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<!DOCTYPE article PUBLIC "-//NLM//DTD Journal Publishing with OASIS Tables v3.0 20080202//EN" "journalpub-oasis3.dtd">
<article xmlns:xlink="http://www.w3.org/1999/xlink" xmlns:mml="http://www.w3.org/1998/Math/MathML" xmlns:oasis="http://docs.oasis-open.org/ns/oasis-exchange/table" xml:lang="en" dtd-version="3.0" article-type="research-article"><?xmltex \makeatother\@nolinetrue\makeatletter?>
  <front>
    <journal-meta><journal-id journal-id-type="publisher">BG</journal-id><journal-title-group>
    <journal-title>Biogeosciences</journal-title>
    <abbrev-journal-title abbrev-type="publisher">BG</abbrev-journal-title><abbrev-journal-title abbrev-type="nlm-ta">Biogeosciences</abbrev-journal-title>
  </journal-title-group><issn pub-type="epub">1726-4189</issn><publisher>
    <publisher-name>Copernicus Publications</publisher-name>
    <publisher-loc>Göttingen, Germany</publisher-loc>
  </publisher></journal-meta>
    <article-meta>
      <article-id pub-id-type="doi">10.5194/bg-18-5767-2021</article-id><title-group><article-title><?xmltex \hack{\vspace*{-1mm}}?>Model simulations of arctic biogeochemistry and permafrost extent are highly sensitive to the implemented snow scheme in LPJ-GUESS</article-title><alt-title>Model simulations are highly sensitive to the implemented snow scheme in LPJ-GUESS</alt-title>
      </title-group><?xmltex \runningtitle{Model simulations are highly sensitive to the implemented snow scheme in LPJ-GUESS}?><?xmltex \runningauthor{A.~Pongracz~et~al.}?>
      <contrib-group>
        <contrib contrib-type="author" corresp="yes" rid="aff1">
          <name><surname>Pongracz</surname><given-names>Alexandra</given-names></name>
          <email>alexandra.pongracz@nateko.lu.se</email>
        <ext-link>https://orcid.org/0000-0001-8840-5077</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff1">
          <name><surname>Wårlind</surname><given-names>David</given-names></name>
          
        <ext-link>https://orcid.org/0000-0002-6257-0338</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff1">
          <name><surname>Miller</surname><given-names>Paul A.</given-names></name>
          
        </contrib>
        <contrib contrib-type="author" corresp="no" rid="aff1 aff2">
          <name><surname>Parmentier</surname><given-names>Frans-Jan W.</given-names></name>
          
        <ext-link>https://orcid.org/0000-0003-2952-7706</ext-link></contrib>
        <aff id="aff1"><label>1</label><institution>Department of Physical Geography and Ecosystem Science, Lund University, Lund, Sweden</institution>
        </aff>
        <aff id="aff2"><label>2</label><institution>Centre for Biogeochemistry in the Anthropocene, Department of Geosciences, University of Oslo, Oslo, Norway</institution>
        </aff>
      </contrib-group>
      <author-notes><corresp id="corr1">Alexandra Pongracz (alexandra.pongracz@nateko.lu.se)</corresp></author-notes><pub-date><day>26</day><month>October</month><year>2021</year></pub-date>
      
      <volume>18</volume>
      <issue>20</issue>
      <fpage>5767</fpage><lpage>5787</lpage>
      <history>
        <date date-type="received"><day>4</day><month>May</month><year>2021</year></date>
           <date date-type="accepted"><day>2</day><month>October</month><year>2021</year></date>
           <date date-type="rev-recd"><day>24</day><month>September</month><year>2021</year></date>
           <date date-type="rev-request"><day>7</day><month>May</month><year>2021</year></date>
      </history>
      <permissions>
        <copyright-statement>Copyright: © 2021 </copyright-statement>
        <copyright-year>2021</copyright-year>
      <license license-type="open-access"><license-p>This work is licensed under the Creative Commons Attribution 4.0 International License. To view a copy of this licence, visit <ext-link ext-link-type="uri" xlink:href="https://creativecommons.org/licenses/by/4.0/">https://creativecommons.org/licenses/by/4.0/</ext-link></license-p></license></permissions><self-uri xlink:href="https://bg.copernicus.org/articles/.html">This article is available from https://bg.copernicus.org/articles/.html</self-uri><self-uri xlink:href="https://bg.copernicus.org/articles/.pdf">The full text article is available as a PDF file from https://bg.copernicus.org/articles/.pdf</self-uri>
      <abstract><title>Abstract</title>
    <p id="d1e115">The Arctic is warming rapidly, especially in winter, which is causing large-scale reductions in snow cover. Snow is one of the main controls on soil
thermodynamics, and changes in its thickness and extent affect both permafrost thaw and soil biogeochemistry. Since soil respiration during the cold
season potentially offsets carbon uptake during the growing season, it is essential to achieve a realistic simulation of the effect of snow cover on
soil conditions to more accurately project the direction of arctic carbon–climate feedbacks under continued winter warming.</p>
    <p id="d1e118">The Lund–Potsdam–Jena General Ecosystem Simulator (LPJ-GUESS) dynamic vegetation model has used – up until now – a single layer snow scheme,
which underestimated the insulation effect of snow, leading to a cold bias in soil temperature. To address this shortcoming, we developed and
integrated a dynamic, multi-layer snow scheme in LPJ-GUESS. The new snow scheme performs well in simulating the insulation of snow at hundreds of
locations across Russia compared to observations. We show that improving this single physical factor enhanced simulations of permafrost extent
compared to an advanced permafrost product, where the overestimation of permafrost cover decreased from 10 % to 5 % using the new snow
scheme. Besides soil thermodynamics, the new snow scheme resulted in a doubled winter respiration and an overall higher vegetation carbon content.</p>
    <p id="d1e121">This study highlights the importance of a correct representation of snow in ecosystem models to project biogeochemical processes that govern climate
feedbacks. The new dynamic snow scheme is an essential improvement in the simulation of cold season processes, which reduces the uncertainty of
model projections. These developments contribute to a more realistic simulation of arctic carbon–climate feedbacks.</p>
  </abstract>
    </article-meta>
  </front>
<body>
      

      <?xmltex \hack{\newpage}?>
<sec id="Ch1.S1" sec-type="intro">
  <label>1</label><title>Introduction</title>
      <p id="d1e135">The Arctic is undergoing rapid warming, with some of the most pronounced changes occurring during the winter <xref ref-type="bibr" rid="bib1.bibx9 bib1.bibx36" id="paren.1"/>. As a result,
snow thickness, the extent of snow-covered area, and snow season length are decreasing, and this is expected to continue in the future
<xref ref-type="bibr" rid="bib1.bibx1 bib1.bibx21" id="paren.2"/>. Snow is an important abiotic component of the Arctic system, since it provides an insulating cover for vegetation and soil.
Snow insulation is recognized as the primary control over soil thermodynamics <xref ref-type="bibr" rid="bib1.bibx25" id="paren.3"/>, and soil temperature is closely connected to
physical (i.e. permafrost active layer depth) and biogeochemical (i.e. decomposition, greenhouse gas emission) processes
<xref ref-type="bibr" rid="bib1.bibx41" id="paren.4"/>. Observations show that snow cover changes have played a major role in a warming trend of permafrost soils of approximately
0.3 <inline-formula><mml:math id="M1" display="inline"><mml:mrow class="unit"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup><mml:mi mathvariant="normal">C</mml:mi></mml:mrow></mml:math></inline-formula> per decade <xref ref-type="bibr" rid="bib1.bibx6 bib1.bibx1" id="paren.5"/>. This warming may lead to increased microbial
activity, decomposition rates and bioavailability of previously frozen soil carbon. Since permafrost soils contain approximately 1600 <inline-formula><mml:math id="M2" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">Pg</mml:mi></mml:mrow></mml:math></inline-formula>
carbon, accounting for half of the global soil carbon storage <xref ref-type="bibr" rid="bib1.bibx20" id="paren.6"/>, there is ample potential for these changes to lead to the release
of the greenhouse gases <inline-formula><mml:math id="M3" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">CO</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> and methane. This has the potential to accelerate global warming <xref ref-type="bibr" rid="bib1.bibx45" id="paren.7"/>, which underlines the need
for a better understanding of drivers and potential feedbacks to better predict the rate and magnitude of future carbon exchange.</p>
      <?pagebreak page5768?><p id="d1e191">Despite numerous field-based and modelling efforts to date, it is still uncertain whether the Arctic will act as a carbon source or sink in the future
<xref ref-type="bibr" rid="bib1.bibx29 bib1.bibx51" id="paren.8"/>. The predicted future carbon balance varies widely among models – between a loss of 641 <inline-formula><mml:math id="M4" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">Pg</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:mi mathvariant="normal">C</mml:mi></mml:mrow></mml:math></inline-formula> and a gain of
167 <inline-formula><mml:math id="M5" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">Pg</mml:mi><mml:mspace width="0.125em" linebreak="nobreak"/><mml:mi mathvariant="normal">C</mml:mi></mml:mrow></mml:math></inline-formula> under RCP8.5 <xref ref-type="bibr" rid="bib1.bibx31" id="paren.9"/> – depending on the representation and level of detail of key processes such as soil temperature
and vegetation dynamics <xref ref-type="bibr" rid="bib1.bibx45 bib1.bibx31" id="paren.10"/>. One of the key goals of model development is to decrease uncertainty of simulations by
refining these processes. While extensive research has been carried out on the mechanics of the growing season, few studies have been directed at cold
season processes. Recent studies suggest that the contribution of the non-growing season to the annual carbon budget may have been underestimated
<xref ref-type="bibr" rid="bib1.bibx43 bib1.bibx28" id="paren.11"/>. A recent meta-analysis by <xref ref-type="bibr" rid="bib1.bibx36" id="text.12"/> found significant wintertime carbon loss and highlighted the large
spread in model simulations of non-growing season greenhouse gas emissions. Models generally underestimated the observed winter flux emissions due to
the inaccuracies in their simulation of cold season respiration. <xref ref-type="bibr" rid="bib1.bibx36" id="text.13"/> stress the need to revise the impact of environmental drivers and
feedbacks in models. Collectively, these efforts demonstrate that the influence of cold season processes on the annual carbon balance is larger than
previously suggested.</p>
      <p id="d1e235">The ability of models to simulate physical and biological processes in the soil is limited by the complexity of their representation of snow. A recent
snow-related model evaluation project analysed the performance of models with different complexity – focusing on variables such as snow-covered area
and snow season length. This SNOWMIP found that a dynamic simulation of internal snowpack processes, such as density and temperature calculations, is
critical to simulate snow thermal profiles <xref ref-type="bibr" rid="bib1.bibx24" id="paren.14"/>. In addition, more complex snow schemes perform better when simulating cold season
processes <xref ref-type="bibr" rid="bib1.bibx49 bib1.bibx46 bib1.bibx53" id="paren.15"/>. To balance computational efficiency and the need for detail, most ecosystem models use an
intermediate-complexity multi-layer snow module <xref ref-type="bibr" rid="bib1.bibx49 bib1.bibx24" id="paren.16"/>. Such schemes may not capture fine-scale internal snowpack processes
such as the evolution of high-density wind slab layers, but they are complex enough to simulate key physical processes – compaction, freeze–thaw
cycles, and liquid water retention – that influence the thermal dampening property of snow. Since LPJ-GUESS had a single-layer static snow
representation, it was found to deviate from observational records of air–soil temperature relationships – simulating cooler winter conditions and
performing poorly when compared to eight land surface models <xref ref-type="bibr" rid="bib1.bibx53" id="paren.17"/>. This showed, combined with previous research, that the snow representation
in LPJ-GUESS needed to be revised to better capture Arctic cold season conditions.</p>
      <p id="d1e250">The primary aim of this study is to improve LPJ-GUESS's simulation of the insulating effect of snow. By developing and integrating a dynamic
intermediate-complexity snow scheme, we also aim to improve the soil temperature and biogeochemistry simulation. To investigate the effect of the new
snow scheme in LPJ-GUESS, we set out to quantify the impact on physical variables, i.e. the direct impact of snow insulation on soil temperature and
permafrost conditions. To further evaluate the snow-related influence on biogeochemistry – such as changes in growing season length – we analyse a set
of biogeochemical variables. Due to differences in soil temperature, we expect to see changes in ecosystem productivity, heterotrophic respiration, and
soil carbon pools. Moreover, we analyse the changes to vegetation dynamics and composition. The updates to the model will allow us to
assess other snow-related processes and feedbacks on a global scale, such as the impact on surface albedo and food access to herbivores, in the future.</p>
</sec>
<sec id="Ch1.S2">
  <label>2</label><title>Materials and methods</title>
      <p id="d1e261">LPJ-GUESS is a process-based dynamic vegetation model, widely applied on regional and global scales <xref ref-type="bibr" rid="bib1.bibx47 bib1.bibx48" id="paren.18"/>. For this study, we
used a customized arctic version of LPJ-GUESS 4.0 (subversion 9905). The model simulates soil freeze–thaw processes and is applicable to studies of
processes at northern high latitudes <xref ref-type="bibr" rid="bib1.bibx33" id="paren.19"/>. In this study we restrict simulations to the northern circumpolar region (above 60<inline-formula><mml:math id="M6" display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula>
latitude) with a spatial resolution of 0.5<inline-formula><mml:math id="M7" display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula> <inline-formula><mml:math id="M8" display="inline"><mml:mo>×</mml:mo></mml:math></inline-formula> 0.5<inline-formula><mml:math id="M9" display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula>. The CRUNCEP global reanalysis climate dataset version 7 was used as input
for all of our model simulations <xref ref-type="bibr" rid="bib1.bibx50" id="paren.20"/>. We ran the model with a 500-year spin-up period to establish an equilibrium vegetation state and
a 40 000-year offline spin-up period for soil conditions.</p>
      <p id="d1e308">LPJ-GUESS simulates vegetation dynamics on individual and patch scales, taking into account growth, competition for resources, and disturbances. This
feature makes it possible to assess how changes in environmental conditions affect vegetation distribution and composition. In this study, we applied
15 plant functional types (PFTs) that characterize arctic ecosystems (see Table S3 in the Supplement). Permafrost dynamics follow <xref ref-type="bibr" rid="bib1.bibx54" id="text.21"/> and
are simulated using the physical characteristics of 15 soil layers, each 10 <inline-formula><mml:math id="M10" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">cm</mml:mi></mml:mrow></mml:math></inline-formula> thick. Soil thermodynamics is governed by climate and snow
conditions, and the thermal properties of each soil layer depend on the ice, water, air, mineral, and organic soil fractions. The layer-specific
thermal properties define the rate of heat transfer through the soil column. For more details on the model structure, see <xref ref-type="bibr" rid="bib1.bibx47 bib1.bibx48" id="text.22"/>, <xref ref-type="bibr" rid="bib1.bibx54" id="text.23"/>, and references therein. To assess the newly developed intermediate-complexity snow scheme's performance and influence, we
conducted simulations with both the old <italic>Static</italic> and the new <italic>Dynamic</italic> snow schemes.</p>
<sec id="Ch1.S2.SS1">
  <label>2.1</label><title>Static snow scheme</title>
      <?pagebreak page5769?><p id="d1e342">The Static snow scheme, which has been in use in LPJ-GUESS until now, treats snow as a single layer with constant values for thermodynamic
parameters. Snowfall is simulated on any given day when precipitation (<inline-formula><mml:math id="M11" display="inline"><mml:mi>P</mml:mi></mml:math></inline-formula>, <inline-formula><mml:math id="M12" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">mm</mml:mi></mml:mrow></mml:math></inline-formula>) occurs and air temperature (<inline-formula><mml:math id="M13" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mi>k</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M14" display="inline"><mml:mrow class="unit"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup><mml:mi mathvariant="normal">C</mml:mi></mml:mrow></mml:math></inline-formula>) is at
or below <inline-formula><mml:math id="M15" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mo>max⁡</mml:mo></mml:msub></mml:mrow></mml:math></inline-formula> (<inline-formula><mml:math id="M16" display="inline"><mml:mrow class="unit"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup><mml:mi mathvariant="normal">C</mml:mi></mml:mrow></mml:math></inline-formula>) – which is the temperature maximum at which precipitation occurs in snow form. <inline-formula><mml:math id="M17" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mi>k</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> denotes the
temperature of a layer <inline-formula><mml:math id="M18" display="inline"><mml:mi>k</mml:mi></mml:math></inline-formula>, in this case the air layer. Snow density (<inline-formula><mml:math id="M19" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi>k</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M20" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kg</mml:mi><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>) and snow thermal conductivity (<inline-formula><mml:math id="M21" display="inline"><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi>k</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>,
<inline-formula><mml:math id="M22" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">J</mml:mi><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">K</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">s</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>) are constant at 362 and 0.196, respectively. Snow heat capacity (<inline-formula><mml:math id="M23" display="inline"><mml:mrow><mml:msub><mml:mi>C</mml:mi><mml:mi>k</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M24" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">J</mml:mi><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">3</mml:mn></mml:mrow></mml:msup><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">K</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>) is calculated by
Eq. (<xref ref-type="disp-formula" rid="Ch1.E1"/>) <xref ref-type="bibr" rid="bib1.bibx16" id="paren.24"/>.
            <disp-formula id="Ch1.E1" content-type="numbered"><label>1</label><mml:math id="M25" display="block"><mml:mrow><mml:msub><mml:mi>C</mml:mi><mml:mi>k</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">1000</mml:mn><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi>k</mml:mi></mml:msub><mml:mo>(</mml:mo><mml:mn mathvariant="normal">0.185</mml:mn><mml:mo>+</mml:mo><mml:mn mathvariant="normal">0.00689</mml:mn><mml:msub><mml:mi>T</mml:mi><mml:mi>k</mml:mi></mml:msub><mml:mo>)</mml:mo></mml:mrow></mml:math></disp-formula></p>
      <p id="d1e576">Compaction processes are not represented in the Static scheme. Snowmelt (melt, <inline-formula><mml:math id="M26" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">mm</mml:mi></mml:mrow></mml:math></inline-formula>) is governed by air temperature and
precipitation and follows a linear function as shown in Eq. (<xref ref-type="disp-formula" rid="Ch1.E2"/>) <xref ref-type="bibr" rid="bib1.bibx12" id="paren.25"/>.
            <disp-formula id="Ch1.E2" content-type="numbered"><label>2</label><mml:math id="M27" display="block"><mml:mrow><mml:mtext>melt</mml:mtext><mml:mo>=</mml:mo><mml:mo>(</mml:mo><mml:mn mathvariant="normal">1.5</mml:mn><mml:mo>+</mml:mo><mml:mn mathvariant="normal">0.007</mml:mn><mml:mo>⋅</mml:mo><mml:mi>P</mml:mi><mml:mo>)</mml:mo><mml:mo>(</mml:mo><mml:msub><mml:mi>T</mml:mi><mml:mi>k</mml:mi></mml:msub><mml:mo>-</mml:mo><mml:msub><mml:mi>T</mml:mi><mml:mo>max⁡</mml:mo></mml:msub><mml:mo>)</mml:mo></mml:mrow></mml:math></disp-formula></p>
      <p id="d1e632">The snowpack is homogeneous in its physical properties, and neither internal processes nor seasonal dynamics are simulated using the Static
scheme. Using this approach, snow conditions are assumed to be uniform across the Arctic regardless of air temperature regime or seasonal snow
dynamics. Due to the heterogeneity in Arctic surface and local climatic conditions, this scheme has a limited ability to represent the variability in
high-latitude ecosystems – this is the main shortcoming of the Static snow scheme that this study sets out to improve.</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F1"><?xmltex \currentcnt{1}?><?xmltex \def\figurename{Figure}?><label>Figure 1</label><caption><p id="d1e638">Snowpack structure and physical properties. Where <inline-formula><mml:math id="M28" display="inline"><mml:mi>z</mml:mi></mml:math></inline-formula> shows layer depth, <inline-formula><mml:math id="M29" display="inline"><mml:mi>W</mml:mi></mml:math></inline-formula> water content, <inline-formula><mml:math id="M30" display="inline"><mml:mi>I</mml:mi></mml:math></inline-formula> ice content, <inline-formula><mml:math id="M31" display="inline"><mml:mi mathvariant="italic">ρ</mml:mi></mml:math></inline-formula> density, <inline-formula><mml:math id="M32" display="inline"><mml:mi>C</mml:mi></mml:math></inline-formula> heat capacity, <inline-formula><mml:math id="M33" display="inline"><mml:mi>K</mml:mi></mml:math></inline-formula> thermal conductivity, <inline-formula><mml:math id="M34" display="inline"><mml:mi>D</mml:mi></mml:math></inline-formula> thermal diffusivity, and <inline-formula><mml:math id="M35" display="inline"><mml:mi>T</mml:mi></mml:math></inline-formula> layer temperature. A detailed list of the used variables can be found in Table <xref ref-type="table" rid="App1.Ch1.S1.T3"/>.</p></caption>
          <?xmltex \igopts{width=207.705118pt}?><graphic xlink:href="https://bg.copernicus.org/articles/18/5767/2021/bg-18-5767-2021-f01.png"/>

        </fig>

</sec>
<sec id="Ch1.S2.SS2">
  <label>2.2</label><title>Dynamic snow scheme</title>
      <p id="d1e714">The schematic structure of the multi-layer snow scheme is shown in Fig. <xref ref-type="fig" rid="Ch1.F1"/>. The occurrence of snowfall on any given day depends on air
temperature and precipitation, using the same principle as for the Static scheme. Fresh snow density (<inline-formula><mml:math id="M36" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mtext>fresh</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula>,
<inline-formula><mml:math id="M37" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kg</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">3</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>) is calculated by taking into account air temperature and wind speed, following Eq. (<xref ref-type="disp-formula" rid="Ch1.E3"/>), where <inline-formula><mml:math id="M38" display="inline"><mml:mi>a</mml:mi></mml:math></inline-formula>, <inline-formula><mml:math id="M39" display="inline"><mml:mi>b</mml:mi></mml:math></inline-formula>, and <inline-formula><mml:math id="M40" display="inline"><mml:mi>c</mml:mi></mml:math></inline-formula> are
scaling parameters defined by <xref ref-type="bibr" rid="bib1.bibx49" id="text.26"/> (for parameter values see Table <xref ref-type="table" rid="App1.Ch1.S1.T3"/> in the Appendix).
            <disp-formula id="Ch1.E3" content-type="numbered"><label>3</label><mml:math id="M41" display="block"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mtext>fresh</mml:mtext></mml:msub><mml:mo>=</mml:mo><mml:mi>a</mml:mi><mml:mo>+</mml:mo><mml:mi>b</mml:mi><mml:mo>⋅</mml:mo><mml:msub><mml:mi>T</mml:mi><mml:mo>max⁡</mml:mo></mml:msub><mml:mo>+</mml:mo><mml:mi>c</mml:mi><mml:msubsup><mml:mi>U</mml:mi><mml:mn mathvariant="normal">10</mml:mn><mml:mn mathvariant="normal">0.5</mml:mn></mml:msubsup></mml:mrow></mml:math></disp-formula></p>
      <p id="d1e812"><inline-formula><mml:math id="M42" display="inline"><mml:mrow><mml:msub><mml:mi>U</mml:mi><mml:mn mathvariant="normal">10</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> denotes the 10 <inline-formula><mml:math id="M43" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">m</mml:mi></mml:mrow></mml:math></inline-formula> height wind speed (<inline-formula><mml:math id="M44" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">m</mml:mi><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">s</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>), following the detailed snowpack model Crocus <xref ref-type="bibr" rid="bib1.bibx49" id="paren.27"/>. To avoid
unrealistically low snow density values that may occur in rare cases, the density minimum is set to 100 <inline-formula><mml:math id="M45" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kg</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">3</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>.</p>
      <p id="d1e870">The new snow scheme simulates internal snowpack dynamics with up to five snow layers, taking into consideration each layer's depth. Fresh snow either
initiates a snowpack or is added to already existing snow layers. If the freshly fallen snow is added to the snowpack, the physical properties of the
snow layer are updated. The number and thickness of snow layers are defined according to predefined thresholds: a new snow layer is initialized when
an existing layer exceeds twice the prescribed threshold height (2 <inline-formula><mml:math id="M46" display="inline"><mml:mo>×</mml:mo></mml:math></inline-formula> 100 <inline-formula><mml:math id="M47" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">mm</mml:mi></mml:mrow></mml:math></inline-formula>). If a single snow layer exists but does not reach the
minimum height (set to 50 <inline-formula><mml:math id="M48" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">mm</mml:mi></mml:mrow></mml:math></inline-formula>), the shallow snow layer properties are combined with the top soil layer. Thereafter, their properties (ice,
air, and liquid water fractions and heat capacity) are scaled using weighted averages based on the layer's ice, water, and air fractions for the sake of
computational stability. In the case where all five layers exceed the prescribed maximum threshold, the bottom layer accumulates snow in order to
preserve and align vertical resolution near the surface of the snowpack. The snow layer density (<inline-formula><mml:math id="M49" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi>k</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>) and depth (<inline-formula><mml:math id="M50" display="inline"><mml:mrow><mml:msub><mml:mi>z</mml:mi><mml:mi>k</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M51" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">m</mml:mi></mml:mrow></mml:math></inline-formula>) relationship
is described by Eq. (<xref ref-type="disp-formula" rid="Ch1.E4"/>), where <inline-formula><mml:math id="M52" display="inline"><mml:mrow><mml:msub><mml:mi>I</mml:mi><mml:mi>k</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> (<inline-formula><mml:math id="M53" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kg</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>) defines the ice content of a layer <xref ref-type="bibr" rid="bib1.bibx26" id="paren.28"/>.
            <disp-formula id="Ch1.E4" content-type="numbered"><label>4</label><mml:math id="M54" display="block"><mml:mrow><mml:msub><mml:mi>z</mml:mi><mml:mi>k</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:mstyle displaystyle="true"><mml:mfrac style="display"><mml:mrow><mml:msub><mml:mi>I</mml:mi><mml:mi>k</mml:mi></mml:msub></mml:mrow><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi>k</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle></mml:mrow></mml:math></disp-formula></p>
      <p id="d1e986">The density of a snow layer changes through compaction, which is simulated by two processes: (1) mechanical compaction due to pressure from the
overlying snow layers as shown in Eq. (<xref ref-type="disp-formula" rid="Ch1.E5"/>) <xref ref-type="bibr" rid="bib1.bibx4" id="paren.29"/>.
            <disp-formula id="Ch1.E5" content-type="numbered"><label>5</label><mml:math id="M55" display="block"><mml:mrow><mml:mstyle displaystyle="true"><mml:mfrac style="display"><mml:mrow><mml:mo>∂</mml:mo><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi>k</mml:mi></mml:msub></mml:mrow><mml:mrow><mml:mo>∂</mml:mo><mml:mi>t</mml:mi></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>=</mml:mo><mml:mstyle displaystyle="true"><mml:mfrac style="display"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi>k</mml:mi></mml:msub><mml:mi>g</mml:mi><mml:msub><mml:mi>M</mml:mi><mml:mi>k</mml:mi></mml:msub></mml:mrow><mml:mrow><mml:msub><mml:mi mathvariant="italic">η</mml:mi><mml:mi>k</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle><mml:mi>exp⁡</mml:mi><mml:mfenced close=")" open="("><mml:mrow><mml:mstyle displaystyle="true"><mml:mfrac style="display"><mml:mrow><mml:msub><mml:mi>k</mml:mi><mml:mi>s</mml:mi></mml:msub></mml:mrow><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mo>max⁡</mml:mo></mml:msub></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>-</mml:mo><mml:mstyle displaystyle="true"><mml:mfrac style="display"><mml:mrow><mml:msub><mml:mi>k</mml:mi><mml:mi>s</mml:mi></mml:msub></mml:mrow><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mi>k</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>-</mml:mo><mml:mstyle displaystyle="true"><mml:mfrac style="display"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi>k</mml:mi></mml:msub></mml:mrow><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mn mathvariant="normal">0</mml:mn></mml:msub></mml:mrow></mml:mfrac></mml:mstyle></mml:mrow></mml:mfenced></mml:mrow></mml:math></disp-formula></p>
      <?pagebreak page5770?><p id="d1e1088">The increase in the snow layer's density (<inline-formula><mml:math id="M56" display="inline"><mml:mrow><mml:mo>∂</mml:mo><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi>k</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>) depends on the mass of overlying layers (<inline-formula><mml:math id="M57" display="inline"><mml:mrow><mml:msub><mml:mi>M</mml:mi><mml:mi>k</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M58" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kg</mml:mi></mml:mrow></mml:math></inline-formula>). <inline-formula><mml:math id="M59" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">η</mml:mi><mml:mi>k</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>
(<inline-formula><mml:math id="M60" display="inline"><mml:mrow><mml:msup><mml:mn mathvariant="normal">10</mml:mn><mml:mn mathvariant="normal">6</mml:mn></mml:msup></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M61" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">Pa</mml:mi><mml:mspace width="0.125em" linebreak="nobreak"/><mml:mi mathvariant="normal">s</mml:mi></mml:mrow></mml:math></inline-formula>) denotes the compactive viscosity factor, <inline-formula><mml:math id="M62" display="inline"><mml:mrow><mml:msub><mml:mi>k</mml:mi><mml:mi mathvariant="normal">s</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> is an empirical constant defined by <xref ref-type="bibr" rid="bib1.bibx4" id="text.30"/> with a value of
4000 <inline-formula><mml:math id="M63" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">K</mml:mi></mml:mrow></mml:math></inline-formula>, and <inline-formula><mml:math id="M64" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mn mathvariant="normal">0</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> is a reference density (50 <inline-formula><mml:math id="M65" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kg</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>). Snow density may also change by (2) phase changes as a result of
freeze–thaw processes within the layers. If a layer's snow or liquid water content changed during freeze–thaw events, its depth and density properties
are recalculated, taking into account the snow and ice fractions of the layer as shown in Eq. (<xref ref-type="disp-formula" rid="Ch1.E4"/>).</p>
      <p id="d1e1210">In contrast to the Static formulation, phase changes within the snow layers depend on the layer's internal temperature, and this controls the
melting process in the Dynamic snow scheme. This development enables the simulation of mid-winter melt events and ensures an improved
representation of internal snowpack thermodynamics. Upon melt, each layer can retain a fraction of liquid water based on Eq. (<xref ref-type="disp-formula" rid="Ch1.E6"/>), where
rw<inline-formula><mml:math id="M66" display="inline"><mml:msub><mml:mi/><mml:mo>min⁡</mml:mo></mml:msub></mml:math></inline-formula> and rw<inline-formula><mml:math id="M67" display="inline"><mml:msub><mml:mi/><mml:mo>max⁡</mml:mo></mml:msub></mml:math></inline-formula> are empirical constants and <inline-formula><mml:math id="M68" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">t</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> is a reference density <xref ref-type="bibr" rid="bib1.bibx52 bib1.bibx2" id="paren.31"/>.
            <disp-formula id="Ch1.E6" content-type="numbered"><label>6</label><mml:math id="M69" display="block"><mml:mtable class="split" rowspacing="0.2ex" displaystyle="true" columnalign="right left"><mml:mtr><mml:mtd><mml:mrow><mml:msub><mml:mi>W</mml:mi><mml:mrow><mml:mtext>cap</mml:mtext><mml:mo>,</mml:mo><mml:mo>max⁡</mml:mo></mml:mrow></mml:msub><mml:mo>=</mml:mo></mml:mrow></mml:mtd><mml:mtd><mml:mrow><mml:mspace width="0.25em" linebreak="nobreak"/><mml:msub><mml:mi>I</mml:mi><mml:mi>k</mml:mi></mml:msub><mml:mo mathsize="2.0em">[</mml:mo><mml:msub><mml:mi mathvariant="normal">rw</mml:mi><mml:mo>min⁡</mml:mo></mml:msub><mml:mo>+</mml:mo><mml:mo>(</mml:mo><mml:msub><mml:mi mathvariant="normal">rw</mml:mi><mml:mo>max⁡</mml:mo></mml:msub><mml:mo>-</mml:mo><mml:msub><mml:mi mathvariant="normal">rw</mml:mi><mml:mo>min⁡</mml:mo></mml:msub><mml:mo>)</mml:mo></mml:mrow></mml:mtd></mml:mtr><mml:mtr><mml:mtd/><mml:mtd><mml:mrow><mml:mo>⋅</mml:mo><mml:mo movablelimits="false">max⁡</mml:mo><mml:mfenced close=")" open="("><mml:mrow><mml:mn mathvariant="normal">0</mml:mn><mml:mo>,</mml:mo><mml:mstyle displaystyle="true"><mml:mfrac style="display"><mml:mrow><mml:mo>(</mml:mo><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">t</mml:mi></mml:msub><mml:mo>-</mml:mo><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi>k</mml:mi></mml:msub><mml:mo>)</mml:mo></mml:mrow><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">t</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle></mml:mrow></mml:mfenced><mml:mo mathsize="2.0em">]</mml:mo></mml:mrow></mml:mtd></mml:mtr></mml:mtable></mml:math></disp-formula></p>
      <p id="d1e1342">If the liquid water content (<inline-formula><mml:math id="M70" display="inline"><mml:mrow><mml:msub><mml:mi>W</mml:mi><mml:mi>k</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M71" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">mm</mml:mi></mml:mrow></mml:math></inline-formula>) of a layer exceeds the maximum water holding capacity (<inline-formula><mml:math id="M72" display="inline"><mml:mrow><mml:msub><mml:mi>W</mml:mi><mml:mrow><mml:mtext>cap</mml:mtext><mml:mo>,</mml:mo><mml:mo>max⁡</mml:mo></mml:mrow></mml:msub></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M73" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">mm</mml:mi></mml:mrow></mml:math></inline-formula>), water is
passed to the layer underneath following a simple bucket model. Rain-on-snow events (ROSs) are simulated if it rains while a snowpack is present. The
energy of rainwater may induce phase changes in the snow layers. The overflow liquid water is forwarded to the underlying snow layers and lastly to
the top soil layer to percolate to the soil or to be discharged as runoff.</p>
      <p id="d1e1388">Each layer is characterized thermodynamically by the following physical properties: density, temperature, thermal conductivity, heat capacity, and
diffusivity (<inline-formula><mml:math id="M74" display="inline"><mml:mrow><mml:msub><mml:mi>D</mml:mi><mml:mi>k</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M75" display="inline"><mml:mrow class="unit"><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msup><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">d</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>). Thermal conductivity is calculated using density as shown in Eq. (<xref ref-type="disp-formula" rid="Ch1.E7"/>) <xref ref-type="bibr" rid="bib1.bibx4" id="paren.32"/>,
following a power function (<inline-formula><mml:math id="M76" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">J</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">3</mml:mn></mml:mrow></mml:msup><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">K</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>).
            <disp-formula id="Ch1.E7" content-type="numbered"><label>7</label><mml:math id="M77" display="block"><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi>k</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:mn mathvariant="normal">2.22</mml:mn><mml:msup><mml:mfenced close=")" open="("><mml:mstyle displaystyle="true"><mml:mfrac style="display"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:msub><mml:mi/><mml:mi>k</mml:mi></mml:msub></mml:msub></mml:mrow><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mn mathvariant="normal">0</mml:mn></mml:msub></mml:mrow></mml:mfrac></mml:mstyle></mml:mfenced><mml:mn mathvariant="normal">1.88</mml:mn></mml:msup></mml:mrow></mml:math></disp-formula></p>
      <p id="d1e1488">Heat capacity is determined by taking into account snow layer density and temperature according to Eq. (<xref ref-type="disp-formula" rid="Ch1.E1"/>). The snow diffusivity is
calculated by Eq. (<xref ref-type="disp-formula" rid="Ch1.E8"/>). Soil and snow layer temperatures are computed, taking into account each layer's thermal conductivity, heat
capacity, and height, using the Crank–Nicolson finite difference method to solve Eq. (<xref ref-type="disp-formula" rid="Ch1.E9"/>) <xref ref-type="bibr" rid="bib1.bibx26" id="paren.33"/>.

                <disp-formula specific-use="gather" content-type="numbered"><mml:math id="M78" display="block"><mml:mtable displaystyle="true"><mml:mlabeledtr id="Ch1.E8"><mml:mtd><mml:mtext>8</mml:mtext></mml:mtd><mml:mtd><mml:mrow><mml:mstyle class="stylechange" displaystyle="true"/><mml:msub><mml:mi>D</mml:mi><mml:mi>k</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:mstyle displaystyle="true"><mml:mfrac style="display"><mml:mrow><mml:mi>K</mml:mi><mml:msub><mml:mi/><mml:mi>k</mml:mi></mml:msub></mml:mrow><mml:mrow><mml:msub><mml:mi>C</mml:mi><mml:mi>k</mml:mi></mml:msub></mml:mrow></mml:mfrac></mml:mstyle></mml:mrow></mml:mtd></mml:mlabeledtr><mml:mlabeledtr id="Ch1.E9"><mml:mtd><mml:mtext>9</mml:mtext></mml:mtd><mml:mtd><mml:mrow><mml:mstyle displaystyle="true"><mml:mfrac style="display"><mml:mrow><mml:mo>∂</mml:mo><mml:mi>T</mml:mi></mml:mrow><mml:mrow><mml:mo>∂</mml:mo><mml:mi>t</mml:mi></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>=</mml:mo><mml:mstyle displaystyle="true"><mml:mfrac style="display"><mml:mo>∂</mml:mo><mml:mrow><mml:mo>∂</mml:mo><mml:mi>z</mml:mi></mml:mrow></mml:mfrac></mml:mstyle><mml:mfenced open="(" close=")"><mml:mrow><mml:mi>D</mml:mi><mml:mo>(</mml:mo><mml:mi>k</mml:mi><mml:mo>)</mml:mo><mml:mstyle displaystyle="true"><mml:mfrac style="display"><mml:mrow><mml:mo>∂</mml:mo><mml:mi>T</mml:mi></mml:mrow><mml:mrow><mml:mo>∂</mml:mo><mml:mi>z</mml:mi></mml:mrow></mml:mfrac></mml:mstyle></mml:mrow></mml:mfenced></mml:mrow></mml:mtd></mml:mlabeledtr></mml:mtable></mml:math></disp-formula></p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F2"><?xmltex \currentcnt{2}?><?xmltex \def\figurename{Figure}?><label>Figure 2</label><caption><p id="d1e1588">Steps in the daily computational cycle for the Dynamic snow scheme. Blue arrows indicate workflow in case a snowpack is present on the ground, while orange arrows show steps when a snowpack is absent.</p></caption>
          <?xmltex \igopts{width=236.157874pt}?><graphic xlink:href="https://bg.copernicus.org/articles/18/5767/2021/bg-18-5767-2021-f02.png"/>

        </fig>

      <p id="d1e1597">The computational cycle ends by rearranging the layers based on the depth thresholds, taking into account the potential liquid water content. First we
re-calculate each snow layer's depth based on the amount of snow and liquid water using Eq. (<xref ref-type="disp-formula" rid="Ch1.E4"/>). We then re-arrange the layers by using the leaky-bucket
approach, where the snow layers are filled up from the bottom layer (closest to the surface). If the threshold depth is reached, a new snow layer is
initiated and the process continues until the total depth of the snowpack is distributed to the specific snow layers. The overflow meltwater is passed
to the soil for percolation after this step. This cycle is repeated each day when there is a snow or rain-on-snow event. The daily snow cycle of the
Dynamic snow scheme is depicted in Fig. <xref ref-type="fig" rid="Ch1.F2"/>.</p>
      <p id="d1e1604">Besides the changes in the representation of snow, the calculation of heterotrophic respiration below 0 <inline-formula><mml:math id="M79" display="inline"><mml:mrow class="unit"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup><mml:mi mathvariant="normal">C</mml:mi></mml:mrow></mml:math></inline-formula> was changed following a
recent data synthesis <xref ref-type="bibr" rid="bib1.bibx36" id="paren.34"/> to better represent arctic conditions. This adjustment was implemented for both the Static and
Dynamic schemes. The minimum decomposition temperature was set to <inline-formula><mml:math id="M80" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>20 <inline-formula><mml:math id="M81" display="inline"><mml:mrow class="unit"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup><mml:mi mathvariant="normal">C</mml:mi></mml:mrow></mml:math></inline-formula>, and the Q10 value was set to 2.9. A comparison between the
old and new functions is shown in the Supplement (Fig. S1 in the Supplement). This adjustment led to higher soil respiration in both schemes during
the cold season compared to the old model set-up.</p>
      <p id="d1e1641">The implemented processes and physical representations are simpler than in dedicated, high-resolution snow models – such as Crocus and SNOWPACK
<xref ref-type="bibr" rid="bib1.bibx27 bib1.bibx49" id="paren.35"/> – but reflect the model improvements identified as being most important in previous model inter-comparison
studies <xref ref-type="bibr" rid="bib1.bibx24" id="paren.36"/>. These improvements enable us to simulate a more realistic range of snow conditions and soil thermal conditions across the
Arctic.</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F3"><?xmltex \currentcnt{3}?><?xmltex \def\figurename{Figure}?><label>Figure 3</label><caption><p id="d1e1652">Snowpack dynamics at the Zackenberg GeoBasis station. Density values for the layers are extrapolated – from three and five layers for the observational and modelled data, respectively. The colours of the snowpack indicate snow density.</p></caption>
          <?xmltex \igopts{width=199.169291pt}?><graphic xlink:href="https://bg.copernicus.org/articles/18/5767/2021/bg-18-5767-2021-f03.png"/>

        </fig>

</sec>
<?pagebreak page5771?><sec id="Ch1.S2.SS3">
  <label>2.3</label><title>Simulation set-up</title>
      <p id="d1e1670">The performance of LPJ-GUESS using both snow schemes was compared at the site and regional levels. Modelled properties were compared to observational
datasets, when available. We quantified the correspondence between simulated and measured variables using statistical methods.
<list list-type="custom"><list-item><label>i.</label>
      <p id="d1e1675"><italic>Site-level comparison</italic>.</p>
      <p id="d1e1680">To highlight the differences in snowpack dynamics using the two snow schemes, we compiled a detailed single-site model–data comparison of the
internal snowpack structure (Fig. <xref ref-type="fig" rid="Ch1.F3"/>).  Afterwards, we ran the model for five well-studied northern high-latitude sites in order to
identify how well the two snow schemes can simulate snow and soil temperature at the site level. These sites are Abisko, Zackenberg, Bayelva, Kytalyk,
and Samoylov – see site details in Table S1 in the Supplement. Measurements of snow depth and soil temperature were sorted and averaged on a daily
basis – 10 years for the simulations and all available years for observations at each site. Model outputs were examined and compared to these
time series to evaluate the snow schemes' ability to simulate snow depth and soil temperature seasonality adequately.</p></list-item><list-item><label>ii.</label>
      <p id="d1e1686"><italic>Regional simulations</italic>.</p>
      <p id="d1e1691">We conducted simulations for a set of Russian sites (256 sites) which were part of the study by <xref ref-type="bibr" rid="bib1.bibx53" id="text.37"/>, as a follow-up, and re-evaluate
the snow insulation effect in LPJ-GUESS over a large region. First, snow depth and soil temperature data were sorted monthly for each site for the
years 1980–2000. Site observations were provided by the All-Russian Research Institute of Hydrometeorological Information – World Data Centre
(RIHMI-WDC; <uri>http://meteo.ru/</uri>, last access: 3 January 2019). Following this, averages were calculated for December, January,
and February. The difference between soil (25 <inline-formula><mml:math id="M82" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">cm</mml:mi></mml:mrow></mml:math></inline-formula> depth) and air temperature – henceforth <inline-formula><mml:math id="M83" display="inline"><mml:mrow><mml:mi mathvariant="normal">Δ</mml:mi><mml:mi>T</mml:mi></mml:mrow></mml:math></inline-formula> – was used as a proxy to evaluate the
strength of the model-simulated insulation effect. Snow depth, soil temperature, and <inline-formula><mml:math id="M84" display="inline"><mml:mrow><mml:mi mathvariant="normal">Δ</mml:mi><mml:mi>T</mml:mi></mml:mrow></mml:math></inline-formula> series were grouped according to air temperature to
evaluate the insulation capacity under different temperature regimes.</p></list-item><list-item><label>iii.</label>
      <p id="d1e1729"><italic>Pan-Arctic simulations</italic>.</p>
      <p id="d1e1734">Finally, we conducted model simulations across the Arctic to assess the effect of changing the snow scheme on selected physical and biogeochemical
variables and vegetation properties. When applicable, variables were averaged over December, January, and February to emphasize the effect on the
winter season. Instead of the absolute results, we show the difference between the set of simulations, calculated as the difference between
Dynamic and Static model outputs.</p></list-item></list></p>
</sec>
</sec>
<sec id="Ch1.S3">
  <label>3</label><title>Results</title>
<sec id="Ch1.S3.SS1">
  <label>3.1</label><title>Site-level simulations</title>
      <p id="d1e1753">Prior to the evaluation of the large-scale performance of the new Dynamic snow scheme, we conducted a single-site comparison to examine the
validity of the results. These detailed snowpack observations from Zackenberg helped to determine whether the Dynamic scheme can simulate
internal snowpack dynamics, snow depth, and snow density.</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F4" specific-use="star"><?xmltex \currentcnt{4}?><?xmltex \def\figurename{Figure}?><label>Figure 4</label><caption><p id="d1e1758">Seasonal cycle of <bold>(a)</bold> snow depth and <bold>(b)</bold> soil temperature at 25 <inline-formula><mml:math id="M85" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">cm</mml:mi></mml:mrow></mml:math></inline-formula> depth for the studied sites, comparing model simulations and observations. Site statistics show the spread of monthly snow depth and soil temperature values for the respective sites – excluding the summer months (June–July–August).</p></caption>
          <?xmltex \igopts{width=455.244094pt}?><graphic xlink:href="https://bg.copernicus.org/articles/18/5767/2021/bg-18-5767-2021-f04.png"/>

        </fig>

      <p id="d1e1781">We established the ability of the new snow scheme to simulate snow conditions by comparing a simulated snowpack with snow depth and density
observations from Zackenberg (2013–2014 snow season). Figure <xref ref-type="fig" rid="Ch1.F3"/> presents the observed and simulated snowpack by the Dynamic and
Static schemes. This figure shows that the Dynamic scheme simulates comparable snow depth and that the simulated snow densities
follow the observed snow density pattern through the snow season. Density values are compared qualitatively, since it is difficult to accurately align
the observational and modelled layer densities. To be consequent, we used global climatic forcing data for all simulations in this study, including
this site-scale comparison. This fact should be taken into account when interpreting the model–data comparison in this section – as some of the
differences may be derived from the differences in climatic data.</p>
      <p id="d1e1787">There are lower densities early in the snow season, with fresh snow having low density, while density increases in<?pagebreak page5772?> late spring, during the melt
season. The Static scheme with constant snow density simulated a somewhat higher-than-observed snow depth. Thermal properties in snow layers
are derived from density, and this is especially important in the Dynamic snow scheme. Dynamically simulated density translates to more
realistic thermal conductivity dynamics, which governs the rate of heat transfer through the snowpack. This feature is essential in simulating a
reliable atmosphere–snow–soil heat transfer interaction. The difference in snow depth between the Static and Dynamic simulations is
most visible at the end of the snow season, before the start of snowmelt – as indicated in Fig. <xref ref-type="fig" rid="Ch1.F3"/>, bottom panel.</p>
      <p id="d1e1792">Overall, the new scheme reproduces the snow dynamics over the cold season better than the Static scheme. Taken together, these results
suggest that the Dynamic scheme is skilled in simulating the snowpack's internal structure and dynamics. Since the Static scheme
has a constant snow density throughout the snow season, the Dynamic scheme is expected to better capture the seasonal behaviour of snow and
soil conditions. The Zackenberg site comparison indicated that the Dynamic scheme successfully integrated these key processes affecting the
density over the snow season. In this study, we used a global climate forcing dataset, which may explain some of the observed model–observation
differences. The mismatch between snow observations and simulations is influenced by the use of the global model forcing dataset instead of
site-specific temperature, precipitation, or snowfall time series.</p>

<?xmltex \floatpos{t}?><table-wrap id="Ch1.T1" specific-use="star"><?xmltex \currentcnt{1}?><label>Table 1</label><caption><p id="d1e1798">RMSE for soil temperature and snow depth for the applied snow schemes for the single site simulations.</p></caption><oasis:table frame="topbot"><oasis:tgroup cols="7">
     <oasis:colspec colnum="1" colname="col1" align="left"/>
     <oasis:colspec colnum="2" colname="col2" align="left"/>
     <oasis:colspec colnum="3" colname="col3" align="right"/>
     <oasis:colspec colnum="4" colname="col4" align="right"/>
     <oasis:colspec colnum="5" colname="col5" align="right"/>
     <oasis:colspec colnum="6" colname="col6" align="right"/>
     <oasis:colspec colnum="7" colname="col7" align="right"/>
     <oasis:thead>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1"/>
         <oasis:entry colname="col2">Snow scheme</oasis:entry>
         <oasis:entry colname="col3">Abisko</oasis:entry>
         <oasis:entry colname="col4">Kytalyk</oasis:entry>
         <oasis:entry colname="col5">Zackenberg</oasis:entry>
         <oasis:entry colname="col6">Samoylov</oasis:entry>
         <oasis:entry colname="col7">Bayelva</oasis:entry>
       </oasis:row>
     </oasis:thead>
     <oasis:tbody>
       <oasis:row>
         <oasis:entry colname="col1">Soil <inline-formula><mml:math id="M86" display="inline"><mml:mi>T</mml:mi></mml:math></inline-formula> (<inline-formula><mml:math id="M87" display="inline"><mml:mrow class="unit"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup><mml:mi mathvariant="normal">C</mml:mi></mml:mrow></mml:math></inline-formula>)</oasis:entry>
         <oasis:entry colname="col2">Static</oasis:entry>
         <oasis:entry colname="col3">6.13</oasis:entry>
         <oasis:entry colname="col4">12.35</oasis:entry>
         <oasis:entry colname="col5">3.17</oasis:entry>
         <oasis:entry colname="col6">14.97</oasis:entry>
         <oasis:entry colname="col7">5.65</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1"/>
         <oasis:entry colname="col2">Dynamic</oasis:entry>
         <oasis:entry colname="col3">1.36</oasis:entry>
         <oasis:entry colname="col4">5.35</oasis:entry>
         <oasis:entry colname="col5">3.34</oasis:entry>
         <oasis:entry colname="col6">4.67</oasis:entry>
         <oasis:entry colname="col7">2.56</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Snow depth (<inline-formula><mml:math id="M88" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">m</mml:mi></mml:mrow></mml:math></inline-formula>)</oasis:entry>
         <oasis:entry colname="col2">Static</oasis:entry>
         <oasis:entry colname="col3">0.17</oasis:entry>
         <oasis:entry colname="col4">0.09</oasis:entry>
         <oasis:entry colname="col5">0.11</oasis:entry>
         <oasis:entry colname="col6">0.06</oasis:entry>
         <oasis:entry colname="col7">0.18</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"/>
         <oasis:entry colname="col2">Dynamic</oasis:entry>
         <oasis:entry colname="col3">0.14</oasis:entry>
         <oasis:entry colname="col4">0.07</oasis:entry>
         <oasis:entry colname="col5">0.10</oasis:entry>
         <oasis:entry colname="col6">0.12</oasis:entry>
         <oasis:entry colname="col7">0.12</oasis:entry>
       </oasis:row>
     </oasis:tbody>
   </oasis:tgroup></oasis:table></table-wrap>

      <p id="d1e1976">We moved to a multi-site analysis to compare the Static and Dynamic snow schemes on five well-documented sites. To assess the
performance of the two snow schemes, we composed seasonal cycles based on monthly averages of (a) snow depth and (b) soil temperature at 25 <inline-formula><mml:math id="M89" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">cm</mml:mi></mml:mrow></mml:math></inline-formula>
depth, shown in Fig. <xref ref-type="fig" rid="Ch1.F4"/>. The corresponding root-mean-squared error (RMSE) for each study site is shown in
Table <xref ref-type="table" rid="Ch1.T1"/>. Generally, the Dynamic scheme shows only minor improvements in the simulation of snow depth. Despite this, modelled
soil temperatures are much closer to the observed values for all sites, especially during the winter months. This behaviour highlights that changing
the internal snowpack dynamics with the Dynamic snow scheme had a significant effect on soil temperature, even when the simulated snow depth
differed marginally. The changes in soil temperature are due to the differences in snow thermal properties, which significantly influenced the
insulation capacity of the snowpack.</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F5" specific-use="star"><?xmltex \currentcnt{5}?><?xmltex \def\figurename{Figure}?><label>Figure 5</label><caption><p id="d1e1993">Comparison of the observed and modelled snow insulation effect at the Russian sites between observations and model simulations using the Dynamic and Static schemes. <bold>(a–c)</bold> Soil temperature and snow depth relationship. <bold>(d–f)</bold> Difference in air–soil temperature and snow depth relationship. Snow depth presented on the horizontal axis is classified in 5 <inline-formula><mml:math id="M90" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">cm</mml:mi></mml:mrow></mml:math></inline-formula> depth bins. Colours indicate different air temperature regimes, and upper and lower bars show the 25th and 75th percentiles.</p></caption>
          <?xmltex \igopts{width=341.433071pt}?><graphic xlink:href="https://bg.copernicus.org/articles/18/5767/2021/bg-18-5767-2021-f05.png"/>

        </fig>

      <?pagebreak page5773?><p id="d1e2017">The implementation of the new snow scheme resulted in significant changes within the snowpack that are responsible for the improved soil temperature
simulation. The Static scheme applies constant snow density and thermal conductivity, which defines the rate of heat transfer through the
snowpack. In the Dynamic scheme, thermal conductivity is dependent on the dynamically updated density; therefore the new scheme can achieve a
more realistic simulation of snow heat transfer dynamics throughout the snow season – depending on environmental conditions. The Dynamic
scheme simulates snow thermal conductivity in a range from 0.04 to 0.5 <inline-formula><mml:math id="M91" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">W</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">K</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>, which aligns well with literature estimates of
0.021–0.65 <inline-formula><mml:math id="M92" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">W</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">K</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>. This feature enables the simulation of a wide range of conditions across the Arctic, as opposed to the general
conditions assumed by the Static scheme.</p>
      <p id="d1e2072">The statistical comparison (<italic>site statistics</italic>) shows that there is a smaller variance of modelled values of soil temperature using the
Dynamic snow scheme, which indicates an improvement in comparison to the Static simulations' outputs. The RMSE
(Table <xref ref-type="table" rid="Ch1.T1"/>) also shows that the Dynamic scheme provides an improved fit of simulated soil temperature and snow depth at most
sites. Overall, we conclude that, with the Dynamic scheme, the model is able to simulate snow and soil temperatures that correspond better
with the observed ranges.</p>

<?xmltex \floatpos{t}?><table-wrap id="Ch1.T2" specific-use="star"><?xmltex \currentcnt{2}?><label>Table 2</label><caption><p id="d1e2083">Pan-arctic mean values for the studied variables for the Static and Dynamic simulations and their respective differences.</p></caption><oasis:table frame="topbot"><oasis:tgroup cols="7">
     <oasis:colspec colnum="1" colname="col1" align="left"/>
     <oasis:colspec colnum="2" colname="col2" align="left"/>
     <oasis:colspec colnum="3" colname="col3" align="right"/>
     <oasis:colspec colnum="4" colname="col4" align="right"/>
     <oasis:colspec colnum="5" colname="col5" align="right"/>
     <oasis:colspec colnum="6" colname="col6" align="right"/>
     <oasis:colspec colnum="7" colname="col7" align="left"/>
     <oasis:thead>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">Variable</oasis:entry>
         <oasis:entry colname="col2">Unit</oasis:entry>
         <oasis:entry colname="col3">Static</oasis:entry>
         <oasis:entry colname="col4">Dynamic</oasis:entry>
         <oasis:entry colname="col5">Dynamic <inline-formula><mml:math id="M93" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula> Static</oasis:entry>
         <oasis:entry colname="col6">Abs. diff (%)</oasis:entry>
         <oasis:entry colname="col7">Note of changes</oasis:entry>
       </oasis:row>
     </oasis:thead>
     <oasis:tbody>
       <oasis:row>
         <oasis:entry colname="col1">Snow depth</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M94" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">m</mml:mi></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3">0.36</oasis:entry>
         <oasis:entry colname="col4">0.30</oasis:entry>
         <oasis:entry colname="col5"><inline-formula><mml:math id="M95" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.06</oasis:entry>
         <oasis:entry colname="col6"><inline-formula><mml:math id="M96" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>15</oasis:entry>
         <oasis:entry colname="col7">General decrease</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">ALD</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M97" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">m</mml:mi></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3">0.98</oasis:entry>
         <oasis:entry colname="col4">1.06</oasis:entry>
         <oasis:entry colname="col5">0.07</oasis:entry>
         <oasis:entry colname="col6">7</oasis:entry>
         <oasis:entry colname="col7">Increase in ALD</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Permafrost extent</oasis:entry>
         <oasis:entry colname="col2">10<inline-formula><mml:math id="M98" display="inline"><mml:msup><mml:mi/><mml:mn mathvariant="normal">6</mml:mn></mml:msup></mml:math></inline-formula> <inline-formula><mml:math id="M99" display="inline"><mml:mrow class="unit"><mml:msup><mml:mi mathvariant="normal">km</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msup></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3">1.524</oasis:entry>
         <oasis:entry colname="col4">1.466</oasis:entry>
         <oasis:entry colname="col5"><inline-formula><mml:math id="M100" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.058</oasis:entry>
         <oasis:entry colname="col6">4</oasis:entry>
         <oasis:entry colname="col7">Decrease (TTOP: 13.9)</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><inline-formula><mml:math id="M101" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">SoilT</mml:mi><mml:mi mathvariant="normal">winter</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M102" display="inline"><mml:mrow class="unit"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup><mml:mi mathvariant="normal">C</mml:mi></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3"><inline-formula><mml:math id="M103" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>22.65</oasis:entry>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M104" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>12.63</oasis:entry>
         <oasis:entry colname="col5">10.02</oasis:entry>
         <oasis:entry colname="col6">44</oasis:entry>
         <oasis:entry colname="col7">Increase in <inline-formula><mml:math id="M105" display="inline"><mml:mi>T</mml:mi></mml:math></inline-formula></oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1"><inline-formula><mml:math id="M106" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">SoilT</mml:mi><mml:mi mathvariant="normal">summer</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M107" display="inline"><mml:mrow class="unit"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup><mml:mi mathvariant="normal">C</mml:mi></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3">6.34</oasis:entry>
         <oasis:entry colname="col4">4.46</oasis:entry>
         <oasis:entry colname="col5"><inline-formula><mml:math id="M108" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>1.87</oasis:entry>
         <oasis:entry colname="col6"><inline-formula><mml:math id="M109" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>29</oasis:entry>
         <oasis:entry colname="col7">Decrease in <inline-formula><mml:math id="M110" display="inline"><mml:mi>T</mml:mi></mml:math></inline-formula></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><inline-formula><mml:math id="M111" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">GPP</mml:mi><mml:mi mathvariant="normal">winter</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M112" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">g</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3">0.146</oasis:entry>
         <oasis:entry colname="col4">0.180</oasis:entry>
         <oasis:entry colname="col5">0.035</oasis:entry>
         <oasis:entry colname="col6">24</oasis:entry>
         <oasis:entry colname="col7">Increase in gross production</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><inline-formula><mml:math id="M113" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">GPP</mml:mi><mml:mi mathvariant="normal">summer</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2"/>
         <oasis:entry colname="col3">221.10</oasis:entry>
         <oasis:entry colname="col4">227.72</oasis:entry>
         <oasis:entry colname="col5">6.62</oasis:entry>
         <oasis:entry colname="col6">3</oasis:entry>
         <oasis:entry colname="col7">Increase in gross production</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><inline-formula><mml:math id="M114" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">NPP</mml:mi><mml:mi mathvariant="normal">winter</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2"/>
         <oasis:entry colname="col3"><inline-formula><mml:math id="M115" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>3.05</oasis:entry>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M116" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>5.73</oasis:entry>
         <oasis:entry colname="col5"><inline-formula><mml:math id="M117" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>2.68</oasis:entry>
         <oasis:entry colname="col6"><inline-formula><mml:math id="M118" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>88</oasis:entry>
         <oasis:entry colname="col7">Decrease in productivity</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><inline-formula><mml:math id="M119" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">NPP</mml:mi><mml:mi mathvariant="normal">summer</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2"/>
         <oasis:entry colname="col3">156.15</oasis:entry>
         <oasis:entry colname="col4">156.37</oasis:entry>
         <oasis:entry colname="col5">0.23</oasis:entry>
         <oasis:entry colname="col6">0</oasis:entry>
         <oasis:entry colname="col7">Marginal difference</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><inline-formula><mml:math id="M120" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">Rh</mml:mi><mml:mi mathvariant="normal">winter</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2"/>
         <oasis:entry colname="col3">2.34</oasis:entry>
         <oasis:entry colname="col4">7.77</oasis:entry>
         <oasis:entry colname="col5">5.43</oasis:entry>
         <oasis:entry colname="col6">232</oasis:entry>
         <oasis:entry colname="col7">Increase in Rh</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><inline-formula><mml:math id="M121" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">Rh</mml:mi><mml:mi mathvariant="normal">summer</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2"/>
         <oasis:entry colname="col3">117.81</oasis:entry>
         <oasis:entry colname="col4">103.17</oasis:entry>
         <oasis:entry colname="col5"><inline-formula><mml:math id="M122" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>14.64</oasis:entry>
         <oasis:entry colname="col6"><inline-formula><mml:math id="M123" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>12</oasis:entry>
         <oasis:entry colname="col7">Decrease in Rh</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><inline-formula><mml:math id="M124" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">NEE</mml:mi><mml:mi mathvariant="normal">winter</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2"/>
         <oasis:entry colname="col3">5.49</oasis:entry>
         <oasis:entry colname="col4">13.57</oasis:entry>
         <oasis:entry colname="col5">8.08</oasis:entry>
         <oasis:entry colname="col6">147</oasis:entry>
         <oasis:entry colname="col7">Increased carbon emission</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><inline-formula><mml:math id="M125" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">NEE</mml:mi><mml:mi mathvariant="normal">summer</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2"/>
         <oasis:entry colname="col3"><inline-formula><mml:math id="M126" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>38.32</oasis:entry>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M127" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>53.17</oasis:entry>
         <oasis:entry colname="col5"><inline-formula><mml:math id="M128" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>14.84</oasis:entry>
         <oasis:entry colname="col6"><inline-formula><mml:math id="M129" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>39</oasis:entry>
         <oasis:entry colname="col7">Increased carbon update</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1"><inline-formula><mml:math id="M130" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">NEE</mml:mi><mml:mi mathvariant="normal">annual</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2"/>
         <oasis:entry colname="col3"><inline-formula><mml:math id="M131" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>33.90</oasis:entry>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M132" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>37.16</oasis:entry>
         <oasis:entry colname="col5"><inline-formula><mml:math id="M133" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>3.26</oasis:entry>
         <oasis:entry colname="col6"><inline-formula><mml:math id="M134" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>9.6</oasis:entry>
         <oasis:entry colname="col7">Increased C uptake</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">SoilC</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M135" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kg</mml:mi><mml:mspace width="0.125em" linebreak="nobreak"/><mml:mi mathvariant="normal">C</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3">11.13</oasis:entry>
         <oasis:entry colname="col4">11.12</oasis:entry>
         <oasis:entry colname="col5"><inline-formula><mml:math id="M136" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.01</oasis:entry>
         <oasis:entry colname="col6">0</oasis:entry>
         <oasis:entry colname="col7">Marginal difference</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">VegC</oasis:entry>
         <oasis:entry colname="col2"/>
         <oasis:entry colname="col3">1.86</oasis:entry>
         <oasis:entry colname="col4">2.12</oasis:entry>
         <oasis:entry colname="col5">0.26</oasis:entry>
         <oasis:entry colname="col6">14</oasis:entry>
         <oasis:entry colname="col7">Increase in vegetation C</oasis:entry>
       </oasis:row>
     </oasis:tbody>
   </oasis:tgroup></oasis:table></table-wrap>

</sec>
<sec id="Ch1.S3.SS2">
  <label>3.2</label><title>Russian site simulations</title>
      <p id="d1e2872">Following the Dynamic scheme's improved performance at the site level, we further evaluate the model's performance at the regional scale for
the same sites as in the previous model intercomparison by <xref ref-type="bibr" rid="bib1.bibx53" id="text.38"/> that highlighted shortcomings in the snow scheme of
LPJ-GUESS. Figure <xref ref-type="fig" rid="Ch1.F5"/> shows the snow insulation effect over a set of Russian sites, using the two snow schemes, where the coloured bars
show different temperature regimes. The figure is compiled from 20 winter season average values of near-surface soil temperature (25 <inline-formula><mml:math id="M137" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">cm</mml:mi></mml:mrow></mml:math></inline-formula> depth)
and snow depth per site. Due to the air-temperature-based classification, the number of samples per bin is not balanced, which led to an uneven number
of values allocated to the different groups. The top row of Fig. <xref ref-type="fig" rid="Ch1.F5"/> shows that the Dynamic snow scheme has better skill in
simulating the relationship between soil temperature and snow depth than the Static scheme. It must be noted that there is a clear difference
between the current Static scheme simulations and results reported by <xref ref-type="bibr" rid="bib1.bibx53" id="text.39"/>, which is due to recent updates in the model,
independent of the snow module, and the different climate forcing dataset used in this study.</p>
      <p id="d1e2893">It is apparent from the <inline-formula><mml:math id="M138" display="inline"><mml:mrow><mml:mi mathvariant="normal">Δ</mml:mi><mml:mi>T</mml:mi></mml:mrow></mml:math></inline-formula> and snow depth relationship (Fig. <xref ref-type="fig" rid="Ch1.F5"/>, bottom row) that the Dynamic scheme reproduces the
observed insulation effect well. Unlike the Static scheme, the new snow module can also simulate the different insulation<?pagebreak page5774?> behaviour depending
on the air temperature regimes. The improved performance of the Dynamic scheme is confirmed by the root-mean-squared error (RMSE), shown in
the Supplement (Table S2 in the Supplement). RMSE decreased significantly for both the soil temperature–snow depth and <inline-formula><mml:math id="M139" display="inline"><mml:mrow><mml:mi mathvariant="normal">Δ</mml:mi><mml:mi>T</mml:mi></mml:mrow></mml:math></inline-formula>–snow depth
relationships. This regional analysis confirmed that the new Dynamic snow scheme has an improved skill in simulating winter soil conditions.</p>
</sec>
<sec id="Ch1.S3.SS3">
  <label>3.3</label><title>Pan-Arctic simulations</title>
      <p id="d1e2926">To assess how the two snow schemes differ in simulating seasonal snow across the Arctic, we subtracted output variables from simulations with the
Static module from those with the Dynamic module. We calculated average conditions for winter (December–January–February) and summer
(June–July–August) for the period 1990–2015. The mean pan-arctic seasonal dynamics of snow depth, soil temperature, and upper soil water content are
shown in the Table <xref ref-type="table" rid="Ch1.T2"/>.</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F6" specific-use="star"><?xmltex \currentcnt{6}?><?xmltex \def\figurename{Figure}?><label>Figure 6</label><caption><p id="d1e2933"><bold>(a)</bold> Snow depth difference in winter months and <bold>(b)</bold> maximum ALD difference, calculated by subtracting the Static from Dynamic simulation outputs. Modelled permafrost extent is based on mean annual ground temperature (MAGT) and plotted against the permafrost cover estimate by <xref ref-type="bibr" rid="bib1.bibx38" id="text.40"/> (TTOP model). Simulated absolute snow depth is shown in Fig. S2 in the Supplement and ALD in Fig. S3 in the Supplement.</p></caption>
          <?xmltex \igopts{width=483.69685pt}?><graphic xlink:href="https://bg.copernicus.org/articles/18/5767/2021/bg-18-5767-2021-f06.png"/>

        </fig>

<sec id="Ch1.S3.SS3.SSS1">
  <label>3.3.1</label><title>Impacts on physical variables</title>
      <p id="d1e2957">Figure <xref ref-type="fig" rid="Ch1.F6"/>a shows the difference in simulated wintertime snow depth. The Dynamic scheme shows an overall lower snow depth across the
Arctic with the most pronounced changes in coastal Norway and in western Siberia. On average, the snow depth for the Dynamic scheme is
6 <inline-formula><mml:math id="M140" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">cm</mml:mi></mml:mrow></mml:math></inline-formula> lower due to the implementation of snow-related processes affecting snow density and consequently snow depth.</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F7" specific-use="star"><?xmltex \currentcnt{7}?><?xmltex \def\figurename{Figure}?><label>Figure 7</label><caption><p id="d1e2972">Near-surface soil temperature (25 <inline-formula><mml:math id="M141" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">cm</mml:mi></mml:mrow></mml:math></inline-formula> depth) difference between the Dynamic and Static simulations, for winter <bold>(a)</bold> and summer <bold>(b)</bold> seasons. Differences are calculated by subtracting the Static from Dynamic simulation outputs. The absolute simulated soil temperatures using the two snow schemes are shown in the Supplement in Fig. S4.</p></caption>
            <?xmltex \igopts{width=312.980315pt}?><graphic xlink:href="https://bg.copernicus.org/articles/18/5767/2021/bg-18-5767-2021-f07.png"/>

          </fig>

      <p id="d1e2995">The main aim of developing the new snow scheme was not only to enhance the simulation of snow depth but also to improve the simulation of snowpack
properties that directly affect soil conditions. Therefore, we investigated how the internal structural changes in the representation of snow
influenced soil temperatures. The soil temperature differences shown in Fig. <xref ref-type="fig" rid="Ch1.F7"/> reveal that the new snow scheme influenced the winter
season to a large degree, both within and especially outside of the permafrost region. Winter soil temperatures are higher with the Dynamic
scheme, while it results in a cooler near-surface soil temperature during the summer. A closer look at the monthly soil temperature values in
Fig. <xref ref-type="fig" rid="Ch1.F11"/> showed that spring months are cooler for the Dynamic scheme but that the difference between the two schemes
decreases towards the end of summer. This pattern is more pronounced in the permafrost-underlain regions. This shows that the Static snow
scheme has too little insulation and results in soil temperatures that are too cold during the winter months, as we also show in the site
simulations. Moreover, the Static scheme also does not insulate soils sufficiently during the springtime when air temperatures rise above
0 <inline-formula><mml:math id="M142" display="inline"><mml:mrow class="unit"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup><mml:mi mathvariant="normal">C</mml:mi></mml:mrow></mml:math></inline-formula>, which allows the soil to warm up more quickly even in the presence of a snowpack.</p>
      <p id="d1e3015">The depth to which the top soil thaws during summer, and refreezes in winter, in permafrost areas is called the active layer depth (ALD). The
difference in the seasonal maximum active layer depth for the model simulations is shown in Fig. <xref ref-type="fig" rid="Ch1.F6"/>b. Since the Dynamic scheme had
warmer soil temperatures, the modelled permafrost extent is smaller than with the Static scheme. We compared our model simulations with a
recent satellite-driven permafrost extent estimate by <xref ref-type="bibr" rid="bib1.bibx38" id="text.41"/> – from here on referred to as the TTOP model. Modelled permafrost extent was
defined by the area where the mean annual ground temperature (25 <inline-formula><mml:math id="M143" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">cm</mml:mi></mml:mrow></mml:math></inline-formula> depth) was below zero. The Dynamic scheme's permafrost extent is
much closer to the TTOP model's estimate, while the Static scheme simulates a much larger permafrost extent, as shown in
Fig. <xref ref-type="fig" rid="Ch1.F6"/>c. The Dynamic scheme's computed areal permafrost<?pagebreak page5775?> cover, while improved compared to the Static scheme, still
overestimates the TTOP model estimates by approximately 5 % (see Table <xref ref-type="table" rid="Ch1.T2"/>).</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F8" specific-use="star"><?xmltex \currentcnt{8}?><?xmltex \def\figurename{Figure}?><label>Figure 8</label><caption><p id="d1e3037">Mean fractional water content of the upper soil column in May, June, and July, using the Static <bold>(a)</bold> and Dynamic <bold>(b)</bold> schemes and their difference <bold>(c)</bold>. Differences are calculated by subtracting the Static from Dynamic simulation outputs.</p></caption>
            <?xmltex \igopts{width=483.69685pt}?><graphic xlink:href="https://bg.copernicus.org/articles/18/5767/2021/bg-18-5767-2021-f08.png"/>

          </fig>

      <p id="d1e3055">Besides governing the physical state of permafrost, snow and soil temperature also have a large influence on the temporal and spatial patterns of soil
water content.  We show the simulated upper soil column water content in Fig. <xref ref-type="fig" rid="Ch1.F8"/>. This figure shows the mean fractional soil water
content over May–June–July. Soil water content was calculated using the average seasonal liquid water content from the top 50 <inline-formula><mml:math id="M144" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">cm</mml:mi></mml:mrow></mml:math></inline-formula> of the soil
column. Soil water content is represented as the fraction of the available capacity between the wilting point and field capacity, and therefore frozen
water is not included in these values. Figure <xref ref-type="fig" rid="Ch1.F8"/>c shows that there is a higher water availability within the permafrost region using the
Static scheme. Water availability is a key driver of the start of the growing season, nutrient availability, and vegetation dynamics. The
time-series analysis of upper soil water content highlights that the snowmelt rate is not significantly different between the schemes. Still, there is
a large difference in soil temperature dynamics. The Static scheme's soil temperature increases more rapidly during the spring than the
Dynamic scheme's soil temperature (see Fig. <xref ref-type="fig" rid="Ch1.F11"/>a and b). This results in an earlier onset of snowmelt and earlier increase
in soil water availability and nitrogen mineralization. This affects productivity, which we assess in the coming sections. Although the<?pagebreak page5776?> difference in
water content and nitrogen mineralization between the snow schemes converges towards zero as summer progresses, we show that the change in snow scheme
had a lasting effect beyond the cold season.</p>
      <p id="d1e3072">Overall, the new snow scheme had a substantial effect on winter soil temperatures. As a result, summer conditions were also altered by the snow scheme
updates. It is apparent that the largest changes in snow depth and temperature coincide. For instance, along the Norwegian coast and in central
Siberia. Taken together, our results show that the Dynamic snow scheme improved the simulation of physical variables.</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F9" specific-use="star"><?xmltex \currentcnt{9}?><?xmltex \def\figurename{Figure}?><label>Figure 9</label><caption><p id="d1e3077">The difference in simulated soil <bold>(a)</bold> and vegetation carbon pools <bold>(b)</bold> between the two schemes. Differences are calculated by subtracting the Static from Dynamic simulation outputs.</p></caption>
            <?xmltex \igopts{width=312.980315pt}?><graphic xlink:href="https://bg.copernicus.org/articles/18/5767/2021/bg-18-5767-2021-f09.png"/>

          </fig>

      <?xmltex \floatpos{p}?><fig id="Ch1.F10" specific-use="star"><?xmltex \currentcnt{10}?><?xmltex \def\figurename{Figure}?><label>Figure 10</label><caption><p id="d1e3095">The difference in normalized heterotrophic respiration for winter <bold>(a)</bold> and summer <bold>(b)</bold> between the two schemes. Differences between simulated Dynamic and Static winter <bold>(c)</bold> and summer <bold>(d)</bold> NPP. Differences between simulated Dynamic and Static winter <bold>(e)</bold> and summer <bold>(f)</bold> NEE. Differences are calculated by subtracting the Static from Dynamic simulation outputs. Simulations for the two schemes are shown in Figs. S5 (heterotrophic respiration), S6 (NPP), and S7 (NEE) in the Supplement.</p></caption>
            <?xmltex \igopts{width=341.433071pt}?><graphic xlink:href="https://bg.copernicus.org/articles/18/5767/2021/bg-18-5767-2021-f10.png"/>

          </fig>

</sec>
<sec id="Ch1.S3.SS3.SSS2">
  <label>3.3.2</label><title>Impacts on biogeochemical variables</title>
      <p id="d1e3132">Besides the impact on soil thermodynamics, we investigated how key biogeochemical components – such as productivity and carbon pools – were
affected. The changes across seasons and permafrost conditions are summarized in Figs. <xref ref-type="fig" rid="Ch1.F11"/> and <xref ref-type="fig" rid="Ch1.F12"/>.</p>
      <p id="d1e3139">Our simulated soil carbon pools (Fig. S6 in the Supplement) deviate from literature values <xref ref-type="bibr" rid="bib1.bibx20" id="paren.42"/> and are consistently lower across the
Arctic. The main reason for this is the model's representation of soil organic matter processes. Soil carbon and nitrogen are represented by pools
that exist in the top 50 <inline-formula><mml:math id="M145" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">cm</mml:mi></mml:mrow></mml:math></inline-formula> of the soil column <xref ref-type="bibr" rid="bib1.bibx48" id="paren.43"/> and are thus only influenced by near-surface conditions. Moreover, peatlands
are not explicitly represented. The differences in soil carbon between the schemes, as shown in Fig. <xref ref-type="fig" rid="Ch1.F9"/>a, coincide spatially with the
highest differences in soil temperature. This suggests that the changes in soil temperature influence soil carbon in the model and therefore the rate
of respiration from soils as well. Vegetation carbon pools (Fig. <xref ref-type="fig" rid="Ch1.F9"/>b) are higher in the non-permafrost region using the Dynamic
snow scheme (see Table <xref ref-type="table" rid="Ch1.T2"/> for mean values). Since the evaluation of soil carbon is not the focus of this study, soil carbon outputs were
used to normalize the heterotrophic respiration to be able to interpret the relative differences between schemes (Fig. <xref ref-type="fig" rid="Ch1.F10"/>a and b). To do
so, we divided the heterotrophic respiration by the soil carbon estimates for the respective simulations using the two snow schemes.  With the
Dynamic scheme, summer soil respiration decreased across the Arctic. Winter respiration, on the other hand, increased, except for in
eastern Siberia. These changes in soil respiration can be attributed to changes in soil temperature, as shown in Fig. <xref ref-type="fig" rid="Ch1.F7"/>.</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F11" specific-use="star"><?xmltex \currentcnt{11}?><?xmltex \def\figurename{Figure}?><label>Figure 11</label><caption><p id="d1e3169">Seasonal dynamics of snow depth, soil temperature (25 <inline-formula><mml:math id="M146" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">cm</mml:mi></mml:mrow></mml:math></inline-formula> depth), and fractional soil water content within <bold>(a)</bold> and outside of the permafrost region <bold>(b)</bold>. Seasonal dynamics of NEE and NPP within <bold>(c)</bold> and outside of the permafrost region <bold>(d)</bold>.</p></caption>
            <?xmltex \igopts{width=426.791339pt}?><graphic xlink:href="https://bg.copernicus.org/articles/18/5767/2021/bg-18-5767-2021-f11.png"/>

          </fig>

      <p id="d1e3199">The difference in net primary productivity (NPP) between simulations with the two snow schemes for both winter and summer is shown in
Fig. <xref ref-type="fig" rid="Ch1.F10"/>c and d, where positive NPP means more carbon uptake by the vegetation. We note an impact of the different snow schemes on summer
productivity, caused by the different soil thermodynamics and soil water availability during the spring and early summer period. This artefact is also
visible in the simulated pan-Arctic NEE, in Fig. <xref ref-type="fig" rid="Ch1.F10"/>e and f, where negative NEE values indicate a stronger uptake of carbon by
ecosystems. The positive difference in winter NEE (e) shows that there is a higher carbon release in the winter season for the Dynamic scheme
in central Europe, western Siberia, and coastal Norway. The mean winter NEE of the Dynamic scheme more than doubled
(Table <xref ref-type="table" rid="Ch1.T2"/>). Compared to the Static scheme, which relates to both the change in soil respiration and
NPP. Figure <xref ref-type="fig" rid="Ch1.F11"/>c and d show some interesting contrasts regarding the seasonal carbon fluxes. Permafrost-underlain regions
(Fig. <xref ref-type="fig" rid="Ch1.F10"/>c) experience little difference in the simulated NEE. The Dynamic scheme simulates lower peak summer NPP. Winter NEE in
the non-permafrost region is higher using the Dynamic scheme, indicating a larger carbon uptake by the vegetation. On the other hand, we can
observe an increased sink capacity (more negative NEE) during the summer months.</p>
</sec>
<?pagebreak page5777?><sec id="Ch1.S3.SS3.SSS3">
  <label>3.3.3</label><title>Vegetation composition and distribution</title>
      <p id="d1e3220">Vegetation composition and distribution depend on the changes in physical and biochemical variables described in the previous section. Therefore, we
investigated how the application of the two snow schemes affected vegetation distribution to determine if there are shifts in dominant plant
functional types (PFTs) as a result of using different snow schemes. The dominant PFT for each simulated grid cell was determined by selecting the PFT
with the highest maximum leaf area index (LAI) during the simulation years (1990–2015). Using the Dynamic snow scheme, roughly half of the sites are dominated by
summergreen low shrubs and boreal needle-leaved evergreen trees (LSS with 25 % and BNE with 23 %; see Table S3). Prostrate dwarf shrubs,
(SPDS), graminoid and forb tundra (GRT), and boreal needle-leaved summergreen trees (BNS) accounted for 20 %, 8 %, and 7 % dominance,
respectively. For an easier comparison between the Static and Dynamic simulations, PFT classes were grouped into forest, open grass,
shrubs, and no vegetation categories after determining the dominant PFT in each grid cell (classification based on <xref ref-type="bibr" rid="bib1.bibx55" id="altparen.44"/>). This
classification showed that grassland classes dominate (56 %), followed by forest cover (36 %). Shrubs dominate at 29 % of the simulation
sites. There is a negligible number of sites with mostly bare soil. When comparing the spatial pattern of dominant vegetation groups, we noted that
there is only a marginal difference between the Static and Dynamic simulations (see Fig. S10 in the Supplement). Changes in group
dominance between the Static and Dynamic simulations occurred at approximately 10 % of the sites; see Fig. S11 in the
Supplement. The Sankey diagram shows the direction of change between the three groups. <xref ref-type="bibr" rid="bib1.bibx34" id="text.45"/> suggest that increased soil temperature leads
to a shift to an increased forest height and shrub cover. Grass-to-shrub domination change (3.1 %) was the most prevailing change, which indeed
points towards an increase in vegetation height. However, we observe a shrub-to-grass (1.95 %) shift in domination at the same time; therefore, we
cannot conclude the main direction of changes.</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F12" specific-use="star"><?xmltex \currentcnt{12}?><?xmltex \def\figurename{Figure}?><label>Figure 12</label><caption><p id="d1e3231">Relationship between variables during the winter and summer seasons, within and outside of the permafrost region. The colour of the boxes indicates the direction and qualitative magnitude of changes in the variables, based on the relative difference (%) between Dynamic and Static schemes – shown in the boxes below the variables. Variables are shown as follows: GPP: gross primary production (<inline-formula><mml:math id="M147" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">g</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>); NPP: net primary productivity (<inline-formula><mml:math id="M148" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">g</mml:mi><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>); NEE: net ecosystem exchange (<inline-formula><mml:math id="M149" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">g</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>); <inline-formula><mml:math id="M150" display="inline"><mml:mrow><mml:msub><mml:mi>R</mml:mi><mml:mi mathvariant="normal">a</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>: autotrophic respiration (<inline-formula><mml:math id="M151" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">g</mml:mi><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>); <inline-formula><mml:math id="M152" display="inline"><mml:mrow><mml:msub><mml:mi>R</mml:mi><mml:mi mathvariant="normal">h</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>: heterotrophic respiration (<inline-formula><mml:math id="M153" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">g</mml:mi><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>); <inline-formula><mml:math id="M154" display="inline"><mml:mrow><mml:msub><mml:mi>N</mml:mi><mml:mo>min⁡</mml:mo></mml:msub></mml:mrow></mml:math></inline-formula>: net nitrogen mineralization (<inline-formula><mml:math id="M155" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kg</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:mi mathvariant="normal">N</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">ha</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>); soil water content: <italic>fraction</italic>. The relative changes in soil water content (in %) are high in the winter due to the low fractional water content values as we only account for the liquid soil water and do not consider the amount of frozen water in the soil. These changes correspond to small absolute changes in fractional soil water content. <inline-formula><mml:math id="M156" display="inline"><mml:msup><mml:mi/><mml:mo>∗</mml:mo></mml:msup></mml:math></inline-formula> Note: temperature differences are shown as absolute differences for easier readability.</p></caption>
            <?xmltex \igopts{width=398.338583pt}?><graphic xlink:href="https://bg.copernicus.org/articles/18/5767/2021/bg-18-5767-2021-f12.png"/>

          </fig>

</sec>
</sec>
<sec id="Ch1.S3.SS4">
  <label>3.4</label><title>Cause-and-effect relationships</title>
      <p id="d1e3400">As many of the reviewed processes interact with each other in a complex, non-linear manner, change in one variable may not translate to a direct
impact on another variable. To provide an overview of our findings regarding the physical and biogeochemical processes, we created a flow chart
showing observed changes in modelled state variables and their connections. Figure <xref ref-type="fig" rid="Ch1.F12"/> shows the difference between simulations using the
two snow schemes – calculated by subtracting the Static from Dynamic results. Reddish box colours show that the Dynamic
scheme had higher values, and blueish colours show that the Dynamic scheme simulated lower values than the Static scheme. The
lightness and darkness of the colours indicate the magnitude of changes between the winter and summer seasons qualitatively.</p>
      <?pagebreak page5779?><p id="d1e3405">Each box contains the computed difference, and Table <xref ref-type="table" rid="Ch1.T2"/> summarizes the mean changes in these key variables. Considering the spatial pattern
across the Arctic, we conclude that the pattern of changes and differences between the Static and Dynamic simulations vary depending
on the presence or absence of permafrost cover. For a more detailed evaluation, process relationships are therefore divided into permafrost and
non-permafrost regions. Since snow depth only affects these variables indirectly, through insulation, it was not included in the feedback graph. The
choice of snow scheme induced changes in near-surface temperature (<inline-formula><mml:math id="M157" display="inline"><mml:mi>T</mml:mi></mml:math></inline-formula>), which is the key governing factor over these variables. In general, higher
soil temperatures during the winter season prompt a positive response in respiration, soil water content, and vegetation primary productivity. Soil
temperature increased to a greater degree in the non-permafrost region during the winter season. The same increasing pattern is observed for
heterotrophic (<inline-formula><mml:math id="M158" display="inline"><mml:mrow><mml:msub><mml:mi>R</mml:mi><mml:mi mathvariant="normal">h</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>) and autotrophic respiration (<inline-formula><mml:math id="M159" display="inline"><mml:mrow><mml:msub><mml:mi>R</mml:mi><mml:mi mathvariant="normal">a</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>), soil water content, and NEE. Nitrogen mineralization decreased in the
wintertime, with a larger decrease outside of the permafrost region. In contrast, summer months' average soil temperature showed an overall decrease
in the permafrost region. This is due to the different thermal soil and snow dynamics in the two applied snow schemes. We observed rapid heat loss
using the Static scheme, resulting in insufficient insulation during the snow season. The same feature causes soil temperature to rise
rapidly in the spring, when air temperature is already above zero but a snow cover remains present. Faster soil warming leads to increased soil water
availability that affects productivity. Respiration and NEE are slightly reduced for both permafrost and non-permafrost regions in the
Dynamic scheme during the summer. Differences noted for the summer are smaller for all variables than in the winter season.</p>
</sec>
</sec>
<sec id="Ch1.S4">
  <label>4</label><title>Discussion</title>
<sec id="Ch1.S4.SS1">
  <label>4.1</label><title>Snow scheme dynamics</title>
      <p id="d1e3456">The site-level analysis shows that the new Dynamic scheme is able to simulate snow height and density adequately due to the implementation of
physical processes and a dynamic representation of snow properties. The integrated mechanistic compaction scheme and phase changes within snow layers
make it possible to simulate heterogeneous snow density and thermal properties within the snowpack. This influences the simulated snow density
directly by altering snowpack structure. Density regulates heat transport rate through snow layers by affecting thermal conductivity
(Eq. <xref ref-type="disp-formula" rid="Ch1.E7"/>): lower density results in a more insulating cover, whereas higher density and compacted layers are a better heat-transferring
medium and exhibit lower insulation. In the Static scheme, snow density was assumed constant through the snow season and across all study
sites. Such static snow representation is unsatisfactory when simulating arctic conditions <xref ref-type="bibr" rid="bib1.bibx24" id="paren.46"/>. The new snow scheme provides an improved
framework for a mechanistic snow season simulation.</p>
      <p id="d1e3464">The single site simulations (Sect. 3.1) provide reasonably consistent evidence that the new snow scheme's implementation leads to significant changes
in near-surface temperature simulation – especially at Abisko, Bayelva, and Samoylov. As shown by <xref ref-type="bibr" rid="bib1.bibx11" id="text.47"/>, the site-wise model–data
comparison is challenging since point measurements may not be representative of a larger area due to the complexity in topography and vegetation
conditions. The model–observation fit may be improved by using site-specific climatic forcing instead of a global gridded dataset.</p>
      <?pagebreak page5780?><p id="d1e3470">To avoid site-specific problems in the interpretation of simulations, we also evaluated the model at a regional scale. By comparing the results of the
Russian site simulations (Sect. 3.2) with those of <xref ref-type="bibr" rid="bib1.bibx53" id="text.48"/>, we conclude that the development of the representation of snow in LPJ-GUESS
significantly improved air–soil temperature and snow depth–soil temperature relationships. The Dynamic snow scheme's insulation capacity
followed a quasi asymptotic trend, increasing with snow depth and slightly levelling out after reaching the so-called effective depth at
30–40 <inline-formula><mml:math id="M160" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">cm</mml:mi></mml:mrow></mml:math></inline-formula> <xref ref-type="bibr" rid="bib1.bibx46" id="paren.49"/>. The insulation capacity was, in general, slightly lower than observations, with a notable underestimation when
snow depth is below 20 <inline-formula><mml:math id="M161" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">cm</mml:mi></mml:mrow></mml:math></inline-formula>. Nonetheless, these results are a vast improvement over the old Static scheme, as shown in
Fig. <xref ref-type="fig" rid="Ch1.F5"/>.</p>
      <p id="d1e3497">The RMSE values (Table S2) also show that the Dynamic scheme better captured the observed the soil temperature and snow depth relationship than
the Static scheme. RMSE was slightly higher for the coldest air temperature regime for both snow schemes. We note that the Static
schemes' performance differs from what was shown in <xref ref-type="bibr" rid="bib1.bibx53" id="text.50"/>. The reasons for these differences are developments of the model since then in
components other than the snow scheme and also the different meteorological forcing used in this study. Our results indicate that the enhancement of
snow-related processes improves the simulation of soil temperatures in LPJ-GUESS and that the model can be more reliably applied to assess the impact
of environmental changes on the arctic carbon cycle.</p>
</sec>
<sec id="Ch1.S4.SS2">
  <label>4.2</label><title>Impact on physical and biogeochemical variables</title>
      <p id="d1e3511">The changes to the snow insulation capacity in the Dynamic scheme had a significant effect on permafrost conditions. Our pan-Arctic results
showed that the Static scheme simulated near-surface soil temperatures that were too cold in winter and too warm in summer. Permafrost extent
simulated with the Dynamic scheme agreed more closely with the permafrost estimate by <xref ref-type="bibr" rid="bib1.bibx38" id="text.51"/>, as shown in
Fig. <xref ref-type="fig" rid="Ch1.F6"/>. Comparison of these findings with other studies where a new snow scheme was introduced reiterates that the model representation of
snow strongly affects soil temperatures <xref ref-type="bibr" rid="bib1.bibx17 bib1.bibx52" id="paren.52"/>. Reliable soil temperature simulations are essential to study the
permafrost–climate feedback. <xref ref-type="bibr" rid="bib1.bibx6" id="text.53"/> concluded that<?pagebreak page5781?> recent warming trends of permafrost soils are partly due to an increase in snow
insulation, accelerating its degradation. Both field observations and modelling studies have identified this close link between snow and permafrost
conditions <xref ref-type="bibr" rid="bib1.bibx23 bib1.bibx30 bib1.bibx25" id="paren.54"/>. Identifying changes in permafrost-underlain areas is important because of the potential
increase in organic matter decomposition and release of greenhouse gases. These aspects will be further evaluated in LPJ-GUESS with the new snow
scheme.</p>
      <p id="d1e3528">We observed a general decrease in mean NPP during the winter and a marginal difference in the summer. Considering the presence of permafrost, however,
we noted an increase in GPP and NPP for non-permafrost-underlain areas in summer. The significantly warmer winter soil conditions for the
Dynamic scheme caused an increase in heterotrophic respiration – i.e due to faster litter decomposition rates and increased microbial
activity. Accordingly, soil respiration increased during the winter in the non-permafrost region. During the summer, there is an overall minor
decrease in soil respiration due to the lower soil temperatures simulated by the Dynamic scheme. The net effect of the above-discussed
processes is an overall increase in carbon emissions during the winter and an increased uptake during the summer.</p>
      <p id="d1e3531">The impact of the new snow scheme on summer conditions was surprising. These differences were caused by the changes in spring snow and soil
temperatures and soil water availability. During springtime, soils with the Static scheme warm more quickly, due to the lower insulation,
which leads to an earlier thaw and increased soil water availability. The Dynamic scheme simulates a more realistic atmosphere–snow–soil heat
transfer, leading to a slower temperature transition. The difference between the schemes diminishes towards the end of the summer.  Overall, the
simulated pan-arctic carbon fluxes are systematically lower than other published values <xref ref-type="bibr" rid="bib1.bibx13 bib1.bibx44 bib1.bibx29" id="paren.55"/>. <xref ref-type="bibr" rid="bib1.bibx51" id="text.56"/> estimate an annual NEE in the range of <inline-formula><mml:math id="M162" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>46 to <inline-formula><mml:math id="M163" display="inline"><mml:mo>+</mml:mo></mml:math></inline-formula>10 <inline-formula><mml:math id="M164" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">g</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:mi mathvariant="normal">C</mml:mi><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:msup><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">yr</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>, while our simulations have
a mean estimate around <inline-formula><mml:math id="M165" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>35.5 <inline-formula><mml:math id="M166" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">g</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:mi mathvariant="normal">C</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:msup><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">yr</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula> (see Table <xref ref-type="table" rid="Ch1.T2"/>).</p>
      <p id="d1e3622">Besides the carbon fluxes, we also evaluated the simulated annual soil and vegetation carbon pools. Vegetation pools were more different when
applying the Dynamic, while no clear differences were apparent for soil carbon (see Table <xref ref-type="table" rid="Ch1.T2"/>). With the
Dynamic snow scheme, the soil carbon pool is lower within the permafrost region and higher outside of the permafrost region. These results
align well with the sensitivity study by <xref ref-type="bibr" rid="bib1.bibx17" id="text.57"/>, which highlighted that decreased soil carbon stocks can be attributed to a higher
respiration rate and increased microbial decomposition rates.</p>
      <p id="d1e3631">The mean simulated soil carbon content was around 10 <inline-formula><mml:math id="M167" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kg</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:mi mathvariant="normal">C</mml:mi><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula>, which is much lower than the 50–100 <inline-formula><mml:math id="M168" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kg</mml:mi><mml:mspace width="0.125em" linebreak="nobreak"/><mml:mi mathvariant="normal">C</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula> range suggested in literature <xref ref-type="bibr" rid="bib1.bibx20 bib1.bibx19" id="paren.58"/>. This difference is most likely due to the fact that we only simulated grid cells with upland soils,
while peatlands were not represented. The inclusion of peatlands would have led to a larger amount of soil carbon, since these ecosystems are
characterized by waterlogged soils in which decomposition is suppressed – although carbon can be released as methane. Also, all organic matter is
considered to be in the top 0.5 <inline-formula><mml:math id="M169" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">m</mml:mi></mml:mrow></mml:math></inline-formula> of the soil in the current version of LPJ-GUESS and is therefore only affected by the average soil
temperature and moisture conditions down to 0.5 <inline-formula><mml:math id="M170" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">m</mml:mi></mml:mrow></mml:math></inline-formula>, but not by conditions further down. These aspects will be taken into account in ongoing
model development. Our analysis highlights that the observed differences between the Static and Dynamic schemes correlate well with
the spatial pattern of near-surface soil temperature changes. This shows that the changes in soil temperature influence the soil carbon content in the
model. The shortcomings in soil carbon simulation will be addressed and improved in the future, which will enable a more reliable carbon pool
assessment.</p>
</sec>
<sec id="Ch1.S4.SS3">
  <label>4.3</label><title>Impact on vegetation dynamics</title>
      <p id="d1e3701">Satellite-based studies have identified an overall greening trend across the Arctic, in response to a warming from the 1980s until now. However, they
also showed that this greening trend is not uniform and certain areas have actually experienced browning (loss of greenness) during this period
<xref ref-type="bibr" rid="bib1.bibx3 bib1.bibx35" id="paren.59"/>. This may be partly due to damage to vegetation following extreme winter events <xref ref-type="bibr" rid="bib1.bibx42" id="paren.60"/>. At the site level, a
recent study by <xref ref-type="bibr" rid="bib1.bibx37" id="text.61"/> showed that winter thermal conditions are a strong control on vegetation patterns in arctic landscapes. Still,
it is challenging to fully understand vegetation responses to warming solely from remotely sensed data or field observations, due to the scale
dependency of interpreting trends in vegetation dynamics. Moreover, most field sites are highly concentrated in northern Scandinavia and Alaska, which
leaves the full heterogeneity of the arctic and its ecosystems vastly under-sampled <xref ref-type="bibr" rid="bib1.bibx32" id="paren.62"/>. With ecosystem models, we can fill in spatial
gaps, identify feedback loops, and assess potential future changes.</p>
      <p id="d1e3716">Following the assessment of the new snow scheme's impact on biogeochemical variables, we compiled the simulated vegetation conditions with the two
snow schemes. We found that PFT domination changed marginally using the updated snow scheme in some grid cells. The main direction of change is a grass-to-shrub dominance shift in grid cells – shown in Fig. S11. The forest–shrub border did not shift much in most areas. However, the vegetation carbon
pool was higher with the Dynamic scheme, which indicates that even though the changes are minor visually, they affected vegetation
biomass. These comparisons show that changing the snow scheme in LPJ-GUESS affected vegetation distribution and composition, albeit on a small scale.</p>
</sec>
<?pagebreak page5782?><sec id="Ch1.S4.SS4">
  <label>4.4</label><title>Outlook</title>
      <p id="d1e3727">It is well established that the Arctic is highly susceptible to climate change, and the ongoing warming has significant consequences for the arctic
system – even if we implement the most strict mitigation measures <xref ref-type="bibr" rid="bib1.bibx10 bib1.bibx1" id="paren.63"/>. One of these consequences is a change in snow
conditions. In the near future, snow thickness will decrease, caused by air temperature and precipitation changes, inducing a decrease in snow-covered
area in the region <xref ref-type="bibr" rid="bib1.bibx1 bib1.bibx21" id="paren.64"/>. Due to a later onset and earlier spring melt, the snow season is expected to shorten under a changing
climate. Moreover, northern high latitudes are predicted to be rain-dominated in the future <xref ref-type="bibr" rid="bib1.bibx5 bib1.bibx22" id="paren.65"/>. These changes will
strongly influence soil thermodynamics, and the observed and projected changes will have a significant impact on arctic ecosystems
<xref ref-type="bibr" rid="bib1.bibx10" id="paren.66"/>. To be able to provide robust projections of the future, we need to account for a multitude of interlinked processes and
feedbacks. Some of the current key areas are to assess the relative sensitivity of plant productivity to climate change, the development of
decomposition rates, and their net effect on the carbon budget.</p>
      <p id="d1e3742">Besides an assessment of geophysical and biogeochemical processes, LPJ-GUESS can also be used to explore future vegetation trends to assess whether
favourable growing conditions will induce further greening or whether new stressors will prompt local- to regional-scale browning (loss of
productivity) <xref ref-type="bibr" rid="bib1.bibx35" id="paren.67"/>. Studying future vegetation trends across the Arctic is important from a global perspective. A potential decrease in
snow-covered area may significantly decrease surface albedo, which would enhance arctic warming. Consequently, changes in snow dynamics on a local
scale influence carbon fluxes by altering soil thermal conditions and vegetation habitat. Evaluating snow–soil–vegetation feedbacks in future studies
is therefore relevant to further investigate climate change impacts on the Arctic in global-scale land surface modelling.</p>
</sec>
</sec>
<sec id="Ch1.S5" sec-type="conclusions">
  <label>5</label><title>Conclusions</title>
      <p id="d1e3757">This study shows that the representation of snow dynamics in a dynamic vegetation model significantly influences the simulated soil thermodynamics and
related biogeochemical variables. We show, due to the improved snow insulation capacity, that the new Dynamic snow scheme simulates more
realistic soil thermodynamics and permafrost extent than the old Static scheme. The improved simulation of permafrost cover can be attributed
to significantly warmer winter soil temperatures, which compare well to observations across 256 locations in Russia. We further showed the importance
of an accurate snow scheme for the simulation of biogeochemical processes. Our results show that the intermediate-complexity snow scheme had a
significant impact on carbon fluxes. Heterotrophic respiration increased during the winter, which led to an increased carbon release during the cold
season. We also identified differences in soil carbon content between the Static and Dynamic simulations. Although the modelled soil
carbon content was lower than literature values, the spatial pattern of low and high soil carbon content aligns well with observations. A
differentiation between the seasons and accounting for permafrost presence highlighted the differences between the two sets of simulations. Wintertime
carbon emissions were higher using the Dynamic scheme, both within but especially outside of the permafrost region. The differences between
the simulations were larger within permafrost-underlain areas for the physical variables. Besides spatial patterns, we explored seasonal differences,
which showed that summertime conditions were also affected by the representation of snow. In contrast to warmer soils in winters, soils were cooler in
summer using the Dynamic scheme – especially in permafrost-underlain areas – due to a delayed response to snowmelt. These differences
between the old and new snow schemes underline the importance of further developing winter processes as they may significantly affect the annual
carbon budget.</p>
      <p id="d1e3760">These findings contribute to our understanding of the impact of wintertime changes on the arctic carbon cycle. We show that an accurate, dynamic snow
scheme is essential to investigate the full complexity of snow–soil–vegetation relationships. Models are valuable tools to aid our understanding of
large-scale climate change impacts due to the sparse availability of observations in the Arctic. Addressing identified knowledge gaps in models is
imperative to decrease the uncertainty around carbon balance estimates. Due to the large spread of observed and modelled seasonal and inter-annual
cycles of carbon fluxes, it is not yet possible to determine with high certainty whether the Arctic will act as a carbon source or sink in the future
<xref ref-type="bibr" rid="bib1.bibx15 bib1.bibx31" id="paren.68"/>. To decrease uncertainty in simulations, contemporary modelling efforts are directed, on the one hand, at model
inclusion (account for key, still missing processes) and, on the other hand, at refining process formulations using observational data
<xref ref-type="bibr" rid="bib1.bibx29 bib1.bibx15 bib1.bibx14" id="paren.69"/>.</p>
      <?pagebreak page5783?><p id="d1e3769">In this study, we aimed to improve the representation of the cold-season process using non-growing-season observations and findings. This enhances the
versatility and applicability of LPJ-GUESS as a tool to address the remaining uncertainties regarding climate change impacts at northern high
latitudes and its consequences on a global scale. With this model, we have the ability to investigate complex ecosystem interactions under changing
environmental conditions at multiple scales, considering nitrogen cycling, permafrost processes (freeze–thaw cycles, hydrology), stochastic vegetation
dynamics, and also potential land cover and land use changes. Realistic soil temperature simulations are the first step to improve the simulation
of greenhouse gas emissions under different climate scenarios across the Arctic <xref ref-type="bibr" rid="bib1.bibx36" id="paren.70"/>. Our results show that by improving a process that
appears only relevant in winter, such as snow, we not only decrease the uncertainty regarding physical and biogeochemical parameters during the cold
season, but we also improve simulations of soil conditions and the carbon cycle in the growing season. Further developments will aim at improving soil
carbon content simulations and better assessing plant responses to future environmental conditions during the cold season. By accounting for
snow–soil–vegetation interactions in all seasons of the year, we ensure more reliable projections of the future state of vegetation composition,
permafrost stability, and greenhouse gas exchange in a rapidly warming Arctic.</p>
</sec>

      
      </body>
    <back><app-group>

<app id="App1.Ch1.S1">
  <?xmltex \currentcnt{A}?><label>Appendix A</label><title>Simulation details</title>

<?xmltex \floatpos{h!}?><table-wrap id="App1.Ch1.S1.T3"><?xmltex \hack{\hsize\textwidth}?><?xmltex \currentcnt{A1}?><label>Table A1</label><caption><p id="d1e3790">Table with used variables, their description, and units.</p></caption><oasis:table frame="topbot"><oasis:tgroup cols="5">
     <oasis:colspec colnum="1" colname="col1" align="left"/>
     <oasis:colspec colnum="2" colname="col2" align="left"/>
     <oasis:colspec colnum="3" colname="col3" align="right"/>
     <oasis:colspec colnum="4" colname="col4" align="left"/>
     <oasis:colspec colnum="5" colname="col5" align="left"/>
     <oasis:thead>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">Variable</oasis:entry>
         <oasis:entry colname="col2">Name</oasis:entry>
         <oasis:entry colname="col3">Value</oasis:entry>
         <oasis:entry colname="col4">Unit</oasis:entry>
         <oasis:entry colname="col5">Eq. number</oasis:entry>
       </oasis:row>
     </oasis:thead>
     <oasis:tbody>
       <oasis:row>
         <oasis:entry colname="col1"><inline-formula><mml:math id="M171" display="inline"><mml:mi>a</mml:mi></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2">Empirical variable</oasis:entry>
         <oasis:entry colname="col3">109</oasis:entry>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M172" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kg</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">3</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col5">3</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><inline-formula><mml:math id="M173" display="inline"><mml:mi>b</mml:mi></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2">Empirical variable</oasis:entry>
         <oasis:entry colname="col3">6</oasis:entry>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M174" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kg</mml:mi><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">3</mml:mn></mml:mrow></mml:msup><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">K</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col5">3</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><inline-formula><mml:math id="M175" display="inline"><mml:mi>c</mml:mi></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2">Empirical variable</oasis:entry>
         <oasis:entry colname="col3">26</oasis:entry>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M176" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">g</mml:mi><mml:mspace width="0.125em" linebreak="nobreak"/><mml:mi mathvariant="normal">m</mml:mi><mml:msup><mml:mo>-</mml:mo><mml:mrow><mml:mn mathvariant="normal">7</mml:mn><mml:mo>/</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:msup><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">s</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>/</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col5">3</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><inline-formula><mml:math id="M177" display="inline"><mml:mrow><mml:msub><mml:mi>C</mml:mi><mml:mi>k</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2">Thermal heat capacity</oasis:entry>
         <oasis:entry colname="col3"/>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M178" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">J</mml:mi><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">3</mml:mn></mml:mrow></mml:msup><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">K</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col5">1, 8</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><inline-formula><mml:math id="M179" display="inline"><mml:mrow><mml:msub><mml:mi>D</mml:mi><mml:mi>k</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2">Thermal diffusivity</oasis:entry>
         <oasis:entry colname="col3"/>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M180" display="inline"><mml:mrow class="unit"><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msup><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">d</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col5">8, 9</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><inline-formula><mml:math id="M181" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">η</mml:mi><mml:mi>k</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2">Compactive viscosity factor</oasis:entry>
         <oasis:entry colname="col3"/>
         <oasis:entry colname="col4">10<inline-formula><mml:math id="M182" display="inline"><mml:msup><mml:mi/><mml:mn mathvariant="normal">6</mml:mn></mml:msup></mml:math></inline-formula> <inline-formula><mml:math id="M183" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">Pa</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:mi mathvariant="normal">s</mml:mi></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col5">5</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><inline-formula><mml:math id="M184" display="inline"><mml:mi>g</mml:mi></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2">Gravitation</oasis:entry>
         <oasis:entry colname="col3">9.81</oasis:entry>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M185" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">m</mml:mi><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">s</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col5">5</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><inline-formula><mml:math id="M186" display="inline"><mml:mrow><mml:msub><mml:mi>I</mml:mi><mml:mi>k</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2">Ice content</oasis:entry>
         <oasis:entry colname="col3"/>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M187" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kg</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msup></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col5">4, 6</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><inline-formula><mml:math id="M188" display="inline"><mml:mi>k</mml:mi></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2">Layer index</oasis:entry>
         <oasis:entry colname="col3"/>
         <oasis:entry colname="col4"/>
         <oasis:entry colname="col5">1, 2, 4, 5, 6, 7, 8</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><inline-formula><mml:math id="M189" display="inline"><mml:mrow><mml:msub><mml:mi>K</mml:mi><mml:mi>k</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2">Thermal heat conductivity</oasis:entry>
         <oasis:entry colname="col3"/>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M190" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">J</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">K</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">s</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col5">7</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><inline-formula><mml:math id="M191" display="inline"><mml:mrow><mml:msub><mml:mi>k</mml:mi><mml:mi mathvariant="normal">s</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2">Empirical variable</oasis:entry>
         <oasis:entry colname="col3">4000</oasis:entry>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M192" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">K</mml:mi></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col5">5</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><inline-formula><mml:math id="M193" display="inline"><mml:mrow><mml:msub><mml:mi>M</mml:mi><mml:mi>k</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2">Mass of overlaying snow layers</oasis:entry>
         <oasis:entry colname="col3"/>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M194" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kg</mml:mi></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col5">5</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><inline-formula><mml:math id="M195" display="inline"><mml:mi>P</mml:mi></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2">Precipitation</oasis:entry>
         <oasis:entry colname="col3"/>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M196" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">mm</mml:mi></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col5">2</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><inline-formula><mml:math id="M197" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi>k</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2">Layer density</oasis:entry>
         <oasis:entry colname="col3"/>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M198" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kg</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">3</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col5">1, 4, 5, 6, 7</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><inline-formula><mml:math id="M199" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">t</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2">Reference snow density</oasis:entry>
         <oasis:entry colname="col3">400</oasis:entry>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M200" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kg</mml:mi><mml:mspace linebreak="nobreak" width="0.125em"/><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">3</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col5">6</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><inline-formula><mml:math id="M201" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mtext>fresh</mml:mtext></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2">Fresh snow density</oasis:entry>
         <oasis:entry colname="col3"/>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M202" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kg</mml:mi><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">3</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col5">3</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><inline-formula><mml:math id="M203" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mn mathvariant="normal">0</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2">Reference snow density</oasis:entry>
         <oasis:entry colname="col3">50</oasis:entry>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M204" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kg</mml:mi><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">3</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col5">5, 7</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><inline-formula><mml:math id="M205" display="inline"><mml:mrow><mml:mi>r</mml:mi><mml:msub><mml:mi>w</mml:mi><mml:mo>min⁡</mml:mo></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2">Empirical parameter</oasis:entry>
         <oasis:entry colname="col3">0.03</oasis:entry>
         <oasis:entry colname="col4"/>
         <oasis:entry colname="col5">6</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><inline-formula><mml:math id="M206" display="inline"><mml:mrow><mml:mi>r</mml:mi><mml:msub><mml:mi>w</mml:mi><mml:mo>max⁡</mml:mo></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2">Empirical parameter</oasis:entry>
         <oasis:entry colname="col3">0.1</oasis:entry>
         <oasis:entry colname="col4"/>
         <oasis:entry colname="col5">6</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><inline-formula><mml:math id="M207" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mi>k</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2">Layer temperature</oasis:entry>
         <oasis:entry colname="col3"/>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M208" display="inline"><mml:mrow class="unit"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup><mml:mi mathvariant="normal">C</mml:mi></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col5">2, 5, 9</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><inline-formula><mml:math id="M209" display="inline"><mml:mrow><mml:msub><mml:mi>T</mml:mi><mml:mo>max⁡</mml:mo></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2">Threshold for snow–water phase changes</oasis:entry>
         <oasis:entry colname="col3">0</oasis:entry>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M210" display="inline"><mml:mrow class="unit"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup><mml:mi mathvariant="normal">C</mml:mi></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col5">2, 3, 5</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><inline-formula><mml:math id="M211" display="inline"><mml:mrow><mml:msub><mml:mi>U</mml:mi><mml:mn mathvariant="normal">10</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2">Reference wind temperature at 10 <inline-formula><mml:math id="M212" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">m</mml:mi></mml:mrow></mml:math></inline-formula> height</oasis:entry>
         <oasis:entry colname="col3"/>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M213" display="inline"><mml:mrow class="unit"><mml:msup><mml:mi mathvariant="normal">s</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col5">3</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><inline-formula><mml:math id="M214" display="inline"><mml:mrow><mml:msub><mml:mi>z</mml:mi><mml:mi>k</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2">Soil layer depth</oasis:entry>
         <oasis:entry colname="col3"/>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M215" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">m</mml:mi></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col5">4, 9</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><inline-formula><mml:math id="M216" display="inline"><mml:mrow><mml:msub><mml:mi>W</mml:mi><mml:mi>k</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2">Water content</oasis:entry>
         <oasis:entry colname="col3"/>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M217" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kg</mml:mi><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col5">6</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><inline-formula><mml:math id="M218" display="inline"><mml:mrow><mml:msub><mml:mi>W</mml:mi><mml:mrow><mml:mtext>cap</mml:mtext><mml:mo>,</mml:mo><mml:mo>max⁡</mml:mo></mml:mrow></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col2">Max. water holding capacity</oasis:entry>
         <oasis:entry colname="col3"/>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M219" display="inline"><mml:mrow class="unit"><mml:mi mathvariant="normal">kg</mml:mi><mml:mspace width="0.125em" linebreak="nobreak"/><mml:msup><mml:mi mathvariant="normal">m</mml:mi><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col5">6</oasis:entry>
       </oasis:row>
     </oasis:tbody>
   </oasis:tgroup></oasis:table></table-wrap>

<?xmltex \hack{\clearpage}?>
</app>
  </app-group><notes notes-type="codedataavailability"><title>Code and data availability</title>

      <p id="d1e4831">The code version used for this study is stored in a central code repository and will be made accessible upon request. Observational data used in this study can be retrieved from the following sources: Russian site observations (snow depth, air and soil <inline-formula><mml:math id="M220" display="inline"><mml:mi>T</mml:mi></mml:math></inline-formula>): RIHMI-WDC, <uri>http://meteo.ru/</uri>; PAGE21 site observations: <xref ref-type="bibr" rid="bib1.bibx22 bib1.bibx23 bib1.bibx40 bib1.bibx7 bib1.bibx8" id="text.71"/>; Zackenberg snow-related observations: <xref ref-type="bibr" rid="bib1.bibx18" id="text.72"/>; TTOP model permafrost estimate: <xref ref-type="bibr" rid="bib1.bibx39" id="text.73"/>.</p>
  </notes><app-group>
        <supplementary-material position="anchor"><p id="d1e4853">The supplement related to this article is available online at: <inline-supplementary-material xlink:href="https://doi.org/10.5194/bg-18-5767-2021-supplement" xlink:title="pdf">https://doi.org/10.5194/bg-18-5767-2021-supplement</inline-supplementary-material>.</p></supplementary-material>
        </app-group><notes notes-type="authorcontribution"><title>Author contributions</title>

      <p id="d1e4862">AP and FJWP designed the research. Model developments were lead by AP and implemented by AP, DW, and PAM. AP performed the model simulations and analysed the data. AP prepared the paper with contributions from all co-authors.</p>
  </notes><notes notes-type="competinginterests"><title>Competing interests</title>

      <p id="d1e4868">The contact author has declared that neither they nor their co-authors have any competing interests.</p>
  </notes><notes notes-type="disclaimer"><title>Disclaimer</title>

      <p id="d1e4874">Publisher's note: Copernicus Publications remains neutral with regard to jurisdictional claims in published maps and institutional affiliations.</p>
  </notes><ack><title>Acknowledgements</title><p id="d1e4880">We gratefully acknowledge support from the Swedish Research Council (WinterGap, registration no. 2017-05268) and the Research Council of Norway (WINTERPROOF, project no. 274711). David Wårlind has received funding from the European Union's Horizon 2020 research and innovation programme under grant agreement no. 641816 (CRESCENDO). David Wårlind and Paul A. Miller also acknowledge financial support from the Strategic Research Area MERGE and the Swedish national strategic e-Science research programme eSSENCE.</p></ack><notes notes-type="financialsupport"><title>Financial support</title>

      <p id="d1e4886">This research has been supported by the Swedish Research Council (WinterGap, registration no. 2017-05268), the Research Council of Norway (WINTERPROOF, project no. 274711), the European Union's Horizon 2020 research and innovation programme under grant agreement no. 641816 (CRESCENDO), the Strategic Research Area MERGE, and the Swedish national strategic e-Science research programme eSSENCE.</p>
  </notes><notes notes-type="reviewstatement"><title>Review statement</title>

      <p id="d1e4892">This paper was edited by Alexey V. Eliseev and reviewed by two anonymous referees.</p>
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    <!--<article-title-html>Model simulations of arctic biogeochemistry and permafrost extent are highly sensitive to the implemented snow scheme in LPJ-GUESS</article-title-html>
<abstract-html><p>The Arctic is warming rapidly, especially in winter, which is causing large-scale reductions in snow cover. Snow is one of the main controls on soil
thermodynamics, and changes in its thickness and extent affect both permafrost thaw and soil biogeochemistry. Since soil respiration during the cold
season potentially offsets carbon uptake during the growing season, it is essential to achieve a realistic simulation of the effect of snow cover on
soil conditions to more accurately project the direction of arctic carbon–climate feedbacks under continued winter warming.</p><p>The Lund–Potsdam–Jena General Ecosystem Simulator (LPJ-GUESS) dynamic vegetation model has used – up until now – a single layer snow scheme,
which underestimated the insulation effect of snow, leading to a cold bias in soil temperature. To address this shortcoming, we developed and
integrated a dynamic, multi-layer snow scheme in LPJ-GUESS. The new snow scheme performs well in simulating the insulation of snow at hundreds of
locations across Russia compared to observations. We show that improving this single physical factor enhanced simulations of permafrost extent
compared to an advanced permafrost product, where the overestimation of permafrost cover decreased from 10&thinsp;% to 5&thinsp;% using the new snow
scheme. Besides soil thermodynamics, the new snow scheme resulted in a doubled winter respiration and an overall higher vegetation carbon content.</p><p>This study highlights the importance of a correct representation of snow in ecosystem models to project biogeochemical processes that govern climate
feedbacks. The new dynamic snow scheme is an essential improvement in the simulation of cold season processes, which reduces the uncertainty of
model projections. These developments contribute to a more realistic simulation of arctic carbon–climate feedbacks.</p></abstract-html>
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