a site on land. Thus dots may represent terrestrial sites despite
being plotted in the ocean on our maps (although in some cases they
represent charcoal data actually collected from marine cores; see Figs. 1 and 7 for a comparison between location of sites and dots). Large (small)
dots indicate biomass burning anomalies that are (not) significantly
different from 0.</p>
      <p>Global biomass burning during the past millennium (Fig. 5) shows a gradual
long-term decline until the 17th century during the Little Ice Age
(LIA; Mann et al., 2009), as observed in previous reconstructions
(Marlon et al., 2008). This decline is more pronounced in
the Northern than Southern Hemisphere (Fig. 5, top and bottom panels). After
the LIA, global biomass burning increases gradually until the 19th
century, then rapidly until the 20th century. Maximum levels of biomass
burning in the Northern Hemisphere occur prior to maximum levels in the
Southern Hemisphere, and both hemispheres experience sharp declines in
biomass burning during the second half of the 20th century. The maps of
biomass burning show the spatial heterogeneity underlying the composite
curves. Biomass burning in central and eastern North America is highest from
1850 to 1950 CE, for example, whereas burning in western North America is
highest during the most recent period (1950–2010 CE). In contrast, burning
in western and southern Europe is generally higher 1000 years ago than it is
in the past two centuries. Burning in southeast Asia is very high from
1850 to 1950 CE and remains high in several locations for the period
1950–2010 CE where data are available.</p>
      <p>The most recent upturn in fire activity globally, but particularly in the
Northern Hemisphere reconstruction, is supported by a larger data set than
GCDv1. Marlon et al. (2008) used GCDv1 to document the large
decrease in biomass burning in the 20th century, but the reconstruction had
large uncertainties in the trend over the last few decades. The addition of
new records to versions 2 and 3 of the GCD, along with a finer-scale
temporal focus now reveals the most recent increases in fire activity
observed not only in the charcoal data but also in several lines of
independent evidence, including satellite and observational data (Giglio
et al., 2013; Dennison et al., 2014).</p>
      <p>Global biomass burning since the LGM, 21 000 years ago, shows a long-term
increase (Fig. 6) consistent with increasing temperatures, atmospheric
CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula> concentrations, and burnable biomass (Daniau et al., 2012; Martin
Calvo et al., 2014). The reconstructions from GCDv3 (red lines) are very
similar to those from GCDv2 (thin gray lines) for the globe, northern
extratropics (&gt; 30<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula> N latitude), tropics (&gt; 30<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula> N latitude and &lt; 30<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula> S latitude), and
southern extratropics (&lt; 30<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula> S latitude), with the
exception of burning in the northern extratropics during the LGM, which
registers as very low with the additional records in GCDv3 as compared with
GCDv2 (Fig. 6). However, the Northern and Southern hemispheres show somewhat
inverse patterns of burning during the Holocene, with fire increasing
steadily in the northern extratropics during the Holocene, but declining in
the early to mid-Holocene in the tropics and southern extratropics, before
increasing in the late Holocene.</p>
      <p>The gridded maps provide insight into the spatial variations in biomass
burning since the LGM.</p>
      <p>Burning is generally higher in the past millennium than at any time since
the LGM with the exception of central-western South America (Fig. 6), where
some locations had higher than average burning during the mid-Holocene and
below average burning in the past millennium. Levels of burning during the
LGM in turn were generally lower than at later periods, with a few localized
exceptions. Particularly high levels of biomass burning in the past
millennium are observed in many locations in the Southern Hemisphere (e.g.,
New Zealand, central Africa, the Amazon) as well as in parts of the Northern
Hemisphere (e.g., northeastern North America, southern California, and the
southern Iberian Peninsula). The maps also reveal spatial coherence in
regional biomass burning since the LGM, which likely reflects climate
controls on fire in some cases and human controls on fire in others – the
degree of coherence alone cannot distinguish causal mechanisms at this
scale.</p>
</sec>
<sec id="Ch1.S6">
  <title>Using charcoal data in model validation</title>
      <p>The development of the GCD is motivated by the need to understand the
history of fire on Earth and the linkages among fire, climate, vegetation,
and human activities. As the GCD continues to expand, the expectation is
that knowledge of fire histories will become more detailed. Analyzing
charcoal-based fire-history records with modern data from satellites
(e.g., van der Werf et al., 2010; Giglio et al., 2013), fire scars (e.g.,
Girardin and Sauchyn, 2008; Marlon et al., 2012), or historical records
(e.g., Mouillot et al., 2006; Lamarque et al., 2010) is necessary to connect
relative or qualitative variations in biomass burning from charcoal records
(Aleman et al., 2013) to quantitative estimates of burned area or
carbon emissions. To test hypotheses related to drivers of fire activity
over longer timescales, however, research needs to integrate paleofire data
with modeling approaches. As the spatial network of charcoal records become
denser, there is increasing opportunity to identify locations where varying
types of fire records overlap and thus more opportunities to study changes
in fire regimes that span multiple spatial and temporal scales.</p>
      <p>Fire modeling efforts have advanced rapidly in the last decade (Arora and
Boer, 2005; Kloster et al., 2010; Kelley et al., 2014; Lasslop et al., 2014; Yue
et al., 2014; Le Page et al., 2015), providing a better understanding of the
varied impacts that fires have on humans, the biosphere, and the atmosphere
(Harrison et al., 2010), as well as the mechanisms through which
climate changes and human activities affect fire regimes. Simulations of
fire activity using physically based empirical relationships between
flammability and its controlling variables, such as temperature and soil
moisture, have helped identify the global drivers of modern burning
(Arora and Boer, 2005; Kloster et al., 2010; Pechony and Shindell, 2010;
Thonicke et al., 2010; Li et al., 2013; Pfeiffer et al., 2013). Fire modeling
studies have also qualitatively compared paleofire trends with simulated
global fire activity (Pechony and Shindell, 2010; Kloster et al., 2012; Li
et al., 2013), but quantitative testing of the physically based relationships
that drive fire models – the mechanics of the models themselves – has only
focused on modern climate conditions thus far. As a result, large gaps in
knowledge exist about how fire, climate, vegetation, and humans interact
under different climate conditions and over long timescales. Despite the
fact that mechanistic global fire models remain largely untested outside
modern climate parameters, these models are being used to predict the
response of fires to ongoing climate change (Pechony and
Shindell, 2010; Kloster et al., 2012).</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F8"><caption><p>Modeled (filled grid boxes; Brücher et al., 2014)
vs. reconstructed (GCDv3) fire activity at global <bold>(a)</bold> and regional <bold>(b, c)</bold> scales. Both data and model represent millennial anomalies at 6 ka relative
to present (i.e., mean <inline-formula><mml:math display="inline"><mml:mi>z</mml:mi></mml:math></inline-formula> scores for 5.5–6.5 ka minus mean <inline-formula><mml:math display="inline"><mml:mi>z</mml:mi></mml:math></inline-formula> scores for 500 cal yr BP to present). In all panels, green and pink symbols indicate GCD
data that agree or disagree (respectively) with model output in terms of
the sign of the 6–0 ka anomaly. In <bold>(a)</bold> and <bold>(c)</bold> the data are gridded
following methods presented in Sect. 5. In <bold>(b)</bold>, anomalies for individual
GCD sites are plotted, with symbols indicating positive (“<inline-formula><mml:math display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>”) or negative
(“o”) anomalies; records that do not span the full 6 ka interval are shown
(gray squares) but excluded from the analysis.</p></caption>
        <?xmltex \igopts{width=241.848425pt}?><graphic xlink:href="https://bg.copernicus.org/articles/13/3225/2016/bg-13-3225-2016-f08.png"/>

      </fig>

      <p>The fire modeling studies that have explicitly considered paleofire data
provide examples of the challenges in comparing data and models. A study by
Pechony and Shindell (2010) tested a global fire model scheme
within a Global Climate Model simulation of the past millennium, for
example, and found that at coarse spatial scales precipitation was the most
important factor driving multi-centennial variations in fire activity in the
model. However, the spatial patterns underlying these trends, and the extent
to which finer-scale variations match paleofire evidence are unknown.
Moreover, the finding that precipitation is more important than temperature
in driving trends in fire activity globally contradicts analyses of
paleodata (Daniau et al., 2012; Marlon et al., 2012; Power et al., 2012;
Marlon et al., 2013), as well as satellite remote-sensing data
(Bistinas et al., 2013), raising key questions about how
temperature, precipitation, and their interactions affect variations in
global biomass burning. Another fire modeling study (Brücher
et al., 2014) compared model output to paleofire data from the GCD at
regional scales from the mid-Holocene until the pre-industrial era in the
18th century. Kloster et al. (2015) go one step further to test
the sensitivity of the same model to variations in fuel availability, fuel
moisture, and wind speed, as well as their synergy for the same regions and
time period.</p>
      <p>The new approach to gridding GCD data presented here (and included in the
paleofire R package) should help further paleofire data–model comparison studies.
Whereas modeling studies to date have focused on global or regional trends,
the growing number of records in the GCD allows for evaluation of model
performance at finer spatial scales. However, site-specific variability is often
high among charcoal records, and driver data sets for many global fire models
may be of relatively coarse resolution. As a result it is ill-advised to
compare model output to individual charcoal records. The gridded approach
offers a flexible compromise that can be tuned in terms of spatial
resolution depending on data availability, model driver data sets, and other
factors. As an example, we present here a global map of simulated area
burned using the CLIMBA model (Brücher et al., 2014),
overlaid with gridded composite charcoal anomalies from the GCD. CLIMBA
consists of the EMIC CLIMBER-2 (CLIMate and BiosphERe; Petoukhov et al., 2000; Ganopolski et al., 2001) and JSBACH (Raddatz et al.,
2007;
Brovkin et al., 2009; Reick et al., 2013; Schneck et al., 2013), which is the
land component of the Max Planck Institute Earth System Model (MPI-ESM,
Giorgetta et al., 2013). Simulated area burned throughout the Holocene was
treated analogously to GCD data to produce a gridded map of area-burned
anomalies at 6000 BP relative to present (i.e., 6000 BP <inline-formula><mml:math display="inline"><mml:mi>z</mml:mi></mml:math></inline-formula> scores minus 0 BP
<inline-formula><mml:math display="inline"><mml:mi>z</mml:mi></mml:math></inline-formula> scores).</p>
      <p>Overall, data–model agreement is weak, with many grid cells disagreeing in
terms of the sign of the anomaly estimated from the CLIMBA model versus GCD
data (Fig. 8a). However, the exercise shows promise for some regions. In
eastern North America, for example, site-level GCD data are difficult to
reconcile with model output (Fig. 8b), but the gridded data product shows
that both data and model generally agree that 6000 BP was a period of lower
fire activity than present for the region (Fig. 8c). It is beyond the scope
of this paper to evaluate the importance of this agreement, or the causes of
data–model mismatch in other regions throughout the globe. Instead, we
present the example as a proof-of-concept to motivate future studies.
Important basic research topics to pursue include evaluation of
spatiotemporal patterns in data–model comparisons, and a critical assessment
of how uncertainties in both GCD data and fire model output contribute to
the comparisons.</p>
      <p>Using fire-history data from the GCD to constrain fire model simulations, or
conversely, using fire model simulations to understand variability in the
fire-history data from the GCD, requires careful consideration of the
uncertainties associated with both data types. For paleofire records,
quantifying and accounting for age uncertainties is a major concern, but
progress is occurring on this front through the development of Bayesian
age-modeling methods (Blaauw and Christen, 2011; Goring
et al., 2012). Uncertainties in charcoal records also come from the many
natural processes related to charcoal production, transportation, and
deposition, which interact to produce variability in charcoal accumulation
over time. These processes are being studied through field experiments and
calibration studies that will enable the development of higher quality
fire-history reconstructions and a better understanding of uncertainties
(Tinner et al., 2006; Higuera et al., 2011; Aleman et al., 2013). An
important source of uncertainty in global fire models is the
parameterization of the processes most directly controlling fire activity
(e.g., human influence, climate influence; e.g., Pechony and Shindell,
2009; Pfeiffer et al., 2013). The sensitivity of simulated fire activity to
such parameterizations needs to be tested to understand model uncertainty.
Uncertainty in modern fire records arises from any extrapolation or
interpretation beyond the available fire records (Mouillot et al.,
2006) or to limits in the satellite data itself (Giglio et al.,
2013). With detailed considerations of both the limits and uncertainties of
all data sources and model parameterizations, connecting GCD to fire models
represents the natural evolution in the effort to understand fires in the
Earth system.</p>
</sec>
<sec id="Ch1.S7">
  <title>Future recommendations</title>
      <p>There are several research areas that, with further development, would
facilitate rapid integration of fire data and a more comprehensive
understanding of fire across spatiotemporal scales. Here we identify
particular areas that would help address specific barriers to progress in
paleofire research.
<list list-type="bullet"><list-item><p>Charcoal calibration studies in diverse environments. A major limitation of biomass burning reconstructions is that they can
only represent relative changes in burning from an arbitrary baseline.
Calibration studies that relate variability in charcoal accumulation to fire
regime characteristics from historical, fire-scar, satellite, and other
recent data could allow additional information to be obtained from charcoal
records. Given the complexities of charcoal production, transportation and
deposition, it is unlikely that the absolute amount of biomass burning from
a single paleofire time series can be known; however, with a better understanding
of how charcoal abundances relate quantitatively to area burned or other
fire-regime metrics, constraints on paleofire reconstructions can be
established and integrated into models that can then provide quantitative
estimates of variables like area burned and carbon emissions.</p></list-item><list-item><p>Multiproxy studies of paleofire history. Comparisons of paleofire data from multiple sources, such as charcoal,
black carbon, and levoglucosan, are needed to better understand the roles of
changes in area burned, fire frequency, fire type, and emissions in carbon
cycling and the climate system. The combustion of vegetation produces a wide
array of products, but many of these (e.g., ammonium and black carbon) are
not specific to biomass combustion. As a result, developing methods for
effectively comparing different types of data that imperfectly reflect fire
emissions may improve our understanding of fire by providing convergent
evidence for particular features, enhancing the temporal or spatial
resolution of reconstructions, or refining our understanding of proxy source
areas. By improving our ability to compare and integrate diverse sources of
fire-history information, we can more clearly identify and potentially
offset the weaknesses of each particular data type.</p></list-item><list-item><p>Data–model comparisons of paleofire history. A primary motivation for the development of the GCD has been to create
data sets for use in the development and validation of global fire
models. Mechanistic and process-based simulations of fire activity at
multiple spatiotemporal scales necessarily depend on an accurate
understanding of the controls of biomass burning. The GCD can directly
inform fire models on this point. Paleofire data–model comparisons are an
emerging field in many respects. Spatiotemporal comparisons of GCD to fire
model output will help move research forward into deeper analyses of how
uncertainties associated with both the data and the models contribute to our
collective understanding of paleofire history and implications for future
model-based fire projections.</p></list-item><list-item><p>Filling gaps in paleofire data. Data collection from regions that are presently underrepresented in the
GCD (e.g., Africa, the tropics, tundra and heathlands, and the boreal
forests of Eurasia) is essential for learning how fire varied in response to
climate forcings and human activity in the past, particularly in unique
vegetation types and in biodiversity hotspots. Understanding
fire–climate–vegetation interactions can supplement our knowledge from
data-poor areas, but given the contingencies and legacies that land-use
practices have on land cover and disturbance regimes
(McLauchlan et al., 2014), having data from specific
geographic locations is often necessary.</p></list-item><list-item><p>Comparisons between charcoal data and other spatially extensive data sets. The development of large environmental data sets during the past few
decades has opened up a new frontier in global change science. New research
into the interactions among climate, vegetation, human activities and fire
during the Holocene and in the more distant past can now be supported by
large simulated and observed paleoclimate data sets, pollen data sets, and
data on population growth, land-use, and land-cover change. Analyzing these
data sets jointly with the GCD can provide insights into how changes in fire
regimes affect rates of ecological change and biodiversity
(Colombaroli et al., 2012), how fire affects species
migration (Edwards et al., 2015), or whether humans altered the
climate system using fire in the early Holocene (Marlon et al.,
2013).</p></list-item></list></p>
      <p>In addition to the research needs above, several practices could aid in the
development of high quality charcoal-based fire-history reconstructions and
facilitate data integration across labs and therefore across different
environmental contexts. The practices may be more useful to new researchers
entering the field or establishing new labs.
<list list-type="bullet"><list-item><p>Continuous sampling of macroscopic charcoal data. Although many researchers now sample lacustrine sediment continuously and
quantify macroscopic charcoal, many continue to tally microscopic particles,
or to sample discontinuously. Taking the latter approach may be necessary
due to methodological, funding, or other constraints, but when it is
possible, the former approach is more desirable. Research on charcoal
particle size classes supports macroscopic particles (&gt; 100 <inline-formula><mml:math display="inline"><mml:mi mathvariant="normal">µ</mml:mi></mml:math></inline-formula>m) as a reliable indicator of local (within 1–10<inline-formula><mml:math display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula> kilometers of
a study site) fire activity (Whitlock and Bartlein, 2004),
whereas smaller particles integrate biomass burning from a larger spatial
domain (Conedera et al., 2009). If both macroscopic and
microscopic particles can be tallied, they may provide complimentary
evidence of past fire regime change. However, if only one particle size is
collected, analysis of macroscopic charcoal usually provides a better signal
for local fire reconstruction. While continuous sampling is more time and
cost intensive, it facilitates reconstructing event frequency, aligning
multiple cores, detecting unique events, and examining rates of change.</p></list-item><list-item><p>Separating woody and herbaceous charcoal. In environments that may have had grasses as a fuel source, separate
tallying of woody and herbaceous charcoal (e.g., Walsh et al.,
2008) can be of great value (e.g., Daniau et al., 2013) in
identifying temporal variability in fuel types. Additional charcoal
morphotypes can be observed and classified as well (Enache
and Cumming, 2006; Mustaphi and Pisaric, 2014), but the application of these
methods remains largely untested. In the meantime, separate tallies only of
woody and herbaceous charcoal have already been shown to provide reliable
information about fuel sources (e.g., Wooller et al., 2000; Walsh et al.,
2008; Maezumi et al., 2015) and are recommended when possible.</p></list-item><list-item><p>Data sharing and open-source code. The importance of data sharing, and increasingly code sharing, is now
widely recognized in the scientific community (Easterbrook, 2014). Sharing data and code facilitates and encourages
reproducibility, allows comparative data analysis, and promotes scientific
progress in general. Data sharing is also
essential for addressing questions at broad spatial scales, evaluating
alternative laboratory and analytical methods, and ensuring that limited
research funds are used efficiently. Although sharing data and code
introduces overhead costs for data management and archive maintenance, the
benefits to individuals, the scientific community, and the public at large
are increasingly recognized as far outweighing these costs. The research
presented in this paper is just one example of the science that is possible
with data and code sharing; we hope academic institutions, publishers, and
funders continue to encourage and incentivize such practices
(Kattge et al., 2014).</p></list-item></list></p>
</sec>
<sec id="Ch1.S8" sec-type="conclusions">
  <title>Conclusions</title>
      <p>The GCDv3 incorporates 736 charcoal records and can now be gridded globally
for the modeling community to ease future data–model comparisons. Fire-history reconstructions from the GCDv3 demonstrate that increases in biomass
burning since the last glacial period were widespread, as are unusually high
levels of burning over the past several decades. Present-day burning
inferred from the charcoal data is particularly high in western North
America and southeastern Australasia. Detailed reconstructions of temporal
variations in biomass burning during the past 1000 years reveal that a
global biomass burning decline from 1000 to the LIA was more pronounced in
the Northern than Southern Hemisphere. In addition, variations in fire
activity during the past 200 years show very different spatial patterns. In
general, data–model comparisons with paleofire data provide a powerful
method for testing hypotheses about interactions between climate and fire
outside the range of modern climate conditions. Results from such data–model
comparisons will highlight gaps and weaknesses in both data and models,
allowing targeted refinements to be identified and prioritized. We identify
five areas of focus to promote future progress in paleofire research,
including (1) charcoal calibration studies in diverse environments, (2) multiproxy studies of paleofire history, (3) paleofire data–model
comparisons, (4) filling gaps in paleofire data, (5) comparisons between
charcoal data and other large data sets, and (6) enhanced data extraction
from existing cores, like continuous sampling and herbaceous charcoal
identification.</p>
<sec id="Ch1.S8.SSx1" specific-use="unnumbered">
  <title>Data availability</title>
      <p>The complete GCDv1, v2, and v3 (this paper) Microsoft Access database with
all available metadata is stored and available at <uri>paleofire.org</uri>. Supporting
information about the Global Charcoal Database and the Global Palaeofire
Working Group is also available at <uri>paleofire.org</uri>. Site metadata and the charcoal
data are accessible through the paleofire package (Blarquez et al.,
2014) for R (R Development Core Team, 2013).</p>
</sec>
</sec>

      
      </body>
    <back><app-group>
        <supplementary-material position="anchor"><p><bold>The Supplement related to this article is available online at <inline-supplementary-material xlink:href="http://dx.doi.org/10.5194/bg-13-3225-2016-supplement" xlink:title="pdf">doi:10.5194/bg-13-3225-2016-supplement</inline-supplementary-material>.</bold></p></supplementary-material>
        </app-group><ack><title>Acknowledgements</title><p>We thank PAGES working group support for the GPWG. This research and paper
was initiated during the GPWG workshop 2013 held in Franche-Comté and
supported by the UMR Chrono-Environnement, the OREAS project, the University
of Franche-Comté, and the Région Franche-Comté. Jennifer R. Marlon is supported
by NSF grants BCS-1437074 and EF-1241870. Patrick Bartlein and Brian Magi are supported by NSF
grant BCS-1437074. Anne-Laure Daniau is supported by the project PICS CNRS 06484. Philip Higuera was
supported by NSF grant IIA-0966472. Boris Vannière is supported by the project
MISTRALS-PaleoMEX.
<?xmltex \hack{\newline}?><?xmltex \hack{\newline}?>
Edited by: A. Ito</p></ack><ref-list>
    <title>References</title>

      <ref id="bib1.bib1"><label>1</label><mixed-citation>
Aleman, J. C., Blarquez, O., Bentaleb, I., Bonté, P., Brossier, B., Carcaillet, C., Gond, V.,
Gourlet-Fleury, S., Kpolita, A., Lefèvre, I., Oslisly, R., Power, M. J.,
Yongo, O., Bremond, L., and Favier, C.: Tracking land-cover changes with
sedimentary charcoal in the Afrotropics, Holocene, 23, 1853–1862, 2013.</mixed-citation></ref>
      <ref id="bib1.bib2"><label>2</label><mixed-citation>
Arora, V. K. and Boer, G. J.: Fire as an interactive component of dynamic
vegetation models, J. Geophys. Res., 110, 1–20, 2005.</mixed-citation></ref>
      <ref id="bib1.bib3"><label>3</label><mixed-citation>
Bacon, C. R.: Eruptive history of Mount Mazama and Crater Lake Caldera,
Cascade Range, USA, J. Volcanol. Geoth. Res., 18,
57–117, 1983.</mixed-citation></ref>
      <ref id="bib1.bib4"><label>4</label><mixed-citation>
Bartlein, P., Hostetler, S. W., Shafer, S. L., Holman, J. O., and Solomon,
A. M.: Temporal and spatial structure in a daily wildfire-start data set
from the western United States (1986–96), Int. J. Wildland Fire, 17, 8–17, 2008.</mixed-citation></ref>
      <ref id="bib1.bib5"><label>5</label><mixed-citation>
Behling, H.: Late Quaternary environmental changes in the Lagoa da
Curuça region (eastern Amazonia, Brazil) and evidence of Podocarpus in
the Amazon lowland, Veg. Hist. Archaeobot., 10, 175–183,
2001.</mixed-citation></ref>
      <ref id="bib1.bib6"><label>6</label><mixed-citation>Bistinas, I., Oom, D., Saì, A. C. L., Harrison, S. P., Prentice, I. C., and
Pereira, J. M. C.: Relationships between human population density and burned
area at continental and global scales, PLoS ONE, 8, e81188, <ext-link xlink:href="http://dx.doi.org/10.1371/journal.pone.0081188" ext-link-type="DOI">10.1371/journal.pone.0081188</ext-link>, 2013.</mixed-citation></ref>
      <ref id="bib1.bib7"><label>7</label><mixed-citation>
Blaauw, M. and Christen, J. A.: Flexible paleoclimate age-depth models using
an autoregressive gamma process, Bayesian Analysis, 6, 457–474, 2011.</mixed-citation></ref>
      <ref id="bib1.bib8"><label>8</label><mixed-citation>
Blarquez, O., Vannière, B., Marlon, J. R., Daniau, A.-L., Power, M. J.,
Brewer, S., and Bartlein, P. J.: paleofire: An R package to analyse
sedimentary charcoal records from the Global Charcoal Database to
reconstruct past biomass burning, Comput. Geosci., 72, 255–261,
2014.</mixed-citation></ref>
      <ref id="bib1.bib9"><label>9</label><mixed-citation>
Bond, W. J. and Keeley, J. E.: Fire as a global 'herbivore': the ecology and
evolution of flammable ecosystems, Trends Ecol. Evol., 20,
387–394, 2005.</mixed-citation></ref>
      <ref id="bib1.bib10"><label>10</label><mixed-citation>Brovkin, V., Raddatz, T., Reick, C. H., Claussen, M., and Gayler, V.: Global
biogeophysical interactions between forest and climate, Geophys. Res. Lett., 36, L07405, <ext-link xlink:href="http://dx.doi.org/10.1029/2009GL037543" ext-link-type="DOI">10.1029/2009GL037543</ext-link>, 2009.</mixed-citation></ref>
      <ref id="bib1.bib11"><label>11</label><mixed-citation>
Brown, K. J., Clark, J. S., Grimm, E. C., Donovan, J. J., Mueller, P. G.,
Hansen, B. C. S., and Stefanova, I.: Fire cycles in North American interior
grasslands and their relation to prairie drought, P. Natl. Acad. Sci. USA, 102,
8865–8871, 2005.</mixed-citation></ref>
      <ref id="bib1.bib12"><label>12</label><mixed-citation>Brücher, T., Brovkin, V., Kloster, S., Marlon, J. R., and Power, M. J.: Comparing modelled fire dynamics
with charcoal records for the Holocene, Clim. Past, 10, 811–824, <ext-link xlink:href="http://dx.doi.org/10.5194/cp-10-811-2014" ext-link-type="DOI">10.5194/cp-10-811-2014</ext-link>, 2014.</mixed-citation></ref>
      <ref id="bib1.bib13"><label>13</label><mixed-citation>
Burney, D. A.: Late Quaternary stratigraphic charcoal records from
Madagascar, Quaternary Res., 28, 274–280, 1987.</mixed-citation></ref>
      <ref id="bib1.bib14"><label>14</label><mixed-citation>
Carcaillet, C., Bergeron, Y., Richard, P. J. H., Fréchette, B.,
Gauthier, S., and Prairie, Y. T.: Change of fire frequency in the eastern
Canadian boreal forests during the Holocene: does vegetation composition or
climate trigger the fire regime?, J. Ecol., 89, 930–946, 2001a.</mixed-citation></ref>
      <ref id="bib1.bib15"><label>15</label><mixed-citation>
Carcaillet, C., Bouvier, M., Fréchette, B., Larouche, A. C., and
Richard, P. J. H.: Comparison of pollen-slide and sieving methods in
lacustrine charcoal analyses for local and regional fire history, Holocene,
11, 467–476, 2001b.</mixed-citation></ref>
      <ref id="bib1.bib16"><label>16</label><mixed-citation>
Carcaillet, C., Almquist, H., Asnong, H., Bradshaw, R. H. W., Carrión,
J. S., Gaillard, M. J., Gajewski, K., Haas, J. N., Haberle, S. G., Hadorn,
P., Müller, S. D., Richard, P. J. H., Richoz, I., Rösch, M.,
Sánchez Goñi, M. F., Von Stedingk, H., Stevenson, A. C., Talon, B.,
Tardy, C., Tinner, W., Tryterud, E., Wick, L., and Willis, K. J.: Holocene
biomass burning and global dynamics of the carbon cycle, Chemosphere, 49,
845–863, 2002.</mixed-citation></ref>
      <ref id="bib1.bib17"><label>17</label><mixed-citation>
Clark, J. S.: Particle motion and the theory of charcoal analysis: source
area, transport, deposition, and sampling, Quaternary Res., 30, 67–80,
1988.</mixed-citation></ref>
      <ref id="bib1.bib18"><label>18</label><mixed-citation>
Clark, J. S. and Royall, P. D.: Local and regional sediment charcoal evidence for fire regimes in presettlement north-eastern North America,
J. Ecol., 84, 365–382, 1996.</mixed-citation></ref>
      <ref id="bib1.bib19"><label>19</label><mixed-citation>
Clark, J. S. and Patterson, W. A.: Background and local charcoal in
sediments: scales of fire evidence in the paleorecord, in: Sediment records
of biomass burning and global change, edited by: Clark, J. S., Cachier, H., Goldammer,
J. G., and Stocks, B. J., Springer-Verlag, Berlin, 1997.</mixed-citation></ref>
      <ref id="bib1.bib20"><label>20</label><mixed-citation>
Colombaroli, D., Tinner, W., Leeuwen, J. v., Noti, R., Vescovi, E.,
Vanniere, B., Magny, M., Schmidt, R., and Bugmann, H.: Response of
broadleaved evergreen Mediterranean forest vegetation to fire disturbance
during the Holocene: insights from the peri-Adriatic region, J. Biogeogr., 36, 314–326, 2009.</mixed-citation></ref>
      <ref id="bib1.bib21"><label>21</label><mixed-citation>
Colombaroli, D., Vanniere, B., Emmanuel, C., Magny, M., and Tinner, W.:
Fire-vegetation interactions during the Mesolithic-Neolithic transition at
Lago dell'Accesa, Tuscany, Italy, Holocene, 18, 679–692, 2008.</mixed-citation></ref>
      <ref id="bib1.bib22"><label>22</label><mixed-citation>Colombaroli, D., Beckmann, M., van der Knaap, W. O., Curdy, P., and Tinner,
W.: Changes in biodiversity and vegetation composition in the central Swiss
Alps during the transition from pristine forest to first farming, Divers. Distrib.,  19,  157–170,
<ext-link xlink:href="http://dx.doi.org/10.1111/j.1472-4642.2012.00930.x" ext-link-type="DOI">10.1111/j.1472-4642.2012.00930.x</ext-link>, 2013.</mixed-citation></ref>
      <ref id="bib1.bib23"><label>23</label><mixed-citation>
Conedera, M. and Tinner, W.: Long-term fire ecology of Switzerland, Journal
forestier suisse, 161, 424–432, 2010.</mixed-citation></ref>
      <ref id="bib1.bib24"><label>24</label><mixed-citation>
Conedera, M., Tinner, W., Neff, C., Meurer, M., Dickens, A. F., and Krebs,
P.: Reconstructing past fire regimes: methods, applications, and relevance
to fire management and conservation, Quaternary Sci. Rev., 28,
555–576, 2009.</mixed-citation></ref>
      <ref id="bib1.bib25"><label>25</label><mixed-citation>Cordeiro, R. C., Turcq, B. J., Moreira, L. S., de Aragão Rodrigues, R.,
Filho, F. F. L. S., Martins, G. S., Santos, A. B., Barbosa, M., da
Conceição, M. C. G., de Carvalho Rodrigues, R., Evangelista, H.,
Moreira-Turcq, P. F., Penido, Y. P., Sifeddine, A., and Seoane, J. C. S.:
Palaeofires in Amazon: Interplay between Land Use Change and Palaeoclimatic
Events, Palaeogeogr. Palaeocl., 415, 137–151, <ext-link xlink:href="http://dx.doi.org/10.1016/j.palaeo.2014.07.020" ext-link-type="DOI">10.1016/j.palaeo.2014.07.020</ext-link>, 2014.</mixed-citation></ref>
      <ref id="bib1.bib26"><label>26</label><mixed-citation>
Coughlan, M. R. and Petty, A. M.: Linking humans and fire: a proposal for a transdisciplinary fire ecology,
Int. J. Wildland Fire, 21, 477–487, 2012.</mixed-citation></ref>
      <ref id="bib1.bib27"><label>27</label><mixed-citation>
Courtney Mustaphi, C. J. and Pisaric, M. F. J.: Holocene
climate–fire–vegetation interactions at a subalpine watershed in
southeastern British Columbia, Canada, Quaternary Res., 81, 228–239,
2014.</mixed-citation></ref>
      <ref id="bib1.bib28"><label>28</label><mixed-citation>
Cyr, D., Gauthier, S., Bergeron, Y., and Carcaillet, C.: Forest management
is driving the eastern North American boreal forest outside its natural
range of variability, Front. Ecol. Environ., 7, 519–524,
2009.</mixed-citation></ref>
      <ref id="bib1.bib29"><label>29</label><mixed-citation>
Daniau, A.-L., Harrison, S. P., and Bartlein, P. J.: Fire regimes during the
last glacial, Quaternary Sci. Rev., 29, 2918–2930, 2010.</mixed-citation></ref>
      <ref id="bib1.bib30"><label>30</label><mixed-citation>
Daniau, A.-L., Goñi, M. F. S., Martinez, P., Urrego, D. H.,
Bout-Roumazeilles, V., Desprat, S., and Marlon, J. R.: Orbital-scale climate
forcing of grassland burning in southern Africa, P. Natl. Acad. Sci. USA, 110, 5069–5073, 2013.</mixed-citation></ref>
      <ref id="bib1.bib31"><label>31</label><mixed-citation>Daniau, A. L., Bartlein, P., Harrison, S., Prentice, I., Brewer, S.,
Friedlingstein, P., Harrisson Prentice, T., Inoue, J., Izumi, K., and
Marlon, J.: Predictability of biomass burning in response to climate
changes, Global Biogeochem. Cy., 26, GB4007, <ext-link xlink:href="http://dx.doi.org/10.1029/2011GB004249" ext-link-type="DOI">10.1029/2011GB004249</ext-link>, 2012.</mixed-citation></ref>
      <ref id="bib1.bib32"><label>32</label><mixed-citation>
DeBano, L. F.: The role of fire and soil heating on water repellency in
wildland environments: a review, J. Hydrol., 231, 195–206, 2000.</mixed-citation></ref>
      <ref id="bib1.bib33"><label>33</label><mixed-citation>
Delcourt, P. A., Delcourt, H. R., Ison, C. R., Sharp, W. E., and Gremillion,
K. J.: Prehistoric human use of fire, the Eastern Agricultural Complex, and
Appalachian oak-chestnut forests: Paleoecology of Cliff Palace Pond,
Kentucky, Am. Antiquity, 63, 263–278, 1998.</mixed-citation></ref>
      <ref id="bib1.bib34"><label>34</label><mixed-citation>
Dennison, P. E., Brewer, S. C., Arnold, J. D., and Moritz, M. A.: Large
wildfire trends in the western United States, 1984–2011, Geophys. Res. Lett., 41, 2928–2933, 2014.</mixed-citation></ref>
      <ref id="bib1.bib35"><label>35</label><mixed-citation>
Dunnette, P. V., Higuera, P. E., McLauchlan, K. K., Derr, K. M., Briles, C.
E., and Keefe, M. H.: Biogeochemical impacts of wildfires over four
millennia in a Rocky Mountain subalpine watershed, New Phytol., 203,
900–912, 2014.</mixed-citation></ref>
      <ref id="bib1.bib36"><label>36</label><mixed-citation>
Easterbrook, S. M.: Open code for open science?, Nat. Geosci., 7, 779–781,
2014.</mixed-citation></ref>
      <ref id="bib1.bib37"><label>37</label><mixed-citation>
Edwards, M., Franklin-Smith, L., Clarke, C., Baker, J., Hill, S., and
Gallagher, K.: The role of fire in the mid-Holocene arrival and expansion of
lodgepole pine (Pinus contorta var. latifolia Engelm. ex S. Watson) in
Yukon, Canada, Holocene, 25, 64–78, 2015.</mixed-citation></ref>
      <ref id="bib1.bib38"><label>38</label><mixed-citation>
Enache, M. D. and Cumming, B. F.: Tracking recorded fires using charcoal
morphology from the sedimentary sequence of Prosser Lake, British Columbia
(Canada), Quaternary Res., 65, 282–292, 2006.</mixed-citation></ref>
      <ref id="bib1.bib39"><label>39</label><mixed-citation>
Falk, D. A., Heyerdahl, E. K., Brown, P. M., Farris, C., Fulé, P. Z.,
McKenzie, D., Swetnam, T. W., Taylor, A. H., and Van Horne, M. L.:
Multi-scale controls of historical forest-fire regimes: new insights from
fire-scar networks, Front. Ecol. Environ., 9, 446–454,
2011.</mixed-citation></ref>
      <ref id="bib1.bib40"><label>40</label><mixed-citation>Fischer, H., Behrens, M., Bock, M., Richter, U., Schmitt, J., Loulergue, L.,
Chappellaz, J., Spahni, R., Blunier, T., Leuenberger, M., and Stocker, T.
F.: Changing boreal methane sources and constant biomass burning during the
last termination, Nature, 452,  864–867, <ext-link xlink:href="http://dx.doi.org/10.1038/nature06825" ext-link-type="DOI">10.1038/nature06825</ext-link>, 2008.</mixed-citation></ref>
      <ref id="bib1.bib41"><label>41</label><mixed-citation>
Fuller, J. L., Foster, D. R., McLachlan, J. S., and Drake, N.: Impact of
human activity on regional forest composition and dynamics in central New
England, Ecosystems, 1, 76–95, 1998.</mixed-citation></ref>
      <ref id="bib1.bib42"><label>42</label><mixed-citation>
Ganopolski, A., Petoukhov, V., Rahmstorf, S., Brovkin, V., Claussen, M.,
Eliseev, A., and Kubatzki, C.: CLIMBER-2: a climate system model of
intermediate complexity, Part II: model sensitivity, Clim. Dynam., 17,
735–751, 2001.</mixed-citation></ref>
      <ref id="bib1.bib43"><label>43</label><mixed-citation>
Gavin, D. G., Hu, F. S., Lertzman, K., and Corbett, P.: Weak climatic
control of stand-scale fire history during the late Holocene, Ecology, 87,
1722–1732, 2006.</mixed-citation></ref>
      <ref id="bib1.bib44"><label>44</label><mixed-citation>
Gavin, D. G., Hallett, D. J., Hu, F. S., Lertzman, K. P., Prichard, S. J.,
Brown, K. J., Lynch, J. A., Bartlein, P., and Peterson, D. L.: Forest fire
and climate change in western North America: insights from sediment charcoal
records, Front. Ecol. Environ., 5, 499–506, 2007.</mixed-citation></ref>
      <ref id="bib1.bib45"><label>45</label><mixed-citation>
Giglio, L., Randerson, J. T., and van der Werf, G. R.: Analysis of daily,
monthly, and annual burned area using the fourth-generation global fire
emissions database (GFED4), J. Geophys. Res.-Biogeo.,
118, 317–328, 2013.</mixed-citation></ref>
      <ref id="bib1.bib46"><label>46</label><mixed-citation>
Giorgetta, M. A., Jungclaus, J., Reick, C. H., Legutke, S., Bader, J.,
Böttinger, M., Brovkin, V., Crueger, T., Esch, M., and Fieg, K.: Climate
and carbon cycle changes from 1850 to 2100 in MPI ESM simulations for the
Coupled Model Intercomparison Project phase 5, Journal of Advances in Modeling Earth Systems, 5, 572–597, 2013.</mixed-citation></ref>
      <ref id="bib1.bib47"><label>47</label><mixed-citation>
Girardin, M. P. and Sauchyn, D.: Three centuries of annual area burned
variability in northwestern North America inferred from tree rings,
Holocene, 18, 205–214, 2008.</mixed-citation></ref>
      <ref id="bib1.bib48"><label>48</label><mixed-citation>
Goring, S., Williams, J. W., Blois, J. L., Jackson, S. T., Paciorek, C.,
Booth, R. K., Marlon, J. R., Blaauw, M., and Andres, C.: Deposition times in
the northeastern United States during the Holocene: establishing valid
priors for Bayesian age models, Quaternary Sci. Rev., 48, 54–60,
2012.</mixed-citation></ref>
      <ref id="bib1.bib49"><label>49</label><mixed-citation>
Gu, Y. S., Pearsall, D. M., Xie, S. C., and Yu, J. X.: Vegetation and fire
history of a Chinese site in southern tropical Xishuangbanna derived from
phytolith and charcoal records from Holocene sediments, J. Biogeogr., 35, 325–341, 2008.</mixed-citation></ref>
      <ref id="bib1.bib50"><label>50</label><mixed-citation>
Haberle, S. G.: Late quaternary vegetation change in the Tari Basin, Papua
New Guinea, Palaeogeogr. Palaeocl., 137, 1–24,
1998.</mixed-citation></ref>
      <ref id="bib1.bib51"><label>51</label><mixed-citation>
Haberle, S. G. and Ledru, M. P.: Correlations among charcoal records of
fires from the past 16 000 years in Indonesia, Papua New Guinea, and Central
and South America, Quaternary Res., 55, 97–104, 2001.</mixed-citation></ref>
      <ref id="bib1.bib52"><label>52</label><mixed-citation>
Hallett, D., Hills, L. V., and Clague, J. J.: New accelerator mass
spectrometry radiocarbon ages for the Mazama tephra layer from Kootenay
National Park, British Columbia, Canada, Can. J. Earth Sci.,
34, 1202–1209, 1997.</mixed-citation></ref>
      <ref id="bib1.bib53"><label>53</label><mixed-citation>Han, Y. M., Marlon, J. R., Cao, J. J., Jin, Z. D., and An, Z. S.: Holocene
biomass burning trends in China from soot, char and charcoal in lake
sediments, Global Biogeochem. Cy., 26,  GB4017, <ext-link xlink:href="http://dx.doi.org/10.1029/2011GB004197" ext-link-type="DOI">10.1029/2011GB004197</ext-link>, 2012.</mixed-citation></ref>
      <ref id="bib1.bib54"><label>54</label><mixed-citation>
Harley, G. L., Grissino-Mayer, H. D., and Horn, S. P.: Fire history and
forest structure of an endangered subtropical ecosystem in the Florida Keys,
USA, Int. J. Wildland Fire, 104, 1–19, 2012.</mixed-citation></ref>
      <ref id="bib1.bib55"><label>55</label><mixed-citation>
Harrison, S. P., Marlon, J. R., and Bartlein, P. J.: Fire in the Earth
System, in: Changing Climates, Earth Systems and Society, edited by: Dodson, J.,
Springer, Dordrecht, The Netherlands, 2010.</mixed-citation></ref>
      <ref id="bib1.bib56"><label>56</label><mixed-citation>Herridge, V., Birch, S. P., and Law, M.: Open Quaternary: A New, Open Access
J. Quaternary Res., Open Quaternary, 1, <ext-link xlink:href="http://dx.doi.org/10.5334/oq.ad" ext-link-type="DOI">10.5334/oq.ad</ext-link>, 2015.</mixed-citation></ref>
      <ref id="bib1.bib57"><label>57</label><mixed-citation>
Heusser, C. J.: Three late Quaternary pollen diagrams from southern
Patagonia and their palaeoecological implications, Palaeogeogr. Palaeocl., 118, 1–24, 1995.</mixed-citation></ref>
      <ref id="bib1.bib58"><label>58</label><mixed-citation>
Higuera, P., Gavin, D., Bartlein, P., and Hallett, D.: Peak detection in
sediment-charcoal records: impacts of alternative data analysis methods on
fire-history interpretations, Int. J. Wildland Fire, 19,
996–1014, 2010.</mixed-citation></ref>
      <ref id="bib1.bib59"><label>59</label><mixed-citation>
Higuera, P. E., Peters, M. E., Brubaker, L. B., and Gavin, D. G.:
Understanding the origin and analysis of sediment-charcoal records with a
simulation model, Quaternary Sci. Rev., 26, 1790–1809, 2007.</mixed-citation></ref>
      <ref id="bib1.bib60"><label>60</label><mixed-citation>
Higuera, P. E., Whitlock, C., and Gage, J.: Linking tree-ring and
sediment-charcoal records to reconstruct fire occurrence and area burned in
subalpine forests of Yellowstone National Park, USA,  Holocene 21,
327–341, 2011.</mixed-citation></ref>
      <ref id="bib1.bib61"><label>61</label><mixed-citation>
Higuera, P. E., Briles, C. E., and Whitlock, C.: Fire-regime complacency and
sensitivity to centennial-through millennial-scale climate change in Rocky
Mountain subalpine forests, Colorado, USA, J. Ecol., 102,
1429–1441, 2014.</mixed-citation></ref>
      <ref id="bib1.bib62"><label>62</label><mixed-citation>Iglesias, V. and Whitlock, C.: Fire responses to postglacial climate change
and human impact in northern Patagonia (41–43<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula> S), P. Natl. Acad. Sci. USA, 111, E5545–E5554, 2014.</mixed-citation></ref>
      <ref id="bib1.bib63"><label>63</label><mixed-citation>Iglesias, V., Whitlock, C., Bianchi, M. M., Villarosa, G., and Outes, V.:
Holocene climate variability and environmental history at the Patagonian
forest/steppe ecotone: Lago Mosquito (42<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula>29<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>′</mml:mo></mml:msup></mml:math></inline-formula>37.89<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>′</mml:mo><mml:mo>′</mml:mo></mml:mrow></mml:msup></mml:math></inline-formula> S, 71<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula>24<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>′</mml:mo></mml:msup></mml:math></inline-formula>14.57<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>′</mml:mo><mml:mo>′</mml:mo></mml:mrow></mml:msup></mml:math></inline-formula> W) and Laguna del Condor (42<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula>20<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>′</mml:mo></mml:msup></mml:math></inline-formula>47.22<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>′</mml:mo><mml:mo>′</mml:mo></mml:mrow></mml:msup></mml:math></inline-formula> S,
71<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula>17<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>′</mml:mo></mml:msup></mml:math></inline-formula>07.62<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mrow><mml:mo>′</mml:mo><mml:mo>′</mml:mo></mml:mrow></mml:msup></mml:math></inline-formula> W), Holocene, 22, 1297–1307, 2012.</mixed-citation></ref>
      <ref id="bib1.bib64"><label>64</label><mixed-citation>
Jensen, K., Lynch, E. A., Calcote, R., and Hotchkiss, S. C.: Interpretation
of charcoal morphotypes in sediments from Ferry Lake, Wisconsin, USA: do
different plant fuel sources produce distinctive charcoal morphotypes?,
Holocene, 17, 907–915, 2007.</mixed-citation></ref>
      <ref id="bib1.bib65"><label>65</label><mixed-citation>
Kattge, J., Diaz, S., and Wirth, C.: Of carrots and sticks, Nat. Geosci.,
7, 778–779, 2014.</mixed-citation></ref>
      <ref id="bib1.bib66"><label>66</label><mixed-citation>
Kehrwald, N., Whitlock, C., Barbante, C., Brovkin, V., Daniau, A.-L.,
Kaplan, J., Marlon, J. R., Power, M. J., Thonicke, K., and Van Der Werf, G.
R.: Recent advancements in wildfire research, Eos Trans. AGU, 421–423, 2013.</mixed-citation></ref>
      <ref id="bib1.bib67"><label>67</label><mixed-citation>Kelley, D. I., Harrison, S. P., and Prentice, I. C.: Improved simulation of fire-vegetation interactions in the Land surface Processes and
eXchanges dynamic global vegetation model (LPX-Mv1), Geosci. Model Dev., 7, 2411–2433, <ext-link xlink:href="http://dx.doi.org/10.5194/gmd-7-2411-2014" ext-link-type="DOI">10.5194/gmd-7-2411-2014</ext-link>, 2014.</mixed-citation></ref>
      <ref id="bib1.bib68"><label>68</label><mixed-citation>
Kelly, R., Chipman, M. L., Higuera, P. E., Stefanova, I., Brubaker, L. B.,
and Hu, F. S.: Recent burning of boreal forests exceeds fire regime limits
of the past 10 000 years, P. Natl. Acad. Sci. USA,
110, 13055–13060, 2013.</mixed-citation></ref>
      <ref id="bib1.bib69"><label>69</label><mixed-citation>
Keywood, M., Kanakidou, M., Stohl, A., Dentener, F., Grassi, G., Meyer, C.
P., Torseth, K., Edwards, D., Thompson, A. M., Lohmann, U., and Burrows, J.:
Fire in the Air: Biomass Burning Impacts in a Changing Climate, Critical Reviews in Environmental Science and Technology, 43, 40–83, 2013.</mixed-citation></ref>
      <ref id="bib1.bib70"><label>70</label><mixed-citation>Kloster, S., Mahowald, N. M., Randerson, J. T., Thornton, P. E., Hoffman, F. M., Levis, S., Lawrence, P. J.,
Feddema, J. J., Oleson, K. W., and Lawrence, D. M.: Fire dynamics during the 20th century simulated by the Community Land
Model, Biogeosciences, 7, 1877–1902, <ext-link xlink:href="http://dx.doi.org/10.5194/bg-7-1877-2010" ext-link-type="DOI">10.5194/bg-7-1877-2010</ext-link>, 2010.</mixed-citation></ref>
      <ref id="bib1.bib71"><label>71</label><mixed-citation>Kloster, S., Mahowald, N. M., Randerson, J. T., and Lawrence, P. J.: The impacts of climate, land use, and demography on fires
during the 21st century simulated by CLM-CN, Biogeosciences, 9, 509–525, <ext-link xlink:href="http://dx.doi.org/10.5194/bg-9-509-2012" ext-link-type="DOI">10.5194/bg-9-509-2012</ext-link>, 2012.</mixed-citation></ref>
      <ref id="bib1.bib72"><label>72</label><mixed-citation>Kloster, S., Brücher, T., Brovkin, V., and Wilkenskjeld, S.: Controls on fire activity over the Holocene,
Clim. Past, 11, 781–788, <ext-link xlink:href="http://dx.doi.org/10.5194/cp-11-781-2015" ext-link-type="DOI">10.5194/cp-11-781-2015</ext-link>, 2015.</mixed-citation></ref>
      <ref id="bib1.bib73"><label>73</label><mixed-citation>Lamarque, J.-F., Bond, T. C., Eyring, V., Granier, C., Heil, A., Klimont, Z., Lee, D., Liousse, C., Mieville, A.,
Owen, B., Schultz, M. G., Shindell, D., Smith, S. J., Stehfest, E., Van Aardenne, J., Cooper, O. R., Kainuma, M.,
Mahowald, N., McConnell, J. R., Naik, V., Riahi, K., and van Vuuren, D. P.: Historical (1850–2000) gridded anthropogenic and
biomass burning emissions of reactive gases and aerosols: methodology and application, Atmos. Chem. Phys., 10, 7017–7039, <ext-link xlink:href="http://dx.doi.org/10.5194/acp-10-7017-2010" ext-link-type="DOI">10.5194/acp-10-7017-2010</ext-link>, 2010.</mixed-citation></ref>
      <ref id="bib1.bib74"><label>74</label><mixed-citation>
Lasslop, G., Thonicke, K., and Kloster, S.: Spitfire within the mpi earth
system model: Model development and evaluation, Advances in Modeling Earth Systems, 6, 740–755, 2014.</mixed-citation></ref>
      <ref id="bib1.bib75"><label>75</label><mixed-citation>Le Page, Y., Morton, D., Bond-Lamberty, B., Pereira, J. M. C., and Hurtt, G.: HESFIRE: a global fire model to explore the
role of anthropogenic and weather drivers, Biogeosciences, 12, 887–903, <ext-link xlink:href="http://dx.doi.org/10.5194/bg-12-887-2015" ext-link-type="DOI">10.5194/bg-12-887-2015</ext-link>, 2015.</mixed-citation></ref>
      <ref id="bib1.bib76"><label>76</label><mixed-citation>
Lehndorff, E., Wolf, M., Litt, T., Brauer, A., and Amelung, W.: 15 000 years
of black carbon deposition – A post-glacial fire record from maar lake
sediments (Germany), Quaternary Sci. Rev., 110, 15–22, 2015.</mixed-citation></ref>
      <ref id="bib1.bib77"><label>77</label><mixed-citation>Li, F., Levis, S., and Ward, D. S.: Quantifying the role of fire in the Earth system – Part 1: Improved global fire modeling in
the Community Earth System Model (CESM1), Biogeosciences, 10, 2293–2314, <ext-link xlink:href="http://dx.doi.org/10.5194/bg-10-2293-2013" ext-link-type="DOI">10.5194/bg-10-2293-2013</ext-link>, 2013.</mixed-citation></ref>
      <ref id="bib1.bib78"><label>78</label><mixed-citation>Maezumi, S. Y., Power, M. J., Mayle, F. E., McLauchlan, K. K., and Iriarte, J.: Effects of past climate variability on
fire and vegetation in the cerrãdo savanna of the Huanchaca Mesetta, NE Bolivia, Clim. Past, 11, 835–853, <ext-link xlink:href="http://dx.doi.org/10.5194/cp-11-835-2015" ext-link-type="DOI">10.5194/cp-11-835-2015</ext-link>, 2015.</mixed-citation></ref>
      <ref id="bib1.bib79"><label>79</label><mixed-citation>
Mann, M. E., Zhang, Z., Rutherford, S., Bradley, R. S., Hughes, M. K.,
Shindell, D., Ammann, C., Faluvegi, G., and Ni, F.: Global signatures and
dynamical origins of the Little Ice Age and Medieval Climate Anomaly,
Science, 326, 1256–1260, 2009.</mixed-citation></ref>
      <ref id="bib1.bib80"><label>80</label><mixed-citation>
Marlon, J., Bartlein, P. J., and Whitlock, C.: Fire-fuel-climate linkages in
the northwestern USA during the Holocene, Holocene, 16, 1059–1071, 2006.</mixed-citation></ref>
      <ref id="bib1.bib81"><label>81</label><mixed-citation>
Marlon, J., Bartlein, P., Carcaillet, C., Gavin, D. G., Harrison, S. P.,
Higuera, P. E., Joos, F., Power, M. J., and Prentice, C. I.: Climate and
human influences on global biomass burning over the past two millennia,
Nat. Geosci., 1, 697–701, 2008.</mixed-citation></ref>
      <ref id="bib1.bib82"><label>82</label><mixed-citation>
Marlon, J., Bartlein, P., Walsh, M. K., Harrison, S. P., Brown, K. J.,
Edwards, M. E., Higuera, P. E., Power, M. J., Anderson, R. S., Briles, C.
E., Brunelle, A., Carcaillet, C., Daniels, M., Hu, F. S., Lavoie, M., Long,
C. J., Minckley, T., Richard, P. J. H., Scott, A. C., Shafer, D. S., Tinner,
W., Umbanhowar Jr, C. E., and Whitlock, C.: Wildfire responses to abrupt
climate change in North America, P. Natl. Acad. Sci. USA, 106, 2519–2524, 2009.</mixed-citation></ref>
      <ref id="bib1.bib83"><label>83</label><mixed-citation>
Marlon, J. R., Bartlein, P. J., Gavin, D. G., Long, C. J., Anderson, R. S.,
Briles, C. E., Brown, K. J., Colombaroli, D., Hallett, D. J., Power, M. J.,
Scharf, E. A., and Walsh, M. K.: Long-term perspective on wildfires in the
western USA, P. Natl. Acad. Sci. USA, 109,
E535–E543, 2012.</mixed-citation></ref>
      <ref id="bib1.bib84"><label>84</label><mixed-citation>
Marlon, J. R., Bartlein, P. J., Daniau, A.-L., Harrison, S. P., Maezumi, S.
Y., Power, M. J., Tinner, W., and Vanniere, B.: Global biomass burning: a
synthesis and review of Holocene paleofire records and their controls,
Quaternary Sci. Rev., 65, 5–25, 2013.</mixed-citation></ref>
      <ref id="bib1.bib85"><label>85</label><mixed-citation>Martin Calvo, M., Prentice, I. C., and Harrison, S. P.: Climate versus carbon dioxide controls on biomass burning: a model
analysis of the glacial-interglacial contrast, Biogeosciences, 11, 6017–6027, <ext-link xlink:href="http://dx.doi.org/10.5194/bg-11-6017-2014" ext-link-type="DOI">10.5194/bg-11-6017-2014</ext-link>, 2014.</mixed-citation></ref>
      <ref id="bib1.bib86"><label>86</label><mixed-citation>
McConnell, J. R., Edwards, R., Kok, G. L., Flanner, M. G., Zender, C. S.,
Saltzman, E. S., Banta, J. R., Pasteris, D. R., Carter, M. M., and Kahl, J.
D. W.: 20th-century industrial black carbon emissions altered Arctic climate
forcing, Science, 317, 1381–1384, 2007.</mixed-citation></ref>
      <ref id="bib1.bib87"><label>87</label><mixed-citation>
McLauchlan, K. K., Higuera, P. E., Gavin, D. G., Perakis, S. S., Mack, M.
C., Alexander, H., Battles, J., Biondi, F., Buma, B., Colombaroli, D.,
Enders, S. K., Engstrom, D. R., Hu, F. S., Marlon, J. R., Marshall, J.,
McGlone, M., Morris, J. L., Nave, L. E., Shuman, B., Smithwick, E. A. H.,
Urrego, D. H., Wardle, D. A., Williams, C. J., and Williams, J. J.:
Reconstructing Disturbances and Their Biogeochemical Consequences over
Multiple Timescales, BioScience, 64, 105–116, 2014.</mixed-citation></ref>
      <ref id="bib1.bib88"><label>88</label><mixed-citation>
McWethy, D. B., Higuera, P. E., Whitlock, C., Veblen, T. T., Bowman, D. M.
J. S., Cary, G. J., Haberle, S. G., Keane, R. E., Maxwell, B. D., McGlone,
M. S., Perry, G. L. W., Wilmshurst, J. M., Holz, A., and Tepley, A. J.: A
conceptual framework for predicting temperate ecosystem sensitivity to human
impacts on fire regimes, Glob. Ecol. Biogeogr., 22, 900–912,
2013.</mixed-citation></ref>
      <ref id="bib1.bib89"><label>89</label><mixed-citation>
Mooney, S. D., Harrison, S. P., Bartlein, P. J., A.-L., D., Stevenson, J.,
Brownlie, K. C., Buckman, S., Cupper, M., Luly, J., Black, M., Colhoun, E.,
D'Costa, D., Dodson, J., Haberle, S., Hope, G. S., Kershaw, P., Kenyon, C.,
McKenzie, M., and Williams, N.: Late Quaternary fire regimes of Australasia,
Quaternary Sci. Rev., 30, 28–46, 2011.</mixed-citation></ref>
      <ref id="bib1.bib90"><label>90</label><mixed-citation>
Moos, M. T. and Cumming, B. F.: Climate–fire interactions during the
Holocene: a test of the utility of charcoal morphotypes in a sediment core
from the boreal region of north-western Ontario (Canada), Int. J. Wildland Fire, 21, 640–652, 2012.</mixed-citation></ref>
      <ref id="bib1.bib91"><label>91</label><mixed-citation>
Morris, S. E. and Moses, T. A.: Forest fire and the natural soil erosion
regime in the Colorado Front Range, Ann. Assoc. Am. Geogr., 77, 245–254, 1987.</mixed-citation></ref>
      <ref id="bib1.bib92"><label>92</label><mixed-citation>Mouillot, F. and Field, C. B.: Fire history and the global carbon budget: a
1<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula><inline-formula><mml:math display="inline"><mml:mrow><mml:mspace linebreak="nobreak" width="0.125em"/><mml:mo>×</mml:mo></mml:mrow></mml:math></inline-formula> 1<inline-formula><mml:math display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula> fire history reconstruction for the
20th century, Glob. Change Biol., 11, 398–420, 2005.</mixed-citation></ref>
      <ref id="bib1.bib93"><label>93</label><mixed-citation>Mouillot, F., Narasimha, A., Balkanski, Y., Lamarque, J.-F., and Field, C.
B.: Global carbon emissions from biomass burning in the 20th century,
Geophys. Res. Lett., 33, L01801, <ext-link xlink:href="http://dx.doi.org/10.1029/2005GL024707" ext-link-type="DOI">10.1029/2005GL024707</ext-link>, 2006.</mixed-citation></ref>
      <ref id="bib1.bib94"><label>94</label><mixed-citation>
Mouillot, F., Schultz, M. G., Yue, C., Cadule, P., Tansey, K., Ciais, P.,
and Chuvieco, E.: Ten years of global burned area products from spaceborne
remote sensing : a review : analysis of user needs and recommendations for
future developments, Int. J. Appl. Earth Obs., 26, 64–79, 2014.</mixed-citation></ref>
      <ref id="bib1.bib95"><label>95</label><mixed-citation>
Munoz, S. E., Mladenoff, D. J., Schroeder, S., and Williams, J. W.: Defining
the spatial patterns of historical land use associated with the indigenous
societies of eastern North America, J. Biogeogr., 41, 2195–2210,
2014.</mixed-citation></ref>
      <ref id="bib1.bib96"><label>96</label><mixed-citation>
Mustaphi, C. J. C. and Pisaric, M. F. J.: A classification for macroscopic
charcoal morphologies found in Holocene lacustrine sediments, Prog. Phys. Geogr., 38, 734–754, 2014.</mixed-citation></ref>
      <ref id="bib1.bib97"><label>97</label><mixed-citation>
Neumann, F. H., Botha, G. A., and Scott, L.: 18 000 years of grassland
evolution in the summer rainfall region of South Africa: evidence from
Mahwaqa Mountain, KwaZulu-Natal, Veg. Hist. Archaeobot., 23,
665–681, 2014.</mixed-citation></ref>
      <ref id="bib1.bib98"><label>98</label><mixed-citation>
Pechony, O. and Shindell, D. T.: Fire parameterization on a global scale,
J. Geophys. Res., 114, 1–10, 2009.</mixed-citation></ref>
      <ref id="bib1.bib99"><label>99</label><mixed-citation>Pechony, O. and Shindell, D.: Driving forces of global wildfires over the
past millennium and the forthcoming century, P. Natl. Acad. Sci. USA,  107, 19167–19170,
<ext-link xlink:href="http://dx.doi.org/10.1073/pnas.1003669107" ext-link-type="DOI">10.1073/pnas.1003669107</ext-link>, 2010.</mixed-citation></ref>
      <ref id="bib1.bib100"><label>100</label><mixed-citation>
Perry, G. L. W., Wilmshurst, J. M., McGlone, M. S., McWethy, D. B., and
Whitlock, C.: Explaining fire-driven landscape transformation during the
Initial Burning Period of New Zealand's prehistory, Glob. Change Biol.,
18, 1609–1621, 2012.</mixed-citation></ref>
      <ref id="bib1.bib101"><label>101</label><mixed-citation>
Petoukhov, V., Ganopolski, A., Brovkin, V., Claussen, M., Eliseev, A.,
Kubatzki, C., and Rahmstorf, S.: CLIMBER-2: a climate system model of
intermediate complexity, Part I: model description and performance for
present climate, Clim. Dynam., 16, 1–17, 2000.</mixed-citation></ref>
      <ref id="bib1.bib102"><label>102</label><mixed-citation>Pfeiffer, M., Spessa, A., and Kaplan, J. O.: A model for global biomass burning in preindustrial time: LPJ-LMfire (v1.0),
Geosci. Model Dev., 6, 643–685, <ext-link xlink:href="http://dx.doi.org/10.5194/gmd-6-643-2013" ext-link-type="DOI">10.5194/gmd-6-643-2013</ext-link>, 2013.</mixed-citation></ref>
      <ref id="bib1.bib103"><label>103</label><mixed-citation>
Pierce, J., Meyer, G., and Jull, A.: Fire-induced erosion and
millennial-scale climate change in northern ponderosa pine forests, Nature,
432, 87–90, 2004.</mixed-citation></ref>
      <ref id="bib1.bib104"><label>104</label><mixed-citation>
Power, M. J., Marlon, J., Ortiz, N., Bartlein, P. J., Harrison, S. P.,
Mayle, F. E., Ballouche, A., Bradshaw, R. H. W., Carcaillet, C., Cordova,
C., Mooney, S., Moreno, P. I., Prentice, I. C., Thonicke, K., Tinner, W.,
Whitlock, C., Zhang, Y., Zhao, Y., Ali, A. A., Anderson, R. S., Beer, R.,
Behling, H., Briles, C., Brown, K. J., Brunelle, A., Bush, M., Camill, P.,
Chu, G. Q., Clark, J., Colombaroli, D., Connor, S., Daniau, A.-L., Daniels,
M., Dodson, J., Doughty, E., Edwards, M. E., Finsinger, W., Foster, D.,
Frechette, J., Gaillard, M.-J., Gavin, D. G., Gobet, E., Haberle, S.,
Hallett, D. J., Higuera, P., Hope, G., Horn, S., Inoue, J., Kaltenrieder,
P., Kennedy, L., Kong, Z. C., Larsen, C., Long, C. J., Lynch, J., Lynch, E.
A., McGlone, M., Meeks, S., Mensing, S., Meyer, G., Minckley, T., Mohr, J.,
Nelson, D. M., New, J., Newnham, R., Noti, R., Oswald, W., Pierce, J.,
Richard, P. J. H., Rowe, C., Goñi, M. F. S., Shuman, B. N., Takahara,
H., Toney, J., Turney, C., Urrego-Sanchez, D. H., Umbanhowar, C.,
Vandergoes, M., Vanniere, B., Vescovi, E., Walsh, M., Wang, X., Williams,
N., Wilmshurst, J., and Zhang, J. H.: Changes in fire regimes since the Last
Glacial Maximum: An assessment based on a global synthesis and analysis of
charcoal data, Clim. Dynam., 30, 887–907, 2008.</mixed-citation></ref>
      <ref id="bib1.bib105"><label>105</label><mixed-citation>
Power, M. J., Marlon, J. R., Bartlein, P. J., and Harrison, S. P.: Fire
history and the Global Charcoal Database: A new tool for hypothesis testing
and data exploration, Palaeogeogr. Palaeocl.,
291, 52–59, 2010.</mixed-citation></ref>
      <ref id="bib1.bib106"><label>106</label><mixed-citation>
Power, M. J., Mayle, F. E., Bartlein, P. J., Marlon, J. R., Anderson, R. S.,
Behling, H., Brown, K. J., Carcaillet, C., Colombaroli, D., Gavin, D. G.,
Hallett, D. J., Horn, S. P., Kennedy, L. M., Lane, C. S., Long, C. J.,
Moreno, P. I., Paitre, C., Robinson, G., Taylor, Z., and Walsh, M. K.:
Climatic control of the biomass-burning decline in the Americas after AD
1500, Holocene, 23, 3–13, 2012.</mixed-citation></ref>
      <ref id="bib1.bib107"><label>107</label><mixed-citation>
Quintana-Krupinski, N., Marlon, J. R., Nishri, A., Street, J. H., and
Paytan, A.: Climatic and human controls on the late Holocene fire history of
northern Israel, Quaternary Res., 80, 396–405, 2013.</mixed-citation></ref>
      <ref id="bib1.bib108"><label>108</label><mixed-citation>
R Development Core Team: R: A language and environment for statistical
computing, Vienna, Austria, 2013.</mixed-citation></ref>
      <ref id="bib1.bib109"><label>109</label><mixed-citation>
Raddatz, T., Reick, C., Knorr, W., Kattge, J., Roeckner, E., Schnur, R.,
Schnitzler, K.-G., Wetzel, P., and Jungclaus, J.: Will the tropical land
biosphere dominate the climate–carbon cycle feedback during the
twenty-first century?, Clim. Dynam., 29, 565–574, 2007.</mixed-citation></ref>
      <ref id="bib1.bib110"><label>110</label><mixed-citation>
Randerson, J. T., Liu, H., Flanner, M. G., Chambers, S. D., Jin, Y., Hess,
P. G., Pfister, G., Mack, M. C., Treseder, K. K., Welp, L. R., Chapin, F.
S., Harden, J. W., Goulden, M. L., Lyons, E., Neff, J. C., Schuur, E. A. G.,
and Zender, C. S.: The impact of boreal forest fire on climate warming,
Science, 314, 1130–1132, 2006.</mixed-citation></ref>
      <ref id="bib1.bib111"><label>111</label><mixed-citation>
Reick, C., Raddatz, T., Brovkin, V., and Gayler, V.: Representation of
natural and anthropogenic land cover change in MPI ESM, Journal of Advances
in Modeling Earth Systems, 5, 459–482, 2013.</mixed-citation></ref>
      <ref id="bib1.bib112"><label>112</label><mixed-citation>
Saleh, R., Robinson, E. S., Tkacik, D. S., Ahern, A. T., Liu, S., Aiken, A.
C., Sullivan, R. C., Presto, A. A., Dubey, M. K., Yokelson, R. J., Donahue,
N. M., and Robinson, A. L.: Brownness of organics in aerosols from biomass
burning linked to their black carbon content, Nat. Geosci., 7,
647–650, 2014.</mixed-citation></ref>
      <ref id="bib1.bib113"><label>113</label><mixed-citation>
Savarino, J. and Legrand, M.: High northern latitude forest fires and
vegetation emissions over the last millennium inferred from the chemistry of
a central Greenlabd ice core, J. Geophys. Res., 103,
8267–8279, 1998.</mixed-citation></ref>
      <ref id="bib1.bib114"><label>114</label><mixed-citation>Schneck, R., Reick, C. H., and Raddatz, T.: Land contribution to natural
CO<inline-formula><mml:math display="inline"><mml:msub><mml:mi/><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:math></inline-formula>
variability on time scales of centuries, Journal of Advances in Modeling
Earth Systems, 5, 354–365, 2013.</mixed-citation></ref>
      <ref id="bib1.bib115"><label>115</label><mixed-citation>
Shakesby, R. A. and Doerr, S. H.: Wildfire as a hydrological and
geomorphological agent, Earth-Sci. Rev., 74, 269–307, 2006.</mixed-citation></ref>
      <ref id="bib1.bib116"><label>116</label><mixed-citation>
Swain, A. M.: A History of Fire and Vegetation in Northeastern Minnesota as
Recorded in Lake Sediments, Quaternary Res., 3, 383–396, 1973.</mixed-citation></ref>
      <ref id="bib1.bib117"><label>117</label><mixed-citation>
Tan, Z. and Huang, C. C.: Holocene wildfire history in loess tableland in
the middle reaches of the Yellow River of China, Holocene, 23,
1466–1476, 2013.</mixed-citation></ref>
      <ref id="bib1.bib118"><label>118</label><mixed-citation>
Thevenon, F. and Anselmetti, F. S.: Charcoal and fly-ash particles from Lake
Lucerne sediments (Central Switzerland) characterized by image analysis:
anthropologic, stratigraphic and environmental implications, Quaternary Sci. Rev., 26, 2631–2643, 2007.</mixed-citation></ref>
      <ref id="bib1.bib119"><label>119</label><mixed-citation>Thonicke, K., Spessa, A., Prentice, I. C., Harrison, S. P., Dong, L., and Carmona-Moreno, C.: The influence of vegetation,
fire spread and fire behaviour on biomass burning and trace gas emissions: results from a process-based model, Biogeosciences, 7,
1991–2011, <ext-link xlink:href="http://dx.doi.org/10.5194/bg-7-1991-2010" ext-link-type="DOI">10.5194/bg-7-1991-2010</ext-link>, 2010.</mixed-citation></ref>
      <ref id="bib1.bib120"><label>120</label><mixed-citation>
Tinner, W., Hofstetter, S., Zeugin, F., Conedera, M., Wohlgemuth, T.,
Zimmermann, L., and Zweifel, R.: Long-distance transport of macroscopic
charcoal by an intensive crown fire in the Swiss Alps - implications for
fire history reconstruction, Holocene, 16, 287–292, 2006.</mixed-citation></ref>
      <ref id="bib1.bib121"><label>121</label><mixed-citation>
Tweiten, M. A., Hotchkiss, S. C., Booth, R. K., Calcote, R. R., and Lynch,
E. A.: The response of a jack pine forest to late-Holocene climate
variability in northwestern Wisconsin, Holocene, 19, 1049–1061, 2009.</mixed-citation></ref>
      <ref id="bib1.bib122"><label>122</label><mixed-citation>van der Werf, G. R., Randerson, J. T., Giglio, L., Collatz, G. J., Kasibhatla, P. S., and Arellano Jr., A. F.:
Interannual variability in global biomass burning emissions from 1997 to 2004, Atmos. Chem. Phys., 6, 3423–3441, <ext-link xlink:href="http://dx.doi.org/10.5194/acp-6-3423-2006" ext-link-type="DOI">10.5194/acp-6-3423-2006</ext-link>, 2006.</mixed-citation></ref>
      <ref id="bib1.bib123"><label>123</label><mixed-citation>van der Werf, G. R., Randerson, J. T., Giglio, L., Collatz, G. J., Mu, M., Kasibhatla, P. S., Morton, D. C.,
DeFries, R. S., Jin, Y., and van Leeuwen, T. T.: Global fire emissions and the contribution of deforestation, savanna,
forest, agricultural, and peat fires (1997–2009), Atmos. Chem. Phys., 10, 11707–11735, <ext-link xlink:href="http://dx.doi.org/10.5194/acp-10-11707-2010" ext-link-type="DOI">10.5194/acp-10-11707-2010</ext-link>, 2010.</mixed-citation></ref>
      <ref id="bib1.bib124"><label>124</label><mixed-citation>
Vanniere, B., Colombaroli, D., Chapron, E., Leroux, A., Tinner, W., and
Magny, M.: Climate versus human-driven fire regimes in Mediterranean
landscapes: the Holocene record of Lago dell'Accesa (Tuscany, Italy),
Quaternary Sci. Rev., 27, 1181–1196, 2008.</mixed-citation></ref>
      <ref id="bib1.bib125"><label>125</label><mixed-citation>
Vanniere, B., Power, M. J., Roberts, N., Tinner, W., Carrión, J., Magny,
M., Bartlein, P., Colombaroli, D., Daniau, A. L., Finsinger, W., Gil-Romera,
G., Kaltenrieder, P., Pini, R., Sadori, L., Turner, R., Valsecchi, V., and
Vescovi, E.: Circum-Mediterranean fire activity and climate changes during
the mid-Holocene environmental transition (8500–2500 cal. BP),  Holocene,
21, 53–73, 2011.</mixed-citation></ref>
      <ref id="bib1.bib126"><label>126</label><mixed-citation>
Verardo, D. J., Froelich, P. N., and McIntyre, A.: Determination of organic
carbon and nitrogen in marine sediments using the Carlo Erba NA-1500
Analyzer, Deep-Sea Res. Pt. I, 37,
157–165, 1990.</mixed-citation></ref>
      <ref id="bib1.bib127"><label>127</label><mixed-citation>
Walsh, M. K., Whitlock, C., and Bartlein, P. J.: A 14 300-year-long record
of fire-vegetation-climate linkages at Battle Ground Lake, southwestern
Washington, Quaternary Res., 70, 251–264, 2008.</mixed-citation></ref>
      <ref id="bib1.bib128"><label>128</label><mixed-citation>
Walsh, M. K., Marlon, J. R., Goring, S. J., Brown, K. J., and Gavin, D. G.:
A Regional Perspective on Holocene Fire–Climate–Human Interactions in the
Pacific Northwest of North America, Ann. Assoc. Am. Geogr., 105, 1135–1157, 2015.</mixed-citation></ref>
      <ref id="bib1.bib129"><label>129</label><mixed-citation>
Wang, Z., Chappellaz, J., Park, K., and Mak, J. E.: Large variations in
southern hemisphere biomass burning during the last 650 years, Science, 330,
1663–1666, 2010.</mixed-citation></ref>
      <ref id="bib1.bib130"><label>130</label><mixed-citation>Ward, D. S., Kloster, S., Mahowald, N. M., Rogers, B. M., Randerson, J. T., and Hess, P. G.:
The changing radiative forcing of fires: global model estimates for past, present and future,
Atmos. Chem. Phys., 12, 10857–10886, <ext-link xlink:href="http://dx.doi.org/10.5194/acp-12-10857-2012" ext-link-type="DOI">10.5194/acp-12-10857-2012</ext-link>, 2012.</mixed-citation></ref>
      <ref id="bib1.bib131"><label>131</label><mixed-citation>
Whitlock, C. and Bartlein, P. J.: Holocene fire activity as a record of past
environmental change, in: Developments in Quaternary Science, edited by: Gillespie, A.
R., Porter, S. C., and Atwater, B. F., Elsevier, Amsterdam, 2004.</mixed-citation></ref>
      <ref id="bib1.bib132"><label>132</label><mixed-citation>
Whitlock, C., Shafer, S. L., and Marlon, J.: The role of climate and
vegetation change in shaping past and future fire regimes in the
northwestern US and the implications for ecosystem management, Forest
Ecol. Manag., 178, 5–21, 2003.</mixed-citation></ref>
      <ref id="bib1.bib133"><label>133</label><mixed-citation>Whitlock, C., Moreno, P. I., and Bartlein, P.: Climatic controls of Holocene
fire patterns in southern South America, Quaternary Res., 68, 28–36,
2007.
 </mixed-citation></ref><?xmltex \hack{\newpage}?>
      <ref id="bib1.bib134"><label>134</label><mixed-citation>
Williams, A. N., Mooney, S. D., Sisson, S. A., and Marlon, J.: Exploring the
relationship between Aboriginal population indices and fire in Australia
over the last 20 000 years, Palaeogeogr. Palaeocl., 432, 49–57, 2015.</mixed-citation></ref>
      <ref id="bib1.bib135"><label>135</label><mixed-citation>
Winkler, M. G.: Charcoal analysis for paleoenvironmental interpretation: a
chemical assay, Quaternary Res., 23, 313–326, 1985.</mixed-citation></ref>
      <ref id="bib1.bib136"><label>136</label><mixed-citation>
Wooller, M. J., Street-Perrott, F. A., and Agnew, A. D. Q.: Late Quaternary
fires and grassland palaeoecology of Mount Kenya, East Africa: evidence from
charred grass cuticles in lake sediments, Palaeogeogr. Palaeocl., 164, 207–230, 2000.</mixed-citation></ref>
      <ref id="bib1.bib137"><label>137</label><mixed-citation>Yue, C., Ciais, P., Cadule, P., Thonicke, K., Archibald, S., Poulter, B., Hao, W. M., Hantson, S.,
Mouillot, F., Friedlingstein, P., Maignan, F., and Viovy, N.: Modelling the role of fires in the terrestrial
carbon balance by incorporating SPITFIRE into the global vegetation model ORCHIDEE – Part 1: simulating historical global burned
area and fire regimes, Geosci. Model Dev., 7, 2747–2767, <ext-link xlink:href="http://dx.doi.org/10.5194/gmd-7-2747-2014" ext-link-type="DOI">10.5194/gmd-7-2747-2014</ext-link>, 2014.</mixed-citation></ref>
      <ref id="bib1.bib138"><label>138</label><mixed-citation>
Zdanowicz, C. M., Zielinski, G. A., and Germani, M. S.: Mount Mazama
eruption: Calendrical age verified and atmospheric impact assessed, Geology,
27, 621–624, 1999.</mixed-citation></ref>
      <ref id="bib1.bib139"><label>139</label><mixed-citation>Zennaro, P., Kehrwald, N., McConnell, J. R., Schüpbach, S., Maselli, O. J., Marlon, J., Vallelonga, P.,
Leuenberger, D., Zangrando, R., Spolaor, A., Borrotti, M., Barbaro, E., Gambaro, A., and Barbante, C.:
Fire in ice: two millennia of boreal forest fire history from the Greenland NEEM ice core, Clim. Past, 10, 1905–1924, <ext-link xlink:href="http://dx.doi.org/10.5194/cp-10-1905-2014" ext-link-type="DOI">10.5194/cp-10-1905-2014</ext-link>, 2014.</mixed-citation></ref>

  </ref-list><app-group content-type="float"><app><title/>

    </app></app-group></back>
    <!--<article-title-html>Reconstructions of biomass burning from sediment-charcoal records to improve
data–model comparisons</article-title-html>
<abstract-html><p class="p">The location, timing, spatial extent, and frequency of wildfires are
changing rapidly in many parts of the world, producing substantial impacts
on ecosystems, people, and potentially climate. Paleofire records based on
charcoal accumulation in sediments enable modern changes in biomass burning
to be considered in their long-term context. Paleofire records also provide
insights into the causes and impacts of past wildfires and emissions when
analyzed in conjunction with other paleoenvironmental data and with fire
models. Here we present new 1000-year and 22 000-year trends and gridded
biomass burning reconstructions based on the Global Charcoal Database
version 3 (GCDv3), which includes 736 charcoal records (57 more than in
version 2). The new gridded reconstructions reveal the spatial patterns
underlying the temporal trends in the data, allowing insights into likely
controls on biomass burning at regional to global scales. In the most recent
few decades, biomass burning has sharply increased in both hemispheres but
especially in the north, where charcoal fluxes are now higher than at any
other time during the past 22 000 years. We also discuss methodological
issues relevant to data–model comparisons and identify areas for future
research. Spatially gridded versions of the global data set from GCDv3 are
provided to facilitate comparison with and validation of global fire
simulations.</p></abstract-html>
<ref-html id="bib1.bib1"><label>1</label><mixed-citation>
Aleman, J. C., Blarquez, O., Bentaleb, I., Bonté, P., Brossier, B., Carcaillet, C., Gond, V.,
Gourlet-Fleury, S., Kpolita, A., Lefèvre, I., Oslisly, R., Power, M. J.,
Yongo, O., Bremond, L., and Favier, C.: Tracking land-cover changes with
sedimentary charcoal in the Afrotropics, Holocene, 23, 1853–1862, 2013.
</mixed-citation></ref-html>
<ref-html id="bib1.bib2"><label>2</label><mixed-citation>
Arora, V. K. and Boer, G. J.: Fire as an interactive component of dynamic
vegetation models, J. Geophys. Res., 110, 1–20, 2005.
</mixed-citation></ref-html>
<ref-html id="bib1.bib3"><label>3</label><mixed-citation>
Bacon, C. R.: Eruptive history of Mount Mazama and Crater Lake Caldera,
Cascade Range, USA, J. Volcanol. Geoth. Res., 18,
57–117, 1983.
</mixed-citation></ref-html>
<ref-html id="bib1.bib4"><label>4</label><mixed-citation>
Bartlein, P., Hostetler, S. W., Shafer, S. L., Holman, J. O., and Solomon,
A. M.: Temporal and spatial structure in a daily wildfire-start data set
from the western United States (1986–96), Int. J. Wildland Fire, 17, 8–17, 2008.
</mixed-citation></ref-html>
<ref-html id="bib1.bib5"><label>5</label><mixed-citation>
Behling, H.: Late Quaternary environmental changes in the Lagoa da
Curuça region (eastern Amazonia, Brazil) and evidence of Podocarpus in
the Amazon lowland, Veg. Hist. Archaeobot., 10, 175–183,
2001.
</mixed-citation></ref-html>
<ref-html id="bib1.bib6"><label>6</label><mixed-citation>
Bistinas, I., Oom, D., Saì, A. C. L., Harrison, S. P., Prentice, I. C., and
Pereira, J. M. C.: Relationships between human population density and burned
area at continental and global scales, PLoS ONE, 8, e81188, <a href="http://dx.doi.org/10.1371/journal.pone.0081188" target="_blank">doi:10.1371/journal.pone.0081188</a>, 2013.
</mixed-citation></ref-html>
<ref-html id="bib1.bib7"><label>7</label><mixed-citation>
Blaauw, M. and Christen, J. A.: Flexible paleoclimate age-depth models using
an autoregressive gamma process, Bayesian Analysis, 6, 457–474, 2011.
</mixed-citation></ref-html>
<ref-html id="bib1.bib8"><label>8</label><mixed-citation>
Blarquez, O., Vannière, B., Marlon, J. R., Daniau, A.-L., Power, M. J.,
Brewer, S., and Bartlein, P. J.: paleofire: An R package to analyse
sedimentary charcoal records from the Global Charcoal Database to
reconstruct past biomass burning, Comput. Geosci., 72, 255–261,
2014.
</mixed-citation></ref-html>
<ref-html id="bib1.bib9"><label>9</label><mixed-citation>
Bond, W. J. and Keeley, J. E.: Fire as a global 'herbivore': the ecology and
evolution of flammable ecosystems, Trends Ecol. Evol., 20,
387–394, 2005.
</mixed-citation></ref-html>
<ref-html id="bib1.bib10"><label>10</label><mixed-citation>
Brovkin, V., Raddatz, T., Reick, C. H., Claussen, M., and Gayler, V.: Global
biogeophysical interactions between forest and climate, Geophys. Res. Lett., 36, L07405, <a href="http://dx.doi.org/10.1029/2009GL037543" target="_blank">doi:10.1029/2009GL037543</a>, 2009.
</mixed-citation></ref-html>
<ref-html id="bib1.bib11"><label>11</label><mixed-citation>
Brown, K. J., Clark, J. S., Grimm, E. C., Donovan, J. J., Mueller, P. G.,
Hansen, B. C. S., and Stefanova, I.: Fire cycles in North American interior
grasslands and their relation to prairie drought, P. Natl. Acad. Sci. USA, 102,
8865–8871, 2005.
</mixed-citation></ref-html>
<ref-html id="bib1.bib12"><label>12</label><mixed-citation>
Brücher, T., Brovkin, V., Kloster, S., Marlon, J. R., and Power, M. J.: Comparing modelled fire dynamics
with charcoal records for the Holocene, Clim. Past, 10, 811–824, <a href="http://dx.doi.org/10.5194/cp-10-811-2014" target="_blank">doi:10.5194/cp-10-811-2014</a>, 2014.
</mixed-citation></ref-html>
<ref-html id="bib1.bib13"><label>13</label><mixed-citation>
Burney, D. A.: Late Quaternary stratigraphic charcoal records from
Madagascar, Quaternary Res., 28, 274–280, 1987.
</mixed-citation></ref-html>
<ref-html id="bib1.bib14"><label>14</label><mixed-citation>
Carcaillet, C., Bergeron, Y., Richard, P. J. H., Fréchette, B.,
Gauthier, S., and Prairie, Y. T.: Change of fire frequency in the eastern
Canadian boreal forests during the Holocene: does vegetation composition or
climate trigger the fire regime?, J. Ecol., 89, 930–946, 2001a.
</mixed-citation></ref-html>
<ref-html id="bib1.bib15"><label>15</label><mixed-citation>
Carcaillet, C., Bouvier, M., Fréchette, B., Larouche, A. C., and
Richard, P. J. H.: Comparison of pollen-slide and sieving methods in
lacustrine charcoal analyses for local and regional fire history, Holocene,
11, 467–476, 2001b.
</mixed-citation></ref-html>
<ref-html id="bib1.bib16"><label>16</label><mixed-citation>
Carcaillet, C., Almquist, H., Asnong, H., Bradshaw, R. H. W., Carrión,
J. S., Gaillard, M. J., Gajewski, K., Haas, J. N., Haberle, S. G., Hadorn,
P., Müller, S. D., Richard, P. J. H., Richoz, I., Rösch, M.,
Sánchez Goñi, M. F., Von Stedingk, H., Stevenson, A. C., Talon, B.,
Tardy, C., Tinner, W., Tryterud, E., Wick, L., and Willis, K. J.: Holocene
biomass burning and global dynamics of the carbon cycle, Chemosphere, 49,
845–863, 2002.
</mixed-citation></ref-html>
<ref-html id="bib1.bib17"><label>17</label><mixed-citation>
Clark, J. S.: Particle motion and the theory of charcoal analysis: source
area, transport, deposition, and sampling, Quaternary Res., 30, 67–80,
1988.
</mixed-citation></ref-html>
<ref-html id="bib1.bib18"><label>18</label><mixed-citation>
Clark, J. S. and Royall, P. D.: Local and regional sediment charcoal evidence for fire regimes in presettlement north-eastern North America,
J. Ecol., 84, 365–382, 1996.
</mixed-citation></ref-html>
<ref-html id="bib1.bib19"><label>19</label><mixed-citation>
Clark, J. S. and Patterson, W. A.: Background and local charcoal in
sediments: scales of fire evidence in the paleorecord, in: Sediment records
of biomass burning and global change, edited by: Clark, J. S., Cachier, H., Goldammer,
J. G., and Stocks, B. J., Springer-Verlag, Berlin, 1997.
</mixed-citation></ref-html>
<ref-html id="bib1.bib20"><label>20</label><mixed-citation>
Colombaroli, D., Tinner, W., Leeuwen, J. v., Noti, R., Vescovi, E.,
Vanniere, B., Magny, M., Schmidt, R., and Bugmann, H.: Response of
broadleaved evergreen Mediterranean forest vegetation to fire disturbance
during the Holocene: insights from the peri-Adriatic region, J. Biogeogr., 36, 314–326, 2009.
</mixed-citation></ref-html>
<ref-html id="bib1.bib21"><label>21</label><mixed-citation>
Colombaroli, D., Vanniere, B., Emmanuel, C., Magny, M., and Tinner, W.:
Fire-vegetation interactions during the Mesolithic-Neolithic transition at
Lago dell'Accesa, Tuscany, Italy, Holocene, 18, 679–692, 2008.
</mixed-citation></ref-html>
<ref-html id="bib1.bib22"><label>22</label><mixed-citation>
Colombaroli, D., Beckmann, M., van der Knaap, W. O., Curdy, P., and Tinner,
W.: Changes in biodiversity and vegetation composition in the central Swiss
Alps during the transition from pristine forest to first farming, Divers. Distrib.,  19,  157–170,
<a href="http://dx.doi.org/10.1111/j.1472-4642.2012.00930.x" target="_blank">doi:10.1111/j.1472-4642.2012.00930.x</a>, 2013.
</mixed-citation></ref-html>
<ref-html id="bib1.bib23"><label>23</label><mixed-citation>
Conedera, M. and Tinner, W.: Long-term fire ecology of Switzerland, Journal
forestier suisse, 161, 424–432, 2010.
</mixed-citation></ref-html>
<ref-html id="bib1.bib24"><label>24</label><mixed-citation>
Conedera, M., Tinner, W., Neff, C., Meurer, M., Dickens, A. F., and Krebs,
P.: Reconstructing past fire regimes: methods, applications, and relevance
to fire management and conservation, Quaternary Sci. Rev., 28,
555–576, 2009.
</mixed-citation></ref-html>
<ref-html id="bib1.bib25"><label>25</label><mixed-citation>
Cordeiro, R. C., Turcq, B. J., Moreira, L. S., de Aragão Rodrigues, R.,
Filho, F. F. L. S., Martins, G. S., Santos, A. B., Barbosa, M., da
Conceição, M. C. G., de Carvalho Rodrigues, R., Evangelista, H.,
Moreira-Turcq, P. F., Penido, Y. P., Sifeddine, A., and Seoane, J. C. S.:
Palaeofires in Amazon: Interplay between Land Use Change and Palaeoclimatic
Events, Palaeogeogr. Palaeocl., 415, 137–151, <a href="http://dx.doi.org/10.1016/j.palaeo.2014.07.020" target="_blank">doi:10.1016/j.palaeo.2014.07.020</a>, 2014.
</mixed-citation></ref-html>
<ref-html id="bib1.bib26"><label>26</label><mixed-citation>
Coughlan, M. R. and Petty, A. M.: Linking humans and fire: a proposal for a transdisciplinary fire ecology,
Int. J. Wildland Fire, 21, 477–487, 2012.
</mixed-citation></ref-html>
<ref-html id="bib1.bib27"><label>27</label><mixed-citation>
Courtney Mustaphi, C. J. and Pisaric, M. F. J.: Holocene
climate–fire–vegetation interactions at a subalpine watershed in
southeastern British Columbia, Canada, Quaternary Res., 81, 228–239,
2014.
</mixed-citation></ref-html>
<ref-html id="bib1.bib28"><label>28</label><mixed-citation>
Cyr, D., Gauthier, S., Bergeron, Y., and Carcaillet, C.: Forest management
is driving the eastern North American boreal forest outside its natural
range of variability, Front. Ecol. Environ., 7, 519–524,
2009.
</mixed-citation></ref-html>
<ref-html id="bib1.bib29"><label>29</label><mixed-citation>
Daniau, A.-L., Harrison, S. P., and Bartlein, P. J.: Fire regimes during the
last glacial, Quaternary Sci. Rev., 29, 2918–2930, 2010.
</mixed-citation></ref-html>
<ref-html id="bib1.bib30"><label>30</label><mixed-citation>
Daniau, A.-L., Goñi, M. F. S., Martinez, P., Urrego, D. H.,
Bout-Roumazeilles, V., Desprat, S., and Marlon, J. R.: Orbital-scale climate
forcing of grassland burning in southern Africa, P. Natl. Acad. Sci. USA, 110, 5069–5073, 2013.
</mixed-citation></ref-html>
<ref-html id="bib1.bib31"><label>31</label><mixed-citation>
Daniau, A. L., Bartlein, P., Harrison, S., Prentice, I., Brewer, S.,
Friedlingstein, P., Harrisson Prentice, T., Inoue, J., Izumi, K., and
Marlon, J.: Predictability of biomass burning in response to climate
changes, Global Biogeochem. Cy., 26, GB4007, <a href="http://dx.doi.org/10.1029/2011GB004249" target="_blank">doi:10.1029/2011GB004249</a>, 2012.
</mixed-citation></ref-html>
<ref-html id="bib1.bib32"><label>32</label><mixed-citation>
DeBano, L. F.: The role of fire and soil heating on water repellency in
wildland environments: a review, J. Hydrol., 231, 195–206, 2000.
</mixed-citation></ref-html>
<ref-html id="bib1.bib33"><label>33</label><mixed-citation>
Delcourt, P. A., Delcourt, H. R., Ison, C. R., Sharp, W. E., and Gremillion,
K. J.: Prehistoric human use of fire, the Eastern Agricultural Complex, and
Appalachian oak-chestnut forests: Paleoecology of Cliff Palace Pond,
Kentucky, Am. Antiquity, 63, 263–278, 1998.
</mixed-citation></ref-html>
<ref-html id="bib1.bib34"><label>34</label><mixed-citation>
Dennison, P. E., Brewer, S. C., Arnold, J. D., and Moritz, M. A.: Large
wildfire trends in the western United States, 1984–2011, Geophys. Res. Lett., 41, 2928–2933, 2014.
</mixed-citation></ref-html>
<ref-html id="bib1.bib35"><label>35</label><mixed-citation>
Dunnette, P. V., Higuera, P. E., McLauchlan, K. K., Derr, K. M., Briles, C.
E., and Keefe, M. H.: Biogeochemical impacts of wildfires over four
millennia in a Rocky Mountain subalpine watershed, New Phytol., 203,
900–912, 2014.
</mixed-citation></ref-html>
<ref-html id="bib1.bib36"><label>36</label><mixed-citation>
Easterbrook, S. M.: Open code for open science?, Nat. Geosci., 7, 779–781,
2014.
</mixed-citation></ref-html>
<ref-html id="bib1.bib37"><label>37</label><mixed-citation>
Edwards, M., Franklin-Smith, L., Clarke, C., Baker, J., Hill, S., and
Gallagher, K.: The role of fire in the mid-Holocene arrival and expansion of
lodgepole pine (Pinus contorta var. latifolia Engelm. ex S. Watson) in
Yukon, Canada, Holocene, 25, 64–78, 2015.
</mixed-citation></ref-html>
<ref-html id="bib1.bib38"><label>38</label><mixed-citation>
Enache, M. D. and Cumming, B. F.: Tracking recorded fires using charcoal
morphology from the sedimentary sequence of Prosser Lake, British Columbia
(Canada), Quaternary Res., 65, 282–292, 2006.
</mixed-citation></ref-html>
<ref-html id="bib1.bib39"><label>39</label><mixed-citation>
Falk, D. A., Heyerdahl, E. K., Brown, P. M., Farris, C., Fulé, P. Z.,
McKenzie, D., Swetnam, T. W., Taylor, A. H., and Van Horne, M. L.:
Multi-scale controls of historical forest-fire regimes: new insights from
fire-scar networks, Front. Ecol. Environ., 9, 446–454,
2011.
</mixed-citation></ref-html>
<ref-html id="bib1.bib40"><label>40</label><mixed-citation>
Fischer, H., Behrens, M., Bock, M., Richter, U., Schmitt, J., Loulergue, L.,
Chappellaz, J., Spahni, R., Blunier, T., Leuenberger, M., and Stocker, T.
F.: Changing boreal methane sources and constant biomass burning during the
last termination, Nature, 452,  864–867, <a href="http://dx.doi.org/10.1038/nature06825" target="_blank">doi:10.1038/nature06825</a>, 2008.
</mixed-citation></ref-html>
<ref-html id="bib1.bib41"><label>41</label><mixed-citation>
Fuller, J. L., Foster, D. R., McLachlan, J. S., and Drake, N.: Impact of
human activity on regional forest composition and dynamics in central New
England, Ecosystems, 1, 76–95, 1998.
</mixed-citation></ref-html>
<ref-html id="bib1.bib42"><label>42</label><mixed-citation>
Ganopolski, A., Petoukhov, V., Rahmstorf, S., Brovkin, V., Claussen, M.,
Eliseev, A., and Kubatzki, C.: CLIMBER-2: a climate system model of
intermediate complexity, Part II: model sensitivity, Clim. Dynam., 17,
735–751, 2001.
</mixed-citation></ref-html>
<ref-html id="bib1.bib43"><label>43</label><mixed-citation>
Gavin, D. G., Hu, F. S., Lertzman, K., and Corbett, P.: Weak climatic
control of stand-scale fire history during the late Holocene, Ecology, 87,
1722–1732, 2006.
</mixed-citation></ref-html>
<ref-html id="bib1.bib44"><label>44</label><mixed-citation>
Gavin, D. G., Hallett, D. J., Hu, F. S., Lertzman, K. P., Prichard, S. J.,
Brown, K. J., Lynch, J. A., Bartlein, P., and Peterson, D. L.: Forest fire
and climate change in western North America: insights from sediment charcoal
records, Front. Ecol. Environ., 5, 499–506, 2007.
</mixed-citation></ref-html>
<ref-html id="bib1.bib45"><label>45</label><mixed-citation>
Giglio, L., Randerson, J. T., and van der Werf, G. R.: Analysis of daily,
monthly, and annual burned area using the fourth-generation global fire
emissions database (GFED4), J. Geophys. Res.-Biogeo.,
118, 317–328, 2013.
</mixed-citation></ref-html>
<ref-html id="bib1.bib46"><label>46</label><mixed-citation>
Giorgetta, M. A., Jungclaus, J., Reick, C. H., Legutke, S., Bader, J.,
Böttinger, M., Brovkin, V., Crueger, T., Esch, M., and Fieg, K.: Climate
and carbon cycle changes from 1850 to 2100 in MPI ESM simulations for the
Coupled Model Intercomparison Project phase 5, Journal of Advances in Modeling Earth Systems, 5, 572–597, 2013.
</mixed-citation></ref-html>
<ref-html id="bib1.bib47"><label>47</label><mixed-citation>
Girardin, M. P. and Sauchyn, D.: Three centuries of annual area burned
variability in northwestern North America inferred from tree rings,
Holocene, 18, 205–214, 2008.
</mixed-citation></ref-html>
<ref-html id="bib1.bib48"><label>48</label><mixed-citation>
Goring, S., Williams, J. W., Blois, J. L., Jackson, S. T., Paciorek, C.,
Booth, R. K., Marlon, J. R., Blaauw, M., and Andres, C.: Deposition times in
the northeastern United States during the Holocene: establishing valid
priors for Bayesian age models, Quaternary Sci. Rev., 48, 54–60,
2012.
</mixed-citation></ref-html>
<ref-html id="bib1.bib49"><label>49</label><mixed-citation>
Gu, Y. S., Pearsall, D. M., Xie, S. C., and Yu, J. X.: Vegetation and fire
history of a Chinese site in southern tropical Xishuangbanna derived from
phytolith and charcoal records from Holocene sediments, J. Biogeogr., 35, 325–341, 2008.
</mixed-citation></ref-html>
<ref-html id="bib1.bib50"><label>50</label><mixed-citation>
Haberle, S. G.: Late quaternary vegetation change in the Tari Basin, Papua
New Guinea, Palaeogeogr. Palaeocl., 137, 1–24,
1998.
</mixed-citation></ref-html>
<ref-html id="bib1.bib51"><label>51</label><mixed-citation>
Haberle, S. G. and Ledru, M. P.: Correlations among charcoal records of
fires from the past 16 000 years in Indonesia, Papua New Guinea, and Central
and South America, Quaternary Res., 55, 97–104, 2001.
</mixed-citation></ref-html>
<ref-html id="bib1.bib52"><label>52</label><mixed-citation>
Hallett, D., Hills, L. V., and Clague, J. J.: New accelerator mass
spectrometry radiocarbon ages for the Mazama tephra layer from Kootenay
National Park, British Columbia, Canada, Can. J. Earth Sci.,
34, 1202–1209, 1997.
</mixed-citation></ref-html>
<ref-html id="bib1.bib53"><label>53</label><mixed-citation>
Han, Y. M., Marlon, J. R., Cao, J. J., Jin, Z. D., and An, Z. S.: Holocene
biomass burning trends in China from soot, char and charcoal in lake
sediments, Global Biogeochem. Cy., 26,  GB4017, <a href="http://dx.doi.org/10.1029/2011GB004197" target="_blank">doi:10.1029/2011GB004197</a>, 2012.
</mixed-citation></ref-html>
<ref-html id="bib1.bib54"><label>54</label><mixed-citation>
Harley, G. L., Grissino-Mayer, H. D., and Horn, S. P.: Fire history and
forest structure of an endangered subtropical ecosystem in the Florida Keys,
USA, Int. J. Wildland Fire, 104, 1–19, 2012.
</mixed-citation></ref-html>
<ref-html id="bib1.bib55"><label>55</label><mixed-citation>
Harrison, S. P., Marlon, J. R., and Bartlein, P. J.: Fire in the Earth
System, in: Changing Climates, Earth Systems and Society, edited by: Dodson, J.,
Springer, Dordrecht, The Netherlands, 2010.
</mixed-citation></ref-html>
<ref-html id="bib1.bib56"><label>56</label><mixed-citation>
Herridge, V., Birch, S. P., and Law, M.: Open Quaternary: A New, Open Access
J. Quaternary Res., Open Quaternary, 1, <a href="http://dx.doi.org/10.5334/oq.ad" target="_blank">doi:10.5334/oq.ad</a>, 2015.
</mixed-citation></ref-html>
<ref-html id="bib1.bib57"><label>57</label><mixed-citation>
Heusser, C. J.: Three late Quaternary pollen diagrams from southern
Patagonia and their palaeoecological implications, Palaeogeogr. Palaeocl., 118, 1–24, 1995.
</mixed-citation></ref-html>
<ref-html id="bib1.bib58"><label>58</label><mixed-citation>
Higuera, P., Gavin, D., Bartlein, P., and Hallett, D.: Peak detection in
sediment-charcoal records: impacts of alternative data analysis methods on
fire-history interpretations, Int. J. Wildland Fire, 19,
996–1014, 2010.
</mixed-citation></ref-html>
<ref-html id="bib1.bib59"><label>59</label><mixed-citation>
Higuera, P. E., Peters, M. E., Brubaker, L. B., and Gavin, D. G.:
Understanding the origin and analysis of sediment-charcoal records with a
simulation model, Quaternary Sci. Rev., 26, 1790–1809, 2007.
</mixed-citation></ref-html>
<ref-html id="bib1.bib60"><label>60</label><mixed-citation>
Higuera, P. E., Whitlock, C., and Gage, J.: Linking tree-ring and
sediment-charcoal records to reconstruct fire occurrence and area burned in
subalpine forests of Yellowstone National Park, USA,  Holocene 21,
327–341, 2011.
</mixed-citation></ref-html>
<ref-html id="bib1.bib61"><label>61</label><mixed-citation>
Higuera, P. E., Briles, C. E., and Whitlock, C.: Fire-regime complacency and
sensitivity to centennial-through millennial-scale climate change in Rocky
Mountain subalpine forests, Colorado, USA, J. Ecol., 102,
1429–1441, 2014.
</mixed-citation></ref-html>
<ref-html id="bib1.bib62"><label>62</label><mixed-citation>
Iglesias, V. and Whitlock, C.: Fire responses to postglacial climate change
and human impact in northern Patagonia (41–43° S), P. Natl. Acad. Sci. USA, 111, E5545–E5554, 2014.
</mixed-citation></ref-html>
<ref-html id="bib1.bib63"><label>63</label><mixed-citation>
Iglesias, V., Whitlock, C., Bianchi, M. M., Villarosa, G., and Outes, V.:
Holocene climate variability and environmental history at the Patagonian
forest/steppe ecotone: Lago Mosquito (42°29′37.89′′ S, 71°24′14.57′′ W) and Laguna del Condor (42°20′47.22′′ S,
71°17′07.62′′ W), Holocene, 22, 1297–1307, 2012.
</mixed-citation></ref-html>
<ref-html id="bib1.bib64"><label>64</label><mixed-citation>
Jensen, K., Lynch, E. A., Calcote, R., and Hotchkiss, S. C.: Interpretation
of charcoal morphotypes in sediments from Ferry Lake, Wisconsin, USA: do
different plant fuel sources produce distinctive charcoal morphotypes?,
Holocene, 17, 907–915, 2007.
</mixed-citation></ref-html>
<ref-html id="bib1.bib65"><label>65</label><mixed-citation>
Kattge, J., Diaz, S., and Wirth, C.: Of carrots and sticks, Nat. Geosci.,
7, 778–779, 2014.
</mixed-citation></ref-html>
<ref-html id="bib1.bib66"><label>66</label><mixed-citation>
Kehrwald, N., Whitlock, C., Barbante, C., Brovkin, V., Daniau, A.-L.,
Kaplan, J., Marlon, J. R., Power, M. J., Thonicke, K., and Van Der Werf, G.
R.: Recent advancements in wildfire research, Eos Trans. AGU, 421–423, 2013.
</mixed-citation></ref-html>
<ref-html id="bib1.bib67"><label>67</label><mixed-citation>
Kelley, D. I., Harrison, S. P., and Prentice, I. C.: Improved simulation of fire-vegetation interactions in the Land surface Processes and
eXchanges dynamic global vegetation model (LPX-Mv1), Geosci. Model Dev., 7, 2411–2433, <a href="http://dx.doi.org/10.5194/gmd-7-2411-2014" target="_blank">doi:10.5194/gmd-7-2411-2014</a>, 2014.
</mixed-citation></ref-html>
<ref-html id="bib1.bib68"><label>68</label><mixed-citation>
Kelly, R., Chipman, M. L., Higuera, P. E., Stefanova, I., Brubaker, L. B.,
and Hu, F. S.: Recent burning of boreal forests exceeds fire regime limits
of the past 10 000 years, P. Natl. Acad. Sci. USA,
110, 13055–13060, 2013.
</mixed-citation></ref-html>
<ref-html id="bib1.bib69"><label>69</label><mixed-citation>
Keywood, M., Kanakidou, M., Stohl, A., Dentener, F., Grassi, G., Meyer, C.
P., Torseth, K., Edwards, D., Thompson, A. M., Lohmann, U., and Burrows, J.:
Fire in the Air: Biomass Burning Impacts in a Changing Climate, Critical Reviews in Environmental Science and Technology, 43, 40–83, 2013.
</mixed-citation></ref-html>
<ref-html id="bib1.bib70"><label>70</label><mixed-citation>
Kloster, S., Mahowald, N. M., Randerson, J. T., Thornton, P. E., Hoffman, F. M., Levis, S., Lawrence, P. J.,
Feddema, J. J., Oleson, K. W., and Lawrence, D. M.: Fire dynamics during the 20th century simulated by the Community Land
Model, Biogeosciences, 7, 1877–1902, <a href="http://dx.doi.org/10.5194/bg-7-1877-2010" target="_blank">doi:10.5194/bg-7-1877-2010</a>, 2010.
</mixed-citation></ref-html>
<ref-html id="bib1.bib71"><label>71</label><mixed-citation>
Kloster, S., Mahowald, N. M., Randerson, J. T., and Lawrence, P. J.: The impacts of climate, land use, and demography on fires
during the 21st century simulated by CLM-CN, Biogeosciences, 9, 509–525, <a href="http://dx.doi.org/10.5194/bg-9-509-2012" target="_blank">doi:10.5194/bg-9-509-2012</a>, 2012.
</mixed-citation></ref-html>
<ref-html id="bib1.bib72"><label>72</label><mixed-citation>
Kloster, S., Brücher, T., Brovkin, V., and Wilkenskjeld, S.: Controls on fire activity over the Holocene,
Clim. Past, 11, 781–788, <a href="http://dx.doi.org/10.5194/cp-11-781-2015" target="_blank">doi:10.5194/cp-11-781-2015</a>, 2015.
</mixed-citation></ref-html>
<ref-html id="bib1.bib73"><label>73</label><mixed-citation>
Lamarque, J.-F., Bond, T. C., Eyring, V., Granier, C., Heil, A., Klimont, Z., Lee, D., Liousse, C., Mieville, A.,
Owen, B., Schultz, M. G., Shindell, D., Smith, S. J., Stehfest, E., Van Aardenne, J., Cooper, O. R., Kainuma, M.,
Mahowald, N., McConnell, J. R., Naik, V., Riahi, K., and van Vuuren, D. P.: Historical (1850–2000) gridded anthropogenic and
biomass burning emissions of reactive gases and aerosols: methodology and application, Atmos. Chem. Phys., 10, 7017–7039, <a href="http://dx.doi.org/10.5194/acp-10-7017-2010" target="_blank">doi:10.5194/acp-10-7017-2010</a>, 2010.
</mixed-citation></ref-html>
<ref-html id="bib1.bib74"><label>74</label><mixed-citation>
Lasslop, G., Thonicke, K., and Kloster, S.: Spitfire within the mpi earth
system model: Model development and evaluation, Advances in Modeling Earth Systems, 6, 740–755, 2014.
</mixed-citation></ref-html>
<ref-html id="bib1.bib75"><label>75</label><mixed-citation>
Le Page, Y., Morton, D., Bond-Lamberty, B., Pereira, J. M. C., and Hurtt, G.: HESFIRE: a global fire model to explore the
role of anthropogenic and weather drivers, Biogeosciences, 12, 887–903, <a href="http://dx.doi.org/10.5194/bg-12-887-2015" target="_blank">doi:10.5194/bg-12-887-2015</a>, 2015.
</mixed-citation></ref-html>
<ref-html id="bib1.bib76"><label>76</label><mixed-citation>
Lehndorff, E., Wolf, M., Litt, T., Brauer, A., and Amelung, W.: 15 000 years
of black carbon deposition – A post-glacial fire record from maar lake
sediments (Germany), Quaternary Sci. Rev., 110, 15–22, 2015.
</mixed-citation></ref-html>
<ref-html id="bib1.bib77"><label>77</label><mixed-citation>
Li, F., Levis, S., and Ward, D. S.: Quantifying the role of fire in the Earth system – Part 1: Improved global fire modeling in
the Community Earth System Model (CESM1), Biogeosciences, 10, 2293–2314, <a href="http://dx.doi.org/10.5194/bg-10-2293-2013" target="_blank">doi:10.5194/bg-10-2293-2013</a>, 2013.
</mixed-citation></ref-html>
<ref-html id="bib1.bib78"><label>78</label><mixed-citation>
Maezumi, S. Y., Power, M. J., Mayle, F. E., McLauchlan, K. K., and Iriarte, J.: Effects of past climate variability on
fire and vegetation in the cerrãdo savanna of the Huanchaca Mesetta, NE Bolivia, Clim. Past, 11, 835–853, <a href="http://dx.doi.org/10.5194/cp-11-835-2015" target="_blank">doi:10.5194/cp-11-835-2015</a>, 2015.
</mixed-citation></ref-html>
<ref-html id="bib1.bib79"><label>79</label><mixed-citation>
Mann, M. E., Zhang, Z., Rutherford, S., Bradley, R. S., Hughes, M. K.,
Shindell, D., Ammann, C., Faluvegi, G., and Ni, F.: Global signatures and
dynamical origins of the Little Ice Age and Medieval Climate Anomaly,
Science, 326, 1256–1260, 2009.
</mixed-citation></ref-html>
<ref-html id="bib1.bib80"><label>80</label><mixed-citation>
Marlon, J., Bartlein, P. J., and Whitlock, C.: Fire-fuel-climate linkages in
the northwestern USA during the Holocene, Holocene, 16, 1059–1071, 2006.
</mixed-citation></ref-html>
<ref-html id="bib1.bib81"><label>81</label><mixed-citation>
Marlon, J., Bartlein, P., Carcaillet, C., Gavin, D. G., Harrison, S. P.,
Higuera, P. E., Joos, F., Power, M. J., and Prentice, C. I.: Climate and
human influences on global biomass burning over the past two millennia,
Nat. Geosci., 1, 697–701, 2008.
</mixed-citation></ref-html>
<ref-html id="bib1.bib82"><label>82</label><mixed-citation>
Marlon, J., Bartlein, P., Walsh, M. K., Harrison, S. P., Brown, K. J.,
Edwards, M. E., Higuera, P. E., Power, M. J., Anderson, R. S., Briles, C.
E., Brunelle, A., Carcaillet, C., Daniels, M., Hu, F. S., Lavoie, M., Long,
C. J., Minckley, T., Richard, P. J. H., Scott, A. C., Shafer, D. S., Tinner,
W., Umbanhowar Jr, C. E., and Whitlock, C.: Wildfire responses to abrupt
climate change in North America, P. Natl. Acad. Sci. USA, 106, 2519–2524, 2009.
</mixed-citation></ref-html>
<ref-html id="bib1.bib83"><label>83</label><mixed-citation>
Marlon, J. R., Bartlein, P. J., Gavin, D. G., Long, C. J., Anderson, R. S.,
Briles, C. E., Brown, K. J., Colombaroli, D., Hallett, D. J., Power, M. J.,
Scharf, E. A., and Walsh, M. K.: Long-term perspective on wildfires in the
western USA, P. Natl. Acad. Sci. USA, 109,
E535–E543, 2012.
</mixed-citation></ref-html>
<ref-html id="bib1.bib84"><label>84</label><mixed-citation>
Marlon, J. R., Bartlein, P. J., Daniau, A.-L., Harrison, S. P., Maezumi, S.
Y., Power, M. J., Tinner, W., and Vanniere, B.: Global biomass burning: a
synthesis and review of Holocene paleofire records and their controls,
Quaternary Sci. Rev., 65, 5–25, 2013.
</mixed-citation></ref-html>
<ref-html id="bib1.bib85"><label>85</label><mixed-citation>
Martin Calvo, M., Prentice, I. C., and Harrison, S. P.: Climate versus carbon dioxide controls on biomass burning: a model
analysis of the glacial-interglacial contrast, Biogeosciences, 11, 6017–6027, <a href="http://dx.doi.org/10.5194/bg-11-6017-2014" target="_blank">doi:10.5194/bg-11-6017-2014</a>, 2014.
</mixed-citation></ref-html>
<ref-html id="bib1.bib86"><label>86</label><mixed-citation>
McConnell, J. R., Edwards, R., Kok, G. L., Flanner, M. G., Zender, C. S.,
Saltzman, E. S., Banta, J. R., Pasteris, D. R., Carter, M. M., and Kahl, J.
D. W.: 20th-century industrial black carbon emissions altered Arctic climate
forcing, Science, 317, 1381–1384, 2007.
</mixed-citation></ref-html>
<ref-html id="bib1.bib87"><label>87</label><mixed-citation>
McLauchlan, K. K., Higuera, P. E., Gavin, D. G., Perakis, S. S., Mack, M.
C., Alexander, H., Battles, J., Biondi, F., Buma, B., Colombaroli, D.,
Enders, S. K., Engstrom, D. R., Hu, F. S., Marlon, J. R., Marshall, J.,
McGlone, M., Morris, J. L., Nave, L. E., Shuman, B., Smithwick, E. A. H.,
Urrego, D. H., Wardle, D. A., Williams, C. J., and Williams, J. J.:
Reconstructing Disturbances and Their Biogeochemical Consequences over
Multiple Timescales, BioScience, 64, 105–116, 2014.
</mixed-citation></ref-html>
<ref-html id="bib1.bib88"><label>88</label><mixed-citation>
McWethy, D. B., Higuera, P. E., Whitlock, C., Veblen, T. T., Bowman, D. M.
J. S., Cary, G. J., Haberle, S. G., Keane, R. E., Maxwell, B. D., McGlone,
M. S., Perry, G. L. W., Wilmshurst, J. M., Holz, A., and Tepley, A. J.: A
conceptual framework for predicting temperate ecosystem sensitivity to human
impacts on fire regimes, Glob. Ecol. Biogeogr., 22, 900–912,
2013.
</mixed-citation></ref-html>
<ref-html id="bib1.bib89"><label>89</label><mixed-citation>
Mooney, S. D., Harrison, S. P., Bartlein, P. J., A.-L., D., Stevenson, J.,
Brownlie, K. C., Buckman, S., Cupper, M., Luly, J., Black, M., Colhoun, E.,
D'Costa, D., Dodson, J., Haberle, S., Hope, G. S., Kershaw, P., Kenyon, C.,
McKenzie, M., and Williams, N.: Late Quaternary fire regimes of Australasia,
Quaternary Sci. Rev., 30, 28–46, 2011.
</mixed-citation></ref-html>
<ref-html id="bib1.bib90"><label>90</label><mixed-citation>
Moos, M. T. and Cumming, B. F.: Climate–fire interactions during the
Holocene: a test of the utility of charcoal morphotypes in a sediment core
from the boreal region of north-western Ontario (Canada), Int. J. Wildland Fire, 21, 640–652, 2012.
</mixed-citation></ref-html>
<ref-html id="bib1.bib91"><label>91</label><mixed-citation>
Morris, S. E. and Moses, T. A.: Forest fire and the natural soil erosion
regime in the Colorado Front Range, Ann. Assoc. Am. Geogr., 77, 245–254, 1987.
</mixed-citation></ref-html>
<ref-html id="bib1.bib92"><label>92</label><mixed-citation>
Mouillot, F. and Field, C. B.: Fire history and the global carbon budget: a
1°<mspace linebreak="nobreak" width="0.125em"/> ×  1° fire history reconstruction for the
20th century, Glob. Change Biol., 11, 398–420, 2005.
</mixed-citation></ref-html>
<ref-html id="bib1.bib93"><label>93</label><mixed-citation>
Mouillot, F., Narasimha, A., Balkanski, Y., Lamarque, J.-F., and Field, C.
B.: Global carbon emissions from biomass burning in the 20th century,
Geophys. Res. Lett., 33, L01801, <a href="http://dx.doi.org/10.1029/2005GL024707" target="_blank">doi:10.1029/2005GL024707</a>, 2006.
</mixed-citation></ref-html>
<ref-html id="bib1.bib94"><label>94</label><mixed-citation>
Mouillot, F., Schultz, M. G., Yue, C., Cadule, P., Tansey, K., Ciais, P.,
and Chuvieco, E.: Ten years of global burned area products from spaceborne
remote sensing : a review : analysis of user needs and recommendations for
future developments, Int. J. Appl. Earth Obs., 26, 64–79, 2014.
</mixed-citation></ref-html>
<ref-html id="bib1.bib95"><label>95</label><mixed-citation>
Munoz, S. E., Mladenoff, D. J., Schroeder, S., and Williams, J. W.: Defining
the spatial patterns of historical land use associated with the indigenous
societies of eastern North America, J. Biogeogr., 41, 2195–2210,
2014.
</mixed-citation></ref-html>
<ref-html id="bib1.bib96"><label>96</label><mixed-citation>
Mustaphi, C. J. C. and Pisaric, M. F. J.: A classification for macroscopic
charcoal morphologies found in Holocene lacustrine sediments, Prog. Phys. Geogr., 38, 734–754, 2014.
</mixed-citation></ref-html>
<ref-html id="bib1.bib97"><label>97</label><mixed-citation>
Neumann, F. H., Botha, G. A., and Scott, L.: 18 000 years of grassland
evolution in the summer rainfall region of South Africa: evidence from
Mahwaqa Mountain, KwaZulu-Natal, Veg. Hist. Archaeobot., 23,
665–681, 2014.
</mixed-citation></ref-html>
<ref-html id="bib1.bib98"><label>98</label><mixed-citation>
Pechony, O. and Shindell, D. T.: Fire parameterization on a global scale,
J. Geophys. Res., 114, 1–10, 2009.
</mixed-citation></ref-html>
<ref-html id="bib1.bib99"><label>99</label><mixed-citation>
Pechony, O. and Shindell, D.: Driving forces of global wildfires over the
past millennium and the forthcoming century, P. Natl. Acad. Sci. USA,  107, 19167–19170,
<a href="http://dx.doi.org/10.1073/pnas.1003669107" target="_blank">doi:10.1073/pnas.1003669107</a>, 2010.
</mixed-citation></ref-html>
<ref-html id="bib1.bib100"><label>100</label><mixed-citation>
Perry, G. L. W., Wilmshurst, J. M., McGlone, M. S., McWethy, D. B., and
Whitlock, C.: Explaining fire-driven landscape transformation during the
Initial Burning Period of New Zealand's prehistory, Glob. Change Biol.,
18, 1609–1621, 2012.
</mixed-citation></ref-html>
<ref-html id="bib1.bib101"><label>101</label><mixed-citation>
Petoukhov, V., Ganopolski, A., Brovkin, V., Claussen, M., Eliseev, A.,
Kubatzki, C., and Rahmstorf, S.: CLIMBER-2: a climate system model of
intermediate complexity, Part I: model description and performance for
present climate, Clim. Dynam., 16, 1–17, 2000.
</mixed-citation></ref-html>
<ref-html id="bib1.bib102"><label>102</label><mixed-citation>
Pfeiffer, M., Spessa, A., and Kaplan, J. O.: A model for global biomass burning in preindustrial time: LPJ-LMfire (v1.0),
Geosci. Model Dev., 6, 643–685, <a href="http://dx.doi.org/10.5194/gmd-6-643-2013" target="_blank">doi:10.5194/gmd-6-643-2013</a>, 2013.
</mixed-citation></ref-html>
<ref-html id="bib1.bib103"><label>103</label><mixed-citation>
Pierce, J., Meyer, G., and Jull, A.: Fire-induced erosion and
millennial-scale climate change in northern ponderosa pine forests, Nature,
432, 87–90, 2004.
</mixed-citation></ref-html>
<ref-html id="bib1.bib104"><label>104</label><mixed-citation>
Power, M. J., Marlon, J., Ortiz, N., Bartlein, P. J., Harrison, S. P.,
Mayle, F. E., Ballouche, A., Bradshaw, R. H. W., Carcaillet, C., Cordova,
C., Mooney, S., Moreno, P. I., Prentice, I. C., Thonicke, K., Tinner, W.,
Whitlock, C., Zhang, Y., Zhao, Y., Ali, A. A., Anderson, R. S., Beer, R.,
Behling, H., Briles, C., Brown, K. J., Brunelle, A., Bush, M., Camill, P.,
Chu, G. Q., Clark, J., Colombaroli, D., Connor, S., Daniau, A.-L., Daniels,
M., Dodson, J., Doughty, E., Edwards, M. E., Finsinger, W., Foster, D.,
Frechette, J., Gaillard, M.-J., Gavin, D. G., Gobet, E., Haberle, S.,
Hallett, D. J., Higuera, P., Hope, G., Horn, S., Inoue, J., Kaltenrieder,
P., Kennedy, L., Kong, Z. C., Larsen, C., Long, C. J., Lynch, J., Lynch, E.
A., McGlone, M., Meeks, S., Mensing, S., Meyer, G., Minckley, T., Mohr, J.,
Nelson, D. M., New, J., Newnham, R., Noti, R., Oswald, W., Pierce, J.,
Richard, P. J. H., Rowe, C., Goñi, M. F. S., Shuman, B. N., Takahara,
H., Toney, J., Turney, C., Urrego-Sanchez, D. H., Umbanhowar, C.,
Vandergoes, M., Vanniere, B., Vescovi, E., Walsh, M., Wang, X., Williams,
N., Wilmshurst, J., and Zhang, J. H.: Changes in fire regimes since the Last
Glacial Maximum: An assessment based on a global synthesis and analysis of
charcoal data, Clim. Dynam., 30, 887–907, 2008.
</mixed-citation></ref-html>
<ref-html id="bib1.bib105"><label>105</label><mixed-citation>
Power, M. J., Marlon, J. R., Bartlein, P. J., and Harrison, S. P.: Fire
history and the Global Charcoal Database: A new tool for hypothesis testing
and data exploration, Palaeogeogr. Palaeocl.,
291, 52–59, 2010.
</mixed-citation></ref-html>
<ref-html id="bib1.bib106"><label>106</label><mixed-citation>
Power, M. J., Mayle, F. E., Bartlein, P. J., Marlon, J. R., Anderson, R. S.,
Behling, H., Brown, K. J., Carcaillet, C., Colombaroli, D., Gavin, D. G.,
Hallett, D. J., Horn, S. P., Kennedy, L. M., Lane, C. S., Long, C. J.,
Moreno, P. I., Paitre, C., Robinson, G., Taylor, Z., and Walsh, M. K.:
Climatic control of the biomass-burning decline in the Americas after AD
1500, Holocene, 23, 3–13, 2012.
</mixed-citation></ref-html>
<ref-html id="bib1.bib107"><label>107</label><mixed-citation>
Quintana-Krupinski, N., Marlon, J. R., Nishri, A., Street, J. H., and
Paytan, A.: Climatic and human controls on the late Holocene fire history of
northern Israel, Quaternary Res., 80, 396–405, 2013.
</mixed-citation></ref-html>
<ref-html id="bib1.bib108"><label>108</label><mixed-citation>
R Development Core Team: R: A language and environment for statistical
computing, Vienna, Austria, 2013.
</mixed-citation></ref-html>
<ref-html id="bib1.bib109"><label>109</label><mixed-citation>
Raddatz, T., Reick, C., Knorr, W., Kattge, J., Roeckner, E., Schnur, R.,
Schnitzler, K.-G., Wetzel, P., and Jungclaus, J.: Will the tropical land
biosphere dominate the climate–carbon cycle feedback during the
twenty-first century?, Clim. Dynam., 29, 565–574, 2007.
</mixed-citation></ref-html>
<ref-html id="bib1.bib110"><label>110</label><mixed-citation>
Randerson, J. T., Liu, H., Flanner, M. G., Chambers, S. D., Jin, Y., Hess,
P. G., Pfister, G., Mack, M. C., Treseder, K. K., Welp, L. R., Chapin, F.
S., Harden, J. W., Goulden, M. L., Lyons, E., Neff, J. C., Schuur, E. A. G.,
and Zender, C. S.: The impact of boreal forest fire on climate warming,
Science, 314, 1130–1132, 2006.
</mixed-citation></ref-html>
<ref-html id="bib1.bib111"><label>111</label><mixed-citation>
Reick, C., Raddatz, T., Brovkin, V., and Gayler, V.: Representation of
natural and anthropogenic land cover change in MPI ESM, Journal of Advances
in Modeling Earth Systems, 5, 459–482, 2013.
</mixed-citation></ref-html>
<ref-html id="bib1.bib112"><label>112</label><mixed-citation>
Saleh, R., Robinson, E. S., Tkacik, D. S., Ahern, A. T., Liu, S., Aiken, A.
C., Sullivan, R. C., Presto, A. A., Dubey, M. K., Yokelson, R. J., Donahue,
N. M., and Robinson, A. L.: Brownness of organics in aerosols from biomass
burning linked to their black carbon content, Nat. Geosci., 7,
647–650, 2014.
</mixed-citation></ref-html>
<ref-html id="bib1.bib113"><label>113</label><mixed-citation>
Savarino, J. and Legrand, M.: High northern latitude forest fires and
vegetation emissions over the last millennium inferred from the chemistry of
a central Greenlabd ice core, J. Geophys. Res., 103,
8267–8279, 1998.
</mixed-citation></ref-html>
<ref-html id="bib1.bib114"><label>114</label><mixed-citation>
Schneck, R., Reick, C. H., and Raddatz, T.: Land contribution to natural
CO<sub>2</sub>
variability on time scales of centuries, Journal of Advances in Modeling
Earth Systems, 5, 354–365, 2013.
</mixed-citation></ref-html>
<ref-html id="bib1.bib115"><label>115</label><mixed-citation>
Shakesby, R. A. and Doerr, S. H.: Wildfire as a hydrological and
geomorphological agent, Earth-Sci. Rev., 74, 269–307, 2006.
</mixed-citation></ref-html>
<ref-html id="bib1.bib116"><label>116</label><mixed-citation>
Swain, A. M.: A History of Fire and Vegetation in Northeastern Minnesota as
Recorded in Lake Sediments, Quaternary Res., 3, 383–396, 1973.
</mixed-citation></ref-html>
<ref-html id="bib1.bib117"><label>117</label><mixed-citation>
Tan, Z. and Huang, C. C.: Holocene wildfire history in loess tableland in
the middle reaches of the Yellow River of China, Holocene, 23,
1466–1476, 2013.
</mixed-citation></ref-html>
<ref-html id="bib1.bib118"><label>118</label><mixed-citation>
Thevenon, F. and Anselmetti, F. S.: Charcoal and fly-ash particles from Lake
Lucerne sediments (Central Switzerland) characterized by image analysis:
anthropologic, stratigraphic and environmental implications, Quaternary Sci. Rev., 26, 2631–2643, 2007.
</mixed-citation></ref-html>
<ref-html id="bib1.bib119"><label>119</label><mixed-citation>
Thonicke, K., Spessa, A., Prentice, I. C., Harrison, S. P., Dong, L., and Carmona-Moreno, C.: The influence of vegetation,
fire spread and fire behaviour on biomass burning and trace gas emissions: results from a process-based model, Biogeosciences, 7,
1991–2011, <a href="http://dx.doi.org/10.5194/bg-7-1991-2010" target="_blank">doi:10.5194/bg-7-1991-2010</a>, 2010.
</mixed-citation></ref-html>
<ref-html id="bib1.bib120"><label>120</label><mixed-citation>
Tinner, W., Hofstetter, S., Zeugin, F., Conedera, M., Wohlgemuth, T.,
Zimmermann, L., and Zweifel, R.: Long-distance transport of macroscopic
charcoal by an intensive crown fire in the Swiss Alps - implications for
fire history reconstruction, Holocene, 16, 287–292, 2006.
</mixed-citation></ref-html>
<ref-html id="bib1.bib121"><label>121</label><mixed-citation>
Tweiten, M. A., Hotchkiss, S. C., Booth, R. K., Calcote, R. R., and Lynch,
E. A.: The response of a jack pine forest to late-Holocene climate
variability in northwestern Wisconsin, Holocene, 19, 1049–1061, 2009.
</mixed-citation></ref-html>
<ref-html id="bib1.bib122"><label>122</label><mixed-citation>
van der Werf, G. R., Randerson, J. T., Giglio, L., Collatz, G. J., Kasibhatla, P. S., and Arellano Jr., A. F.:
Interannual variability in global biomass burning emissions from 1997 to 2004, Atmos. Chem. Phys., 6, 3423–3441, <a href="http://dx.doi.org/10.5194/acp-6-3423-2006" target="_blank">doi:10.5194/acp-6-3423-2006</a>, 2006.
</mixed-citation></ref-html>
<ref-html id="bib1.bib123"><label>123</label><mixed-citation>
van der Werf, G. R., Randerson, J. T., Giglio, L., Collatz, G. J., Mu, M., Kasibhatla, P. S., Morton, D. C.,
DeFries, R. S., Jin, Y., and van Leeuwen, T. T.: Global fire emissions and the contribution of deforestation, savanna,
forest, agricultural, and peat fires (1997–2009), Atmos. Chem. Phys., 10, 11707–11735, <a href="http://dx.doi.org/10.5194/acp-10-11707-2010" target="_blank">doi:10.5194/acp-10-11707-2010</a>, 2010.
</mixed-citation></ref-html>
<ref-html id="bib1.bib124"><label>124</label><mixed-citation>
Vanniere, B., Colombaroli, D., Chapron, E., Leroux, A., Tinner, W., and
Magny, M.: Climate versus human-driven fire regimes in Mediterranean
landscapes: the Holocene record of Lago dell'Accesa (Tuscany, Italy),
Quaternary Sci. Rev., 27, 1181–1196, 2008.
</mixed-citation></ref-html>
<ref-html id="bib1.bib125"><label>125</label><mixed-citation>
Vanniere, B., Power, M. J., Roberts, N., Tinner, W., Carrión, J., Magny,
M., Bartlein, P., Colombaroli, D., Daniau, A. L., Finsinger, W., Gil-Romera,
G., Kaltenrieder, P., Pini, R., Sadori, L., Turner, R., Valsecchi, V., and
Vescovi, E.: Circum-Mediterranean fire activity and climate changes during
the mid-Holocene environmental transition (8500–2500 cal. BP),  Holocene,
21, 53–73, 2011.
</mixed-citation></ref-html>
<ref-html id="bib1.bib126"><label>126</label><mixed-citation>
Verardo, D. J., Froelich, P. N., and McIntyre, A.: Determination of organic
carbon and nitrogen in marine sediments using the Carlo Erba NA-1500
Analyzer, Deep-Sea Res. Pt. I, 37,
157–165, 1990.
</mixed-citation></ref-html>
<ref-html id="bib1.bib127"><label>127</label><mixed-citation>
Walsh, M. K., Whitlock, C., and Bartlein, P. J.: A 14 300-year-long record
of fire-vegetation-climate linkages at Battle Ground Lake, southwestern
Washington, Quaternary Res., 70, 251–264, 2008.
</mixed-citation></ref-html>
<ref-html id="bib1.bib128"><label>128</label><mixed-citation>
Walsh, M. K., Marlon, J. R., Goring, S. J., Brown, K. J., and Gavin, D. G.:
A Regional Perspective on Holocene Fire–Climate–Human Interactions in the
Pacific Northwest of North America, Ann. Assoc. Am. Geogr., 105, 1135–1157, 2015.
</mixed-citation></ref-html>
<ref-html id="bib1.bib129"><label>129</label><mixed-citation>
Wang, Z., Chappellaz, J., Park, K., and Mak, J. E.: Large variations in
southern hemisphere biomass burning during the last 650 years, Science, 330,
1663–1666, 2010.
</mixed-citation></ref-html>
<ref-html id="bib1.bib130"><label>130</label><mixed-citation>
Ward, D. S., Kloster, S., Mahowald, N. M., Rogers, B. M., Randerson, J. T., and Hess, P. G.:
The changing radiative forcing of fires: global model estimates for past, present and future,
Atmos. Chem. Phys., 12, 10857–10886, <a href="http://dx.doi.org/10.5194/acp-12-10857-2012" target="_blank">doi:10.5194/acp-12-10857-2012</a>, 2012.
</mixed-citation></ref-html>
<ref-html id="bib1.bib131"><label>131</label><mixed-citation>
Whitlock, C. and Bartlein, P. J.: Holocene fire activity as a record of past
environmental change, in: Developments in Quaternary Science, edited by: Gillespie, A.
R., Porter, S. C., and Atwater, B. F., Elsevier, Amsterdam, 2004.
</mixed-citation></ref-html>
<ref-html id="bib1.bib132"><label>132</label><mixed-citation>
Whitlock, C., Shafer, S. L., and Marlon, J.: The role of climate and
vegetation change in shaping past and future fire regimes in the
northwestern US and the implications for ecosystem management, Forest
Ecol. Manag., 178, 5–21, 2003.
</mixed-citation></ref-html>
<ref-html id="bib1.bib133"><label>133</label><mixed-citation>
Whitlock, C., Moreno, P. I., and Bartlein, P.: Climatic controls of Holocene
fire patterns in southern South America, Quaternary Res., 68, 28–36,
2007.

</mixed-citation></ref-html>
<ref-html id="bib1.bib134"><label>134</label><mixed-citation>
Williams, A. N., Mooney, S. D., Sisson, S. A., and Marlon, J.: Exploring the
relationship between Aboriginal population indices and fire in Australia
over the last 20 000 years, Palaeogeogr. Palaeocl., 432, 49–57, 2015.
</mixed-citation></ref-html>
<ref-html id="bib1.bib135"><label>135</label><mixed-citation>
Winkler, M. G.: Charcoal analysis for paleoenvironmental interpretation: a
chemical assay, Quaternary Res., 23, 313–326, 1985.
</mixed-citation></ref-html>
<ref-html id="bib1.bib136"><label>136</label><mixed-citation>
Wooller, M. J., Street-Perrott, F. A., and Agnew, A. D. Q.: Late Quaternary
fires and grassland palaeoecology of Mount Kenya, East Africa: evidence from
charred grass cuticles in lake sediments, Palaeogeogr. Palaeocl., 164, 207–230, 2000.
</mixed-citation></ref-html>
<ref-html id="bib1.bib137"><label>137</label><mixed-citation>
Yue, C., Ciais, P., Cadule, P., Thonicke, K., Archibald, S., Poulter, B., Hao, W. M., Hantson, S.,
Mouillot, F., Friedlingstein, P., Maignan, F., and Viovy, N.: Modelling the role of fires in the terrestrial
carbon balance by incorporating SPITFIRE into the global vegetation model ORCHIDEE – Part 1: simulating historical global burned
area and fire regimes, Geosci. Model Dev., 7, 2747–2767, <a href="http://dx.doi.org/10.5194/gmd-7-2747-2014" target="_blank">doi:10.5194/gmd-7-2747-2014</a>, 2014.
</mixed-citation></ref-html>
<ref-html id="bib1.bib138"><label>138</label><mixed-citation>
Zdanowicz, C. M., Zielinski, G. A., and Germani, M. S.: Mount Mazama
eruption: Calendrical age verified and atmospheric impact assessed, Geology,
27, 621–624, 1999.
</mixed-citation></ref-html>
<ref-html id="bib1.bib139"><label>139</label><mixed-citation>
Zennaro, P., Kehrwald, N., McConnell, J. R., Schüpbach, S., Maselli, O. J., Marlon, J., Vallelonga, P.,
Leuenberger, D., Zangrando, R., Spolaor, A., Borrotti, M., Barbaro, E., Gambaro, A., and Barbante, C.:
Fire in ice: two millennia of boreal forest fire history from the Greenland NEEM ice core, Clim. Past, 10, 1905–1924, <a href="http://dx.doi.org/10.5194/cp-10-1905-2014" target="_blank">doi:10.5194/cp-10-1905-2014</a>, 2014.
</mixed-citation></ref-html>--></article>
