<?xml version="1.0" encoding="UTF-8"?>
<!DOCTYPE article PUBLIC "-//NLM//DTD Journal Publishing with OASIS Tables v3.0 20080202//EN" "https://jats.nlm.nih.gov/nlm-dtd/publishing/3.0/journalpub-oasis3.dtd">
<article xmlns:xlink="http://www.w3.org/1999/xlink" xmlns:mml="http://www.w3.org/1998/Math/MathML" xmlns:oasis="http://docs.oasis-open.org/ns/oasis-exchange/table" xml:lang="en" dtd-version="3.0" article-type="research-article">
  <front>
    <journal-meta><journal-id journal-id-type="publisher">BG</journal-id><journal-title-group>
    <journal-title>Biogeosciences</journal-title>
    <abbrev-journal-title abbrev-type="publisher">BG</abbrev-journal-title><abbrev-journal-title abbrev-type="nlm-ta">Biogeosciences</abbrev-journal-title>
  </journal-title-group><issn pub-type="epub">1726-4189</issn><publisher>
    <publisher-name>Copernicus Publications</publisher-name>
    <publisher-loc>Göttingen, Germany</publisher-loc>
  </publisher></journal-meta>
    <article-meta>
      <article-id pub-id-type="doi">10.5194/bg-23-6687-2026</article-id><title-group><article-title>Using deep learning to assimilate sun-induced fluorescence satellite observations in the ISBA land surface model</article-title><alt-title>Using deep learning to assimilate sun-induced fluorescence satellite observations</alt-title>
      </title-group>
      <contrib-group>
        <contrib contrib-type="author" corresp="no" rid="aff1">
          <name><surname>Vanderbecken</surname><given-names>Pierre</given-names></name>
          
        <ext-link>https://orcid.org/0000-0001-9722-5182</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff1">
          <name><surname>Vural</surname><given-names>Jasmin</given-names></name>
          
        <ext-link>https://orcid.org/0000-0001-8025-0349</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff1">
          <name><surname>Rojas-Muñoz</surname><given-names>Oscar</given-names></name>
          
        <ext-link>https://orcid.org/0000-0003-4444-1783</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff2">
          <name><surname>Garrigues</surname><given-names>Sébastien</given-names></name>
          
        </contrib>
        <contrib contrib-type="author" corresp="no" rid="aff1">
          <name><surname>Bonan</surname><given-names>Bertrand</given-names></name>
          
        <ext-link>https://orcid.org/0000-0002-8808-2201</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff4">
          <name><surname>Bacour</surname><given-names>Cédric</given-names></name>
          
        <ext-link>https://orcid.org/0000-0002-1913-3722</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff3">
          <name><surname>Rascher</surname><given-names>Uwe</given-names></name>
          
        <ext-link>https://orcid.org/0000-0002-9993-4588</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff3">
          <name><surname>Siegmann</surname><given-names>Bastian</given-names></name>
          
        </contrib>
        <contrib contrib-type="author" corresp="no" rid="aff2">
          <name><surname>de Rosnay</surname><given-names>Patricia</given-names></name>
          
        <ext-link>https://orcid.org/0000-0002-7374-3820</ext-link></contrib>
        <contrib contrib-type="author" corresp="yes" rid="aff1">
          <name><surname>Calvet</surname><given-names>Jean-Christophe</given-names></name>
          <email>jean-christophe.calvet@meteo.fr</email>
        <ext-link>https://orcid.org/0000-0001-6425-6492</ext-link></contrib>
        <aff id="aff1"><label>1</label><institution>Météo-France, CNRS, Univ. Toulouse, CNRM, Toulouse, France</institution>
        </aff>
        <aff id="aff2"><label>2</label><institution>ECMWF, Reading, UK and Bonn, Germany</institution>
        </aff>
        <aff id="aff3"><label>3</label><institution>Institute of Bio- and Geosciences, IBG-2: Plant Sciences, Forschungszentrum Jülich GmbH,  Leo-Brandt-Str., 52425 Jülich, Germany</institution>
        </aff>
        <aff id="aff4"><label>4</label><institution>Laboratoire des Sciences du Climat et de l'Environnement, LSCE/IPSL, CEA-CNRS-UVSQ,  Université Paris-Saclay, Gif-sur-Yvette, France</institution>
        </aff>
      </contrib-group>
      <author-notes><corresp id="corr1">Jean-Christophe Calvet (jean-christophe.calvet@meteo.fr)</corresp></author-notes><pub-date><day>23</day><month>September</month><year>2026</year></pub-date>
      
      <volume>23</volume>
      <issue>18</issue>
      <fpage>6687</fpage><lpage>6703</lpage>
      <history>
        <date date-type="received"><day>18</day><month>February</month><year>2026</year></date>
           <date date-type="rev-request"><day>24</day><month>February</month><year>2026</year></date>
           <date date-type="rev-recd"><day>2</day><month>September</month><year>2026</year></date>
           <date date-type="accepted"><day>9</day><month>September</month><year>2026</year></date>
      </history>
      <permissions>
        <copyright-statement>Copyright: © 2026 Pierre Vanderbecken et al.</copyright-statement>
        <copyright-year>2026</copyright-year>
      <license license-type="open-access"><license-p>This work is licensed under the Creative Commons Attribution 4.0 International License. To view a copy of this licence, visit <ext-link ext-link-type="uri" xlink:href="https://creativecommons.org/licenses/by/4.0/">https://creativecommons.org/licenses/by/4.0/</ext-link></license-p></license></permissions><self-uri xlink:href="https://bg.copernicus.org/articles/23/6687/2026/bg-23-6687-2026.html">This article is available from https://bg.copernicus.org/articles/23/6687/2026/bg-23-6687-2026.html</self-uri><self-uri xlink:href="https://bg.copernicus.org/articles/23/6687/2026/bg-23-6687-2026.pdf">The full text article is available as a PDF file from https://bg.copernicus.org/articles/23/6687/2026/bg-23-6687-2026.pdf</self-uri>
      <abstract><title>Abstract</title>

      <p id="d2e190">Accurate representations of the surface and vegetation are critical for simulating the terrestrial CO<sub>2</sub> cycle in response to climate and meteorological conditions. To meet this challenge, an increasing number of satellite missions are being launched which can monitor vegetation conditions and biomass. One is the Copernicus Sentinel-5P mission, which carries the TROPOMI instrument and retrieves Sun-Induced Chlorophyll Fluorescence (SIF). As an indicator of plant photosynthetic activity, SIF can provide critical information for evaluating and parameterising gross carbon flux dynamics in surface models. This study aims to assimilate TROPOMI SIF data into the ISBA (Interactions between Soil, Biosphere and Atmosphere) land surface model developed by Meteo-France, with the objective of directly correcting the representation of Leaf Area Index (LAI) and Gross Primary Production (GPP). To achieve this, we have developed a dedicated observation operator that links the modelled LAI to the TROPOMI SIF daily product. This neural network (NN) operator was developed using deep learning and was trained using observations over Europe. This operator achieved good accuracy, was implemented in a land data assimilation system (LDAS), and was used to assimilate TROPOMI SIF in ISBA using a sequential simplified extended Kalman filter. Specific experiments were conducted to study the assimilation process over the Ebro basin in Spain. This area is known for its irrigated croplands, which are not well represented by ISBA. Some experiments assimilate TROPOMI SIF, some assimilate a Copernicus Land Monitoring Service (CLMS) LAI 10 d product, and some assimilate both. This provided us with a useful point of reference for improving the vegetation simulation. Following SIF assimilation, LAI representation improved across the domain, highlighting heavily irrigated croplands. The gross primary production (GPP) derived from the analysis is closing the gap between the simulated and observed values, though a marked difference remains. When compared with other assimilation experiments, assimilating SIF alone provides a similar benefit to standard assimilation of a CLMS LAI product on LAI and GPP. The best improvements to the LAI and GPP results come from co-assimilating TROPOMI SIF with the CLMS LAI product, which combines the advantages of high-frequency SIF observations and robust 10 d LAI assimilation.</p>
  </abstract>
    
<funding-group>
<award-group id="gs1">
<funding-source>HORIZON EUROPE Digital, Industry and Space</funding-source>
<award-id>101082194</award-id>
<award-id>101082139</award-id>
</award-group>
</funding-group>
</article-meta>
  </front>
<body>
      

<sec id="Ch1.S1" sec-type="intro">
  <label>1</label><title>Introduction</title>
      <p id="d2e211">Monitoring the carbon cycle in near real time at city, regional and national scales helps decision-makers track the effectiveness of environmental policies in the context of climate change mitigation and emission reduction efforts <xref ref-type="bibr" rid="bib1.bibx29" id="paren.1"/>. One of the main efforts at the European level was to develop and maintain a new operational monitoring and verification support capacity (CO<sub>2</sub>MVS) for the carbon cycle. It has been demonstrated that the natural carbon flux of the land surface is the most uncertain component of the carbon budget <xref ref-type="bibr" rid="bib1.bibx1 bib1.bibx23" id="paren.2"/>. Near real-time monitoring requires a joint analysis of the water and carbon cycles. Incorporating new observations of soil moisture and vegetation variables, such as Leaf Area Index (LAI), into land surface models can enhance the precision of these systems <xref ref-type="bibr" rid="bib1.bibx51 bib1.bibx4" id="paren.3"/>. One promising type of observation is the remote sensing of sun-induced chlorophyll fluorescence (SIF) in vegetation  <xref ref-type="bibr" rid="bib1.bibx9 bib1.bibx43" id="paren.4"/>. Solar radiation flux is absorbed by chlorophyll, enabling photosynthesis and fuelling biological fluorescence in the near-infrared and far-red regions of the spectrum <xref ref-type="bibr" rid="bib1.bibx8" id="paren.5"/>. SIF has distinctive spectral maxima that can be extracted from Top of Atmosphere (TOA) radiance measurements taken in space <xref ref-type="bibr" rid="bib1.bibx35 bib1.bibx33" id="paren.6"/>. Recently, SIF was retrieved on the large swath of the Copernicus Sentinel-5 Precursor instrument: TROPOMI <xref ref-type="bibr" rid="bib1.bibx36 bib1.bibx27" id="paren.7"/>. It provides a daily estimate of SIF in the near-infrared spectrum. SIF observations have been shown to correlate strongly with LAI, gross primary production (GPP) and other vegetation monitoring variables <xref ref-type="bibr" rid="bib1.bibx37 bib1.bibx11" id="paren.8"/>. It has also been used to optimise the parameterisation of vegetation simulation models by <xref ref-type="bibr" rid="bib1.bibx5" id="text.9"/>.</p>
      <p id="d2e251">In order to improve operational monitoring of vegetation, <xref ref-type="bibr" rid="bib1.bibx24" id="text.10"/> assimilated the reprocessed 8 d SIF retrieval from raw TROPOMI data into the IFS land surface model developed at ECMWF on a global scale. In this study, we propose to constrain simulations from the ISBA (Interactions between Soil, Biomass and Atmosphere) land surface model, which is developed at Météo-France and available via the SURFEX (SURFace EXTernalisée) platform <xref ref-type="bibr" rid="bib1.bibx42 bib1.bibx19" id="paren.11"/>, by assimilating the raw daily TROPOMI SIF retrievals instead of the 8 d TROPOMI SIF product used by <xref ref-type="bibr" rid="bib1.bibx24" id="text.12"/>. Unlike <xref ref-type="bibr" rid="bib1.bibx24" id="text.13"/>, the ISBA land surface model simulates LAI. As the SIF is not explicitly simulated by ISBA, a new observation operator is required to map the model variables into observation space. A process-based SIF observation operator is complex, requiring computational resources to accurately solve the radiative processes and calibration to reduce associated uncertainties. This makes it difficult to implement such an observation operator in an operational system. An alternative machine learning-based operator for SIF could be more suitable. TROPOMI SIF provides daily global retrievals, which will serve as our database for developing and training a neural network observation operator. <xref ref-type="bibr" rid="bib1.bibx16" id="text.14"/> have successfully achieved the assimilation of new observations in the ISBA land surface model using a neural network observation operator for ASCAT (ESA Advanced Scatterometer) backscattering observations. We adapted this methodology and applied it to the assimilation of TROPOMI SIF.</p>
      <p id="d2e269">This article describes the design of the regional deep learning observation operator for SIF measurements and its use in a sequential assimilation process to constrain simulations produced by the ISBA land and vegetation surface model within the SURFEX platform. The benefits of assimilating TROPOMI SIF data will be evaluated in the Ebro basin. This specific region was chosen because the land surface model does not represent the heavily irrigated croplands resulting from field management. TROPOMI SIF assimilation can correct this. The next section presents the datasets used to train the observation operator and evaluate it later on. The land data assimilation and the deep learning operator are then described in the Methods section. Section 4 is dedicated to the results, beginning with an evaluation of the observation operator and the various assimilation strategies for TROPOMI SIF alongside different observation errors. Section 5 discusses the complementarity of LAI and TROPOMI SIF observations in a co-assimilation framework, and outlines limitations in GPP representation when assimilating only TROPOMI SIF. The final section draws conclusions.</p>
</sec>
<sec id="Ch1.S2">
  <label>2</label><title>Datasets</title>
<sec id="Ch1.S2.SS1">
  <label>2.1</label><title>Domain</title>
      <p id="d2e287">All the datasets for the training of the observation operator are extracted on a large regional domain. The database domain covers Europe, extending from 26° W to 46° E and from 28 to 72° N. The grid is uniform and regular with a resolution of 0.1°. This domain is the same domain as that used by <xref ref-type="bibr" rid="bib1.bibx28" id="text.15"/>. Grid cells of vegetation are selected within this domain using the ECOCLIMAP-II land cover classification <xref ref-type="bibr" rid="bib1.bibx21" id="paren.16"/>, which is available in SURFEX. Figure <xref ref-type="fig" rid="F1"/>a illustrates the most common vegetation type found within each grid cell. In ISBA, urban areas are considered as bare rock. As such, urban areas are also displayed as bare rock in Fig. <xref ref-type="fig" rid="F1"/>. In terms of low vegetation, Europe is dominated by C3-grassland and C3-crops, with forests of mixed deciduous and coniferous trees in higher latitudes. The zoom in Fig. <xref ref-type="fig" rid="F1"/>b is on the Ebro and Rio Segre basins in Spain. This region is well known for its heavily irrigated crop fields, which are usually difficult to model accurately.</p>

      <fig id="F1" specific-use="star"><label>Figure 1</label><caption><p id="d2e304">Dominant Plant Functional Type according to land use maps. <bold>(a)</bold> Domain Europe with ECOCLIMAP-II [26° W to 46° E, 28 to 72° N], the delimited area in red represents the Ebro basin subdomain. <bold>(b)</bold> The Ebro basin subdomain [1° W to 3° E, 39.5 to 43.5° N] with ECOCLIMAP II. <bold>(c)</bold> The Ebro basin subdomain with ECOCLIMAP-SG. Four surface types are shown: grass (C3 grassland, C4 grassland), crops (C3 croplands, C4 croplands, flooded crops), forest (deciduous broadleaf forest, coniferous needleleaf forest, evergreen broadleaf forest), No-veg (bare soil, rocks, snow and ice, peat and wetlands).</p></caption>
          <graphic xlink:href="https://bg.copernicus.org/articles/23/6687/2026/bg-23-6687-2026-f01.png"/>

        </fig>

</sec>
<sec id="Ch1.S2.SS2">
  <label>2.2</label><title>SIF database</title>
      <p id="d2e330">TROPOMI SIF is a high-level product obtained from the atmospheric reflectivity data collected by the Sentinel-5P SWIR instrument as part of the TROPOSIF project <xref ref-type="bibr" rid="bib1.bibx27" id="paren.17"/>. The distributed data ranges from May 2018 to December 2021 for the project, but the same process is now operational on the online data portal that distributes the product. Here, we used the data from the TROPOSIF project. Sentinel-5P operates on a heliosynchronous orbit and, with a swath of 2600 km, passes over a given region at least once a day <xref ref-type="bibr" rid="bib1.bibx55" id="paren.18"/>. The nadir measurement crosses the equator at 13:30 local solar time, achieving a resolution of 7 km <inline-formula><mml:math id="M3" display="inline"><mml:mrow><mml:mo>×</mml:mo><mml:mn mathvariant="normal">3.5</mml:mn></mml:mrow></mml:math></inline-formula> km. The retrieval process involves statistically modelling TOA radiances over vegetation while excluding the signal induced by the underlying bare surface. The MODIS2018 database <xref ref-type="bibr" rid="bib1.bibx22" id="paren.19"/> is used to identify areas of vegetation that are relevant for SIF retrieval. Using prior knowledge of the SIF spectrum, the algorithm extracts the SIF-related part of the TOA radiance signal <xref ref-type="bibr" rid="bib1.bibx26" id="paren.20"/>. This process ensures that the distribution of TROPOMI SIF observations is centred on zero over grid cells where there is only bare soil (i.e. where MODIS classifies the land as having no vegetation). This means that non-physical negative values for TROPOMI SIF can occur. TROPOMI SIF instantaneous estimates are computed according to a spectral window between 743 and 753 nm. The estimates are robust to cloud contamination. Alongside these instantaneous estimates, the Level-2 TROPOMI SIF product provides a corrected value according to day length and solar angle at a given latitude <xref ref-type="bibr" rid="bib1.bibx36" id="paren.21"/>. We will be using the raw SIF retrieval from this corrected Level-2 TROPOMI SIF.</p>
      <p id="d2e359">In this study, since our model domain is defined on a 0.1° resolution regular grid, the raw SIF retrievals were aggregated onto our model grid. Only all positive instantaneous measurements for a given day covering a grid cell in the domain are aggregated to provide a value for that grid cell.</p>
</sec>
<sec id="Ch1.S2.SS3">
  <label>2.3</label><title>LAI and GPP observations</title>
      <p id="d2e370">To complete the training and evaluation of vegetation monitoring, we considered two additional types of vegetation observation: LAI and GPP. The LAI satellite products used here are provided by the Copernicus Land Monitoring Service (CLMS). LAI-GEOV1 <xref ref-type="bibr" rid="bib1.bibx7" id="paren.22"/> is an observation-driven product with a resolution of 1 km, based on measurements from the SPOT-VGT satellite (from 1999 to 2014) and the PROBA-V satellite (from 2014 to June 2020). It provides an estimation of LAI at a global scale every 10 d denoted thereafter as LAI-V1. To align with our daily TROPOMI SIF product, the LAI maps undergo linear time interpolation. Due to instrument issues or excessive cloud cover during observations, random local missing values are displayed on each global map. Time interpolation is only performed between two consecutive LAI observations of the same grid cell separated by 10 d. If an LAI observation is missing, the gap is not filled. As in previous studies <xref ref-type="bibr" rid="bib1.bibx2 bib1.bibx6 bib1.bibx16" id="paren.23"><named-content content-type="post">among others</named-content></xref>, LAI observations are aggregated to the spatial resolution of the model grid points using an arithmetic average, provided that the LAI observations cover at least 50 % of the grid cell area. As the LAI-V1 product was discontinued in June 2020, the CLMS LAI raster at a resolution of 300 m will be used instead. This is derived from Sentinel-3 OLCI (Ocean and Land Colour Instrument) measurements. This provides data every 10 d and is spatially and temporally interpolated in the same way as the LAI-V1 product and, is referred to as LAI-300 thereafter.</p>
      <p id="d2e381">For the GPP, we used the most recent global FLUXCOM-X in V0 product <xref ref-type="bibr" rid="bib1.bibx34 bib1.bibx44" id="paren.24"/>. This provides daily estimates of GPP at a quarter-degree resolution across the globe from 2018 to 2021. The time series of the three observation products are displayed in Fig. <xref ref-type="fig" rid="F2"/>. All of the displayed observations are from the European domain; the GPP from FLUXCOM remains at its original resolution. The expected seasonal cycle is present in all products. The high-frequency variability in median LAI and TROPOMI SIF values is due to changes in data masking by clouds and filtering applied to both datasets. The isolated spike in mid-July 2019, which is present only in the TROPOMI SIF and LAI-V1 datasets, is a rare case in which the Iceland area is the only region observed due to common masking.</p>

      <fig id="F2" specific-use="star"><label>Figure 2</label><caption><p id="d2e391">Time series of the median value of LAI (green), SIF (black) and GPP (blue) observed over the domain shown in Fig. <xref ref-type="fig" rid="F1"/>a.</p></caption>
          <graphic xlink:href="https://bg.copernicus.org/articles/23/6687/2026/bg-23-6687-2026-f02.png"/>

        </fig>

</sec>
</sec>
<sec id="Ch1.S3">
  <label>3</label><title>Methods</title>
<sec id="Ch1.S3.SS1">
  <label>3.1</label><title>ISBA land surface model</title>
      <p id="d2e418">Throughout this study, we use the version 8.1 of the SURFEX platform <xref ref-type="bibr" rid="bib1.bibx42" id="paren.25"/> to model the surface. To model the soil-plant system, we use the ISBA land surface model. Soil moisture and LAI are computed independently for different plant functional types (PFTs). Twelve PFTs are used and distributed according to land use maps. The ISBA model <xref ref-type="bibr" rid="bib1.bibx46 bib1.bibx45" id="paren.26"/> is used in the A-gs configuration <xref ref-type="bibr" rid="bib1.bibx14" id="paren.27"/>, where A stands for CO<sub>2</sub> assimilation by the leaves and gs stands for the stomatal conductance simulated by the model. ISBA-A-gs considers the functional relationship between stomatal aperture and photosynthesis under well-watered conditions, as described in the biochemical A-gs model proposed by <xref ref-type="bibr" rid="bib1.bibx30" id="text.28"/> and <xref ref-type="bibr" rid="bib1.bibx31" id="text.29"/>. For drier conditions, the model also includes a representation of soil moisture stress, as well as two dedicated drought responses: one for herbaceous vegetation and crops, with specifications for C3 and C4 types <xref ref-type="bibr" rid="bib1.bibx12" id="paren.30"/>, and one for forest vegetation <xref ref-type="bibr" rid="bib1.bibx15" id="paren.31"/>. Further details on the ISBA-A-gs can be found in <xref ref-type="bibr" rid="bib1.bibx25" id="text.32"/>. Below the surface, the soil is discretised vertically for both soil temperature and soil moisture. The multi-layer soil model uses a diffusion scheme to solve the one-dimensional Fourier law <xref ref-type="bibr" rid="bib1.bibx18" id="paren.33"/> for soil temperature, and a mixed form of the Richards equation <xref ref-type="bibr" rid="bib1.bibx49" id="paren.34"/> for soil moisture. The ISBA models' simulations can be inaccurate when compared with observations. We address these inaccuracies by assimilating observations into the model.</p>
</sec>
<sec id="Ch1.S3.SS2">
  <label>3.2</label><title>Land Data Assimilation system</title>
      <p id="d2e469">To constrain the ISBA-A-gs model simulations, we assimilate observations sequentially using the global Land Data Assimilation System (LDAS-Monde) described in <xref ref-type="bibr" rid="bib1.bibx3" id="text.35"/>. This process involves updating the control variables in the ISBA model by minimising discrepancies between the variables and observations, and between the variables and a prior estimate. In our case, the control vector comprises the LAI and the soil moisture in the different layers. The results from this process is known as the “analysis”. We use a simplified extended Kalman filter <xref ref-type="bibr" rid="bib1.bibx40" id="paren.36"><named-content content-type="pre">SEKF,</named-content></xref>. Thereafter, the control vector will be referred to as <inline-formula><mml:math id="M5" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="bold-italic">x</mml:mi><mml:mi>i</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>, where the subscript denotes the temporal step, which corresponds to one day since the assimilation is performed daily. The analysis update equation at <inline-formula><mml:math id="M6" display="inline"><mml:mi>t</mml:mi></mml:math></inline-formula> of the Kalman filter from the background control vector at <inline-formula><mml:math id="M7" display="inline"><mml:mrow><mml:mi>t</mml:mi><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:math></inline-formula> is as follows:
          

                <disp-formula id="Ch1.E1" specific-use="gather" content-type="subnumberedsingle"><mml:math id="M8" display="block"><mml:mtable displaystyle="true"><mml:mlabeledtr id="Ch1.E1.2"><mml:mtd><mml:mtext>1a</mml:mtext></mml:mtd><mml:mtd><mml:mrow><mml:mstyle displaystyle="true" class="stylechange"/><mml:msubsup><mml:mi mathvariant="bold-italic">x</mml:mi><mml:mi>t</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msubsup><mml:mo>=</mml:mo><mml:msub><mml:mi mathvariant="script">M</mml:mi><mml:mrow><mml:mi>t</mml:mi><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msub><mml:mo>(</mml:mo><mml:msubsup><mml:mi mathvariant="bold-italic">x</mml:mi><mml:mrow><mml:mi>t</mml:mi><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow><mml:mi mathvariant="normal">a</mml:mi></mml:msubsup><mml:mo>)</mml:mo></mml:mrow></mml:mtd></mml:mlabeledtr><mml:mlabeledtr id="Ch1.E1.3"><mml:mtd><mml:mtext>1b</mml:mtext></mml:mtd><mml:mtd><mml:mrow><mml:mstyle displaystyle="true" class="stylechange"/><mml:msubsup><mml:mi mathvariant="bold-italic">x</mml:mi><mml:mi>t</mml:mi><mml:mi mathvariant="normal">a</mml:mi></mml:msubsup><mml:mo>=</mml:mo><mml:msubsup><mml:mi mathvariant="bold-italic">x</mml:mi><mml:mi>t</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msubsup><mml:mo>+</mml:mo><mml:munder><mml:munder class="underbrace"><mml:mrow><mml:msub><mml:mi mathvariant="bold">K</mml:mi><mml:mi>t</mml:mi></mml:msub><mml:mo>(</mml:mo><mml:msub><mml:mi mathvariant="bold-italic">y</mml:mi><mml:mi>t</mml:mi></mml:msub><mml:mo>-</mml:mo><mml:msub><mml:mi mathvariant="script">H</mml:mi><mml:mi>t</mml:mi></mml:msub><mml:mo>(</mml:mo><mml:msubsup><mml:mi mathvariant="bold-italic">x</mml:mi><mml:mi>t</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msubsup><mml:mo>)</mml:mo><mml:mo>)</mml:mo></mml:mrow><mml:mo mathvariant="normal">︸</mml:mo></mml:munder><mml:mi mathvariant="normal">Increment</mml:mi></mml:munder><mml:mo>,</mml:mo></mml:mrow></mml:mtd></mml:mlabeledtr></mml:mtable></mml:math></disp-formula>

          where the superscripts “<inline-formula><mml:math id="M9" display="inline"><mml:mi>a</mml:mi></mml:math></inline-formula>” and “<inline-formula><mml:math id="M10" display="inline"><mml:mi>b</mml:mi></mml:math></inline-formula>” stand for analysis and background respectively. <inline-formula><mml:math id="M11" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="bold-italic">y</mml:mi><mml:mi>t</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> represents observations. The operators <inline-formula><mml:math id="M12" display="inline"><mml:mi mathvariant="script">M</mml:mi></mml:math></inline-formula> and <inline-formula><mml:math id="M13" display="inline"><mml:mi mathvariant="script">H</mml:mi></mml:math></inline-formula> are respectively the land surface model (i.e. ISBA) and the linear observation operator that maps the control vector into the observation space. The Kalman gain <inline-formula><mml:math id="M14" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="bold">K</mml:mi><mml:mi>t</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> is defined at time <inline-formula><mml:math id="M15" display="inline"><mml:mi>t</mml:mi></mml:math></inline-formula> as follows:

            <disp-formula id="Ch1.E4" content-type="numbered"><label>2</label><mml:math id="M16" display="block"><mml:mrow><mml:msub><mml:mi mathvariant="bold">K</mml:mi><mml:mi>t</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:msubsup><mml:mi mathvariant="bold">BJ</mml:mi><mml:mi>t</mml:mi><mml:mo>⊤</mml:mo></mml:msubsup><mml:mo>(</mml:mo><mml:mi mathvariant="bold">R</mml:mi><mml:mo>+</mml:mo><mml:msub><mml:mi mathvariant="bold">J</mml:mi><mml:mi>i</mml:mi></mml:msub><mml:msubsup><mml:mi mathvariant="bold">BJ</mml:mi><mml:mi>t</mml:mi><mml:mo>⊤</mml:mo></mml:msubsup><mml:msup><mml:mo>)</mml:mo><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:math></disp-formula>

          where <inline-formula><mml:math id="M17" display="inline"><mml:mi mathvariant="bold">B</mml:mi></mml:math></inline-formula> and <inline-formula><mml:math id="M18" display="inline"><mml:mi mathvariant="bold">R</mml:mi></mml:math></inline-formula> are error covariance matrices characterizing the background and observation errors respectively. The operator <inline-formula><mml:math id="M19" display="inline"><mml:mi mathvariant="bold">J</mml:mi></mml:math></inline-formula> is the Jacobian matrix of the partial derivatives and is defined according to Eq. (<xref ref-type="disp-formula" rid="Ch1.E5"/>).

            <disp-formula id="Ch1.E5" content-type="numbered"><label>3</label><mml:math id="M20" display="block"><mml:mrow><mml:msub><mml:mi mathvariant="bold">J</mml:mi><mml:mi>t</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:mstyle displaystyle="true"><mml:mfrac style="display"><mml:mrow><mml:mo>∂</mml:mo><mml:mi mathvariant="script">H</mml:mi><mml:mo>(</mml:mo><mml:msubsup><mml:mi mathvariant="bold-italic">x</mml:mi><mml:mi>t</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msubsup><mml:mo>)</mml:mo></mml:mrow><mml:mrow><mml:mo>∂</mml:mo><mml:mo>(</mml:mo><mml:msubsup><mml:mi mathvariant="bold-italic">x</mml:mi><mml:mrow><mml:mi>t</mml:mi><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow><mml:mi mathvariant="normal">a</mml:mi></mml:msubsup><mml:mo>)</mml:mo></mml:mrow></mml:mfrac></mml:mstyle><mml:mo>=</mml:mo><mml:mstyle displaystyle="true"><mml:mfrac style="display"><mml:mrow><mml:mo>∂</mml:mo><mml:mi mathvariant="script">H</mml:mi><mml:mo>(</mml:mo><mml:msub><mml:mi mathvariant="script">M</mml:mi><mml:mrow><mml:mi>t</mml:mi><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:msub><mml:mo>(</mml:mo><mml:msubsup><mml:mi mathvariant="bold-italic">x</mml:mi><mml:mrow><mml:mi>t</mml:mi><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow><mml:mi mathvariant="normal">a</mml:mi></mml:msubsup><mml:mo>)</mml:mo><mml:mo>)</mml:mo></mml:mrow><mml:mrow><mml:mo>∂</mml:mo><mml:mo>(</mml:mo><mml:msubsup><mml:mi mathvariant="bold-italic">x</mml:mi><mml:mrow><mml:mi>t</mml:mi><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow><mml:mi mathvariant="normal">a</mml:mi></mml:msubsup><mml:mo>)</mml:mo></mml:mrow></mml:mfrac></mml:mstyle></mml:mrow></mml:math></disp-formula>

          This Jacobian is computed using finite differences. This is achieved by perturbing each component of the control vector defined by <xref ref-type="bibr" rid="bib1.bibx50" id="text.37"/>. To simplify the extended Kalman filter, its background error covariance and the observation error covariance matrices are assumed to be diagonal. These covariance values are fixed for a given assimilation window of 24 h, as detailed in <xref ref-type="bibr" rid="bib1.bibx41 bib1.bibx2 bib1.bibx6 bib1.bibx17 bib1.bibx3 bib1.bibx20" id="text.38"><named-content content-type="post">among others</named-content></xref>.</p>
      <p id="d2e871">The background state is computed from the analysis performed over the previous 24 h assimilation window, see <xref ref-type="bibr" rid="bib1.bibx10" id="text.39"/> for more details. Assimilation are thus cycled over a 24 h period like described in the scheme displayed by Fig. <xref ref-type="fig" rid="F3"/>. As example, the baseline LDAS configuration at Meteo-France involves assimilating LAI 10 d retrievals with a fixed relative observation error of 20 % <xref ref-type="bibr" rid="bib1.bibx28 bib1.bibx10 bib1.bibx4" id="paren.40"><named-content content-type="post">among others</named-content></xref>. Since the purpose of assimilation is to bring the model closer to the observations, the difference between the analysis and the observations in observation space – denoted as residuals – should be smaller than the difference between the prior simulation and the observations – denoted as innovation. LAI assimilation does not require a new observation operator to be defined, but SIF assimilation does. In order to assimilate SIF, we have designed a new observation operator based on a neural network that maps ISBA control variables to TROPOMI SIF retrievals.</p>

      <fig id="F3" specific-use="star"><label>Figure 3</label><caption><p id="d2e886">Schematic representation of the neural network architecture integrated in the assimilation framework. <inline-formula><mml:math id="M21" display="inline"><mml:mrow><mml:msup><mml:mi mathvariant="bold-italic">x</mml:mi><mml:mi mathvariant="normal">b</mml:mi></mml:msup></mml:mrow></mml:math></inline-formula> and <inline-formula><mml:math id="M22" display="inline"><mml:mrow><mml:msup><mml:mi mathvariant="bold-italic">x</mml:mi><mml:mi mathvariant="normal">a</mml:mi></mml:msup></mml:mrow></mml:math></inline-formula> are respectively the background and the analysis and represents 8 control variables. The observation assimilated <inline-formula><mml:math id="M23" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="bold-italic">y</mml:mi><mml:mi>i</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> at the <inline-formula><mml:math id="M24" display="inline"><mml:mi>i</mml:mi></mml:math></inline-formula>th cycle in this schematic representation is TROPOMI SIF.</p></caption>
          <graphic xlink:href="https://bg.copernicus.org/articles/23/6687/2026/bg-23-6687-2026-f03.png"/>

        </fig>

</sec>
<sec id="Ch1.S3.SS3">
  <label>3.3</label><title>Neural network observation operator</title>
      <p id="d2e943">For training and evaluation purposes, all areas covered by ocean, water, snow or lakes, or lying above 1500 m a.s.l., or where bare soil, rock, snow or ice are most prevalent according to ECOCLIMAP-II, are removed. Grid cells with no LAI or SIF observations, and grid cells where the soil surface temperature fell below 277.15 K according to the SEEDS project analysis <xref ref-type="bibr" rid="bib1.bibx38" id="paren.41"/> and <xref ref-type="bibr" rid="bib1.bibx52" id="text.42"/>, are also excluded to prevent the inclusion of frozen vegetation and soil.</p>
      <p id="d2e952">Here, we address a regression problem that can be solved using a simple feed-forward neural network (NN) for nonlinear regression. This strategy produced good results for the assimilation of ASCAT backscattering coefficient <inline-formula><mml:math id="M25" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">σ</mml:mi><mml:mn mathvariant="normal">0</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> in  <xref ref-type="bibr" rid="bib1.bibx16" id="text.43"/>, where NNs were trained independently for each grid cell. However, in this study, we will train a single neural network that can be applied to any grid cell. A feed-forward neural network’s basic architecture comprises an input layer, one or more hidden layers of neurons, and an output layer, which provides an estimate. After some testing, we found that the configuration comprising two hidden layers of 128 ReLu-activated neurons strikes the best balance between computational cost and accuracy. As our problem is a regression problem, our output layer comprises a single neuron with linear activation as shown in Fig. <xref ref-type="fig" rid="F3"/>. We added a Gaussian noise layer to the first hidden layer, before the activation layer, to reproduce the effect of a noisy activation layer that would be applied to the inputs. This helps to avoid overfitting during training and enables the duplication of the inputs for data augmentation. This Gaussian layer is only involved during the training process. We expect the neural network to provide smoother estimates than the observations and to filter out the inherent noise of the TROPOMI SIF data. We added a batch normalisation layer at the end of each hidden layer to regularise the learning process.</p>
      <p id="d2e971">The distribution of TROPOMI SIF values resembles a log-normal distribution, with a mode close to zero (see Fig. S1 in the Supplement). In order to prevent the neural network from making non-physical negative predictions and to widen the SIF distribution, it will be trained to estimate the SIF<sub><italic>l</italic></sub> variable. This is the SIF transformed by the strictly increasing function given in Eq. (<xref ref-type="disp-formula" rid="Ch1.E6.7"/>). We applied the following strictly increasing function Eq. (<xref ref-type="disp-formula" rid="Ch1.E6.7"/>) to the TROPOMI SIF values to predict SIF<sub><italic>l</italic></sub>. To reconstruct <inline-formula><mml:math id="M28" display="inline"><mml:mi mathvariant="normal">SIF</mml:mi></mml:math></inline-formula> estimates from the neural network outputs, the reverse equations Eq. (<xref ref-type="disp-formula" rid="Ch1.E6.8"/>) are used.
          

                <disp-formula id="Ch1.E6" specific-use="gather" content-type="subnumberedsingle"><mml:math id="M29" display="block"><mml:mtable displaystyle="true"><mml:mlabeledtr id="Ch1.E6.7"><mml:mtd><mml:mtext>4a</mml:mtext></mml:mtd><mml:mtd><mml:mrow><mml:mstyle class="stylechange" displaystyle="true"/><mml:msub><mml:mi mathvariant="normal">SIF</mml:mi><mml:mi>l</mml:mi></mml:msub><mml:mo>=</mml:mo><mml:mo mathvariant="italic" mathsize="2.0em">{</mml:mo><mml:mtable class="array" columnalign="left left"><mml:mtr><mml:mtd><mml:mrow><mml:mi>ln⁡</mml:mi><mml:mo>(</mml:mo><mml:mi mathvariant="normal">SIF</mml:mi><mml:mo>)</mml:mo><mml:mo>,</mml:mo></mml:mrow></mml:mtd><mml:mtd><mml:mrow><mml:mtext>if</mml:mtext><mml:mspace linebreak="nobreak" width="0.25em"/><mml:mi mathvariant="normal">SIF</mml:mi><mml:mo>≤</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:mtd></mml:mtr><mml:mtr><mml:mtd><mml:mrow><mml:mi>exp⁡</mml:mi><mml:mo>(</mml:mo><mml:mi mathvariant="normal">SIF</mml:mi><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>)</mml:mo><mml:mo>-</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>,</mml:mo></mml:mrow></mml:mtd><mml:mtd><mml:mrow><mml:mtext>if</mml:mtext><mml:mspace linebreak="nobreak" width="0.25em"/><mml:mi mathvariant="normal">SIF</mml:mi><mml:mo>&gt;</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>.</mml:mo></mml:mrow></mml:mtd></mml:mtr></mml:mtable></mml:mrow></mml:mtd></mml:mlabeledtr><mml:mlabeledtr id="Ch1.E6.8"><mml:mtd><mml:mtext>4b</mml:mtext></mml:mtd><mml:mtd><mml:mrow><mml:mstyle displaystyle="true" class="stylechange"/><mml:mi mathvariant="normal">SIF</mml:mi><mml:mspace linebreak="nobreak" width="0.25em"/><mml:mo>=</mml:mo><mml:mo mathsize="2.0em" mathvariant="italic">{</mml:mo><mml:mtable class="array" columnalign="left left"><mml:mtr><mml:mtd><mml:mrow><mml:mi>exp⁡</mml:mi><mml:mo>(</mml:mo><mml:msub><mml:mi mathvariant="normal">SIF</mml:mi><mml:mi>l</mml:mi></mml:msub><mml:mo>)</mml:mo><mml:mo>,</mml:mo></mml:mrow></mml:mtd><mml:mtd><mml:mrow><mml:mtext>if</mml:mtext><mml:mspace width="0.25em" linebreak="nobreak"/><mml:msub><mml:mi mathvariant="normal">SIF</mml:mi><mml:mi>l</mml:mi></mml:msub><mml:mo>≤</mml:mo><mml:mn mathvariant="normal">0</mml:mn></mml:mrow></mml:mtd></mml:mtr><mml:mtr><mml:mtd><mml:mrow><mml:mi>ln⁡</mml:mi><mml:mo>(</mml:mo><mml:msub><mml:mi mathvariant="normal">SIF</mml:mi><mml:mi>l</mml:mi></mml:msub><mml:mo>+</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>)</mml:mo><mml:mo>+</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>,</mml:mo></mml:mrow></mml:mtd><mml:mtd><mml:mrow><mml:mtext>if</mml:mtext><mml:mspace linebreak="nobreak" width="0.25em"/><mml:msub><mml:mi mathvariant="normal">SIF</mml:mi><mml:mi>l</mml:mi></mml:msub><mml:mo>&gt;</mml:mo><mml:mn mathvariant="normal">0</mml:mn><mml:mo>.</mml:mo></mml:mrow></mml:mtd></mml:mtr></mml:mtable></mml:mrow></mml:mtd></mml:mlabeledtr></mml:mtable></mml:math></disp-formula></p>
      <p id="d2e1173">For the loss function, we compare the label value and the prediction of the neural network according to an Huber loss, which follows Eq. (<xref ref-type="disp-formula" rid="Ch1.E9"/>). This loss behave mainly like the usual mean square loss while being less sensitive to outliers due to its more linear behaviour for extreme differences.

            <disp-formula id="Ch1.E9" content-type="numbered"><label>5</label><mml:math id="M30" display="block"><mml:mrow><mml:mi mathvariant="script">L</mml:mi><mml:mo>(</mml:mo><mml:msub><mml:mi mathvariant="normal">SIF</mml:mi><mml:mi>l</mml:mi></mml:msub><mml:mo>,</mml:mo><mml:mi>x</mml:mi><mml:mo>)</mml:mo><mml:mo>=</mml:mo><mml:mo mathvariant="italic" mathsize="2.0em">{</mml:mo><mml:mtable class="array" columnalign="left left"><mml:mtr><mml:mtd><mml:mrow><mml:mstyle displaystyle="false"><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mn mathvariant="normal">1</mml:mn><mml:mn mathvariant="normal">2</mml:mn></mml:mfrac></mml:mstyle></mml:mstyle><mml:mo>(</mml:mo><mml:msub><mml:mi mathvariant="normal">SIF</mml:mi><mml:mi>l</mml:mi></mml:msub><mml:mo>-</mml:mo><mml:mi>H</mml:mi><mml:mo>(</mml:mo><mml:mi>x</mml:mi><mml:mo>)</mml:mo><mml:msup><mml:mo>)</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:msup><mml:mo>,</mml:mo></mml:mrow></mml:mtd><mml:mtd><mml:mrow><mml:mtext>if </mml:mtext><mml:mspace width="0.25em" linebreak="nobreak"/><mml:mo>|</mml:mo><mml:msub><mml:mi mathvariant="normal">SIF</mml:mi><mml:mi>l</mml:mi></mml:msub><mml:mo>-</mml:mo><mml:mi>H</mml:mi><mml:mo>(</mml:mo><mml:mi>x</mml:mi><mml:mo>)</mml:mo><mml:mo>|</mml:mo><mml:mo>≤</mml:mo><mml:mn mathvariant="normal">1</mml:mn></mml:mrow></mml:mtd></mml:mtr><mml:mtr><mml:mtd><mml:mrow><mml:mo>(</mml:mo><mml:mo>|</mml:mo><mml:msub><mml:mi mathvariant="normal">SIF</mml:mi><mml:mi>l</mml:mi></mml:msub><mml:mo>-</mml:mo><mml:mi>H</mml:mi><mml:mo>(</mml:mo><mml:mi>x</mml:mi><mml:mo>)</mml:mo><mml:mo>|</mml:mo><mml:mo>-</mml:mo><mml:mstyle displaystyle="false"><mml:mstyle displaystyle="false"><mml:mfrac style="text"><mml:mn mathvariant="normal">1</mml:mn><mml:mn mathvariant="normal">2</mml:mn></mml:mfrac></mml:mstyle></mml:mstyle><mml:mo>)</mml:mo><mml:mo>,</mml:mo></mml:mrow></mml:mtd><mml:mtd><mml:mrow><mml:mtext>if </mml:mtext><mml:mspace width="0.25em" linebreak="nobreak"/><mml:mo>|</mml:mo><mml:msub><mml:mi mathvariant="normal">SIF</mml:mi><mml:mi>l</mml:mi></mml:msub><mml:mo>-</mml:mo><mml:mi>H</mml:mi><mml:mo>(</mml:mo><mml:mi>x</mml:mi><mml:mo>)</mml:mo><mml:mo>|</mml:mo><mml:mo>&gt;</mml:mo><mml:mn mathvariant="normal">1</mml:mn><mml:mo>.</mml:mo></mml:mrow></mml:mtd></mml:mtr></mml:mtable></mml:mrow></mml:math></disp-formula></p>
      <p id="d2e1338">After a thorough study exposed in the Sect. S1 of the Supplement, the neural network takes only as  predictors the DOY, LAT, LON and the observed LAI-V1. This configuration ensures that SIF assimilation can easily adapt to any land surface model and remain robust in the face of improvements to these models. The NN was trained for at least 100 epochs to ensure convergence of the 18 000 estimated weights, as shown in Fig. S5 of the Supplement. The full data range is from 1 June 2018 to 31 December 2021; however, LAI-V1 is only available until June 2020, meaning that the training and testing datasets include only dates between the beginning of June 2018 and the end of May 2020. The first year (i.e. 1 June 2018 to 31 May 2019) was used for training. Within the training period, 40 % of the training database is effectively used for the training of the weights, while the remaining 60 % is used to validate the neural network internally during the learning process. This division is randomised to avoid spurious temporal correlations. The “training” and “validation” splits are used to check for possible overfitting during training. Even when using only 40 % of the training dataset, the neural network is trained using 14 million input/output pairs. As Fig. S5 in the Supplement shows, using more data for training did not increase accuracy. Data augmentation involves duplicating random training values until the size of the dataset is double the original size. The test dataset, which is unknown to the training model, is used to evaluate the model's ability to generalise. This ranges from 1 June 2019 to 31 May 2020. In summary, the neural network learns using data from one year and is then evaluated using data from a different year.</p>
      <p id="d2e1341">To evaluate thereafter the NN, we will compute the Pearson's correlation coefficient (<inline-formula><mml:math id="M31" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">p</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula>) between the reconstruct SIF from the NN and the observed one, the root mean square error (RMSE) between the reconstruct SIF and the observation, the mean bias  of the difference between the reconstruct SIF and the observed SIF (i.e. experiment minus observations), the unbiased root mean square error (ubRMSE) to have an estimation of the standard deviation of the differences and the extreme values of the differences. These criteria will be computed for both the training and testing sets to ensure that there is no overfitting.</p>
</sec>
<sec id="Ch1.S3.SS4">
  <label>3.4</label><title>Assimilation setup over Ebro basin</title>
      <p id="d2e1363">To limit the computational cost, the assimilation TROPOMI SIF will be done only on a small area of <inline-formula><mml:math id="M32" display="inline"><mml:mrow><mml:mn mathvariant="normal">40</mml:mn><mml:mo>×</mml:mo><mml:mn mathvariant="normal">40</mml:mn></mml:mrow></mml:math></inline-formula> cells within the European domain displayed in Fig. <xref ref-type="fig" rid="F1"/>b. This domain encompasses the Ebro and Rio Segre basins in Spain. This area comprises heavily irrigated croplands which are usually not accurately simulated by the land surface model <xref ref-type="bibr" rid="bib1.bibx32 bib1.bibx39" id="paren.44"/>. Our approach aims to enhance the simulation of these irrigated areas by assimilating TROPOMI SIF data. The assimilation period runs from 1 January 2018 to 31 December 2021. The SEKF from the LDAS model assimilates the observations daily at 13:00 UTC. While the NN operator was trained using European data screened according to ECOCLIMAP-II, the land cover map used for the assimilation experiments was defined using ECOCLIMAP-SG <xref ref-type="bibr" rid="bib1.bibx13" id="paren.45"/>. This land cover is more recent and accurate than the ECOCLIMAP-II land use map, and Fig. <xref ref-type="fig" rid="F1"/>c shows that it displays more croplands over the Ebro domain.</p>
      <p id="d2e1388">In order to evaluate the benefits of assimilating TROPOMI SIF data, we ran an open-loop simulation from 1 January 2018 onwards. This provides a reference point for determining the impact of assimilating SIF data, and it will henceforth be referred to as the “OL experiment” in Table <xref ref-type="table" rid="T1"/>. Another reference experiment we use is the baseline experiment, which is referred to as the LAI<sub>20</sub> experiment in Table <xref ref-type="table" rid="T1"/>. The LAI<sub>20</sub> experiment will use the LDAS to assimilate LAI-V1 from 1 January 2018 to June 2020, and provide a state-of-the-art analysis to evaluate the specific benefits of assimilating SIF data. The initial conditions were spun up for 20 years and shared by these to reference experiment and all SIF assimilation experiment described thereafter. To determine the amount of SIF observation error to consider, three different experiments are run with three different amounts of error: (i) 0.1 mW m<sup>−2</sup> sr<sup>−1</sup> nm<sup>−1</sup>, (ii) 20 % of relative error, and (iii) a combination of the previous two errors with a fixed error of 0.1 mW m<sup>−2</sup> sr<sup>−1</sup> nm<sup>−1</sup> for SIF values below 0.5 mW m<sup>−2</sup> sr<sup>−1</sup> nm<sup>−1</sup> and 20 % of relative error for values above 0.5 mW m<sup>−2</sup> sr<sup>−1</sup> nm<sup>−1</sup>. This configuration will be denoted FR thereafter. Two additional experiments are run to test the co-assimilation of TROPOMI SIF and CLMS LAI 300 m raster for two different values of the SIF observation error and 20 % of relative error for the LAI <xref ref-type="bibr" rid="bib1.bibx6" id="paren.46"/>. All experiment configurations are summarised in Table <xref ref-type="table" rid="T1"/>.</p>

<table-wrap id="T1" specific-use="star"><label>Table 1</label><caption><p id="d2e1568">Overview of the assimilation experiments conducted for TROPOMI SIF on the Ebro basin domain. The table indicates the name of the assimilation experiment (Experiment name), the assimilation of the LAI and which dataset (LAI) with an observation error of 20 %, the assimilation of TROPOMI SIF (SIF) with wich observation error (<inline-formula><mml:math id="M47" display="inline"><mml:mi mathvariant="italic">σ</mml:mi></mml:math></inline-formula>) used.</p></caption><oasis:table frame="topbot"><oasis:tgroup cols="8">
     <oasis:colspec colnum="1" colname="col1" align="left"/>
     <oasis:colspec colnum="2" colname="col2" align="left"/>
     <oasis:colspec colnum="3" colname="col3" align="left" colsep="1"/>
     <oasis:colspec colnum="4" colname="col4" align="center"/>
     <oasis:colspec colnum="5" colname="col5" align="center" colsep="1"/>
     <oasis:colspec colnum="6" colname="col6" align="center"/>
     <oasis:colspec colnum="7" colname="col7" align="center"/>
     <oasis:colspec colnum="8" colname="col8" align="center"/>
     <oasis:thead>
       <oasis:row rowsep="1">
         <oasis:entry namest="col1" nameend="col3" align="center"/>
         <oasis:entry namest="col4" nameend="col5" colsep="1">LAI (m<sup>2</sup> m<sup>−2</sup>) </oasis:entry>
         <oasis:entry namest="col6" nameend="col8">SIF (mW m<sup>−2</sup> sr<sup>−1</sup> nm<sup>−1</sup>) </oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry namest="col1" nameend="col3" align="center" colsep="1">Observation assimilated </oasis:entry>
         <oasis:entry colname="col4">LAI-V1</oasis:entry>
         <oasis:entry colname="col5">LAI-300</oasis:entry>
         <oasis:entry namest="col6" nameend="col8">TROPOMI SIF </oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry rowsep="1" namest="col1" nameend="col3" align="center" colsep="1">Observation error </oasis:entry>
         <oasis:entry colname="col4"><inline-formula><mml:math id="M53" display="inline"><mml:mrow><mml:mi mathvariant="italic">σ</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">20</mml:mn></mml:mrow></mml:math></inline-formula> %</oasis:entry>
         <oasis:entry colname="col5"><inline-formula><mml:math id="M54" display="inline"><mml:mrow><mml:mi mathvariant="italic">σ</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">20</mml:mn></mml:mrow></mml:math></inline-formula> %</oasis:entry>
         <oasis:entry colname="col6"><inline-formula><mml:math id="M55" display="inline"><mml:mrow><mml:mi mathvariant="italic">σ</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0.1</mml:mn></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col7"><inline-formula><mml:math id="M56" display="inline"><mml:mrow><mml:mi mathvariant="italic">σ</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">20</mml:mn></mml:mrow></mml:math></inline-formula> %</oasis:entry>
         <oasis:entry colname="col8"><inline-formula><mml:math id="M57" display="inline"><mml:mrow><mml:mi mathvariant="italic">σ</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">0.1</mml:mn></mml:mrow></mml:math></inline-formula>, SIF <inline-formula><mml:math id="M58" display="inline"><mml:mo>≤</mml:mo></mml:math></inline-formula> 0.5</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Experiment</oasis:entry>
         <oasis:entry colname="col2">Start Date</oasis:entry>
         <oasis:entry colname="col3">End Date</oasis:entry>
         <oasis:entry colname="col4"/>
         <oasis:entry colname="col5"/>
         <oasis:entry colname="col6"/>
         <oasis:entry colname="col7"/>
         <oasis:entry colname="col8"><inline-formula><mml:math id="M59" display="inline"><mml:mrow><mml:mi mathvariant="italic">σ</mml:mi><mml:mo>=</mml:mo><mml:mn mathvariant="normal">20</mml:mn></mml:mrow></mml:math></inline-formula> %, SIF <inline-formula><mml:math id="M60" display="inline"><mml:mi mathvariant="italic">&gt;</mml:mi></mml:math></inline-formula> 0.5</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">name</oasis:entry>
         <oasis:entry colname="col2">(DD/MM/YYYY)</oasis:entry>
         <oasis:entry colname="col3">(DD/MM/YYYY)</oasis:entry>
         <oasis:entry colname="col4"/>
         <oasis:entry colname="col5"/>
         <oasis:entry colname="col6"/>
         <oasis:entry colname="col7"/>
         <oasis:entry colname="col8"/>
       </oasis:row>
     </oasis:thead>
     <oasis:tbody>
       <oasis:row>
         <oasis:entry colname="col1">OL</oasis:entry>
         <oasis:entry colname="col2">01/01/2018</oasis:entry>
         <oasis:entry colname="col3">31/12/2021</oasis:entry>
         <oasis:entry colname="col4">–</oasis:entry>
         <oasis:entry colname="col5">–</oasis:entry>
         <oasis:entry colname="col6">–</oasis:entry>
         <oasis:entry colname="col7">–</oasis:entry>
         <oasis:entry colname="col8">–</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">LAI<sub>20</sub></oasis:entry>
         <oasis:entry colname="col2">01/01/2018</oasis:entry>
         <oasis:entry colname="col3">31/05/2020</oasis:entry>
         <oasis:entry colname="col4">X</oasis:entry>
         <oasis:entry colname="col5">–</oasis:entry>
         <oasis:entry colname="col6">–</oasis:entry>
         <oasis:entry colname="col7">–</oasis:entry>
         <oasis:entry colname="col8">–</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">SIF<sub>01</sub></oasis:entry>
         <oasis:entry colname="col2">01/06/2018</oasis:entry>
         <oasis:entry colname="col3">31/12/2021</oasis:entry>
         <oasis:entry colname="col4">–</oasis:entry>
         <oasis:entry colname="col5">–</oasis:entry>
         <oasis:entry colname="col6">X</oasis:entry>
         <oasis:entry colname="col7">–</oasis:entry>
         <oasis:entry colname="col8">–</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">SIF<sub>20</sub></oasis:entry>
         <oasis:entry colname="col2">01/06/2018</oasis:entry>
         <oasis:entry colname="col3">31/12/2021</oasis:entry>
         <oasis:entry colname="col4">–</oasis:entry>
         <oasis:entry colname="col5">–</oasis:entry>
         <oasis:entry colname="col6">–</oasis:entry>
         <oasis:entry colname="col7">X</oasis:entry>
         <oasis:entry colname="col8">–</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">SIF<sub>FR</sub></oasis:entry>
         <oasis:entry colname="col2">01/06/2018</oasis:entry>
         <oasis:entry colname="col3">31/12/2021</oasis:entry>
         <oasis:entry colname="col4">–</oasis:entry>
         <oasis:entry colname="col5">–</oasis:entry>
         <oasis:entry colname="col6">–</oasis:entry>
         <oasis:entry colname="col7">–</oasis:entry>
         <oasis:entry colname="col8">X</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">SL<sub>20</sub></oasis:entry>
         <oasis:entry colname="col2">01/01/2018</oasis:entry>
         <oasis:entry colname="col3">31/12/2021</oasis:entry>
         <oasis:entry colname="col4">–</oasis:entry>
         <oasis:entry colname="col5">X</oasis:entry>
         <oasis:entry colname="col6">–</oasis:entry>
         <oasis:entry colname="col7">X</oasis:entry>
         <oasis:entry colname="col8">–</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">SL<sub>FR</sub></oasis:entry>
         <oasis:entry colname="col2">01/01/2018</oasis:entry>
         <oasis:entry colname="col3">31/12/2021</oasis:entry>
         <oasis:entry colname="col4">–</oasis:entry>
         <oasis:entry colname="col5">X</oasis:entry>
         <oasis:entry colname="col6">–</oasis:entry>
         <oasis:entry colname="col7">–</oasis:entry>
         <oasis:entry colname="col8">X</oasis:entry>
       </oasis:row>
     </oasis:tbody>
   </oasis:tgroup></oasis:table></table-wrap>

      <p id="d2e2083">The efficiency with which TROPOMI SIF is assimilated into ISBA using the NN operator will be assessed by computing the root mean square error (RMSE) and the Pearson’s correlation coefficient between the analysed fields and the observed values (i.e. LAI and GPP). To ensure the statistical relevance of these evaluations, a <inline-formula><mml:math id="M67" display="inline"><mml:mi>p</mml:mi></mml:math></inline-formula>-value is computed using a Fisher's test when calculating the various Pearson correlations. Any results with a <inline-formula><mml:math id="M68" display="inline"><mml:mi>p</mml:mi></mml:math></inline-formula>-value greater than 0.01 are removed.</p>
</sec>
</sec>
<sec id="Ch1.S4">
  <label>4</label><title>Results</title>
<sec id="Ch1.S4.SS1">
  <label>4.1</label><title>Validation of the observation operator</title>
      <p id="d2e2116">The correlation between TROPOMI SIF and the estimate from the operator is above 0.8 and the RMSE on SIF is around 0.15 mWm<sup>−2</sup> sr<sup>−1</sup> nm<sup>−1</sup> as displayed in Table <xref ref-type="table" rid="T2"/>. However, the neural network's range is smaller than that of the TROPOMI SIF. Both the values closest to and farthest from zero are miss-estimated by the NN. To improve our understanding of the neural network's behaviour, we evaluate its performance on specific TROPOMI SIF samples. We also check whether the neural network's behaviour is sensitive to the range of TROPOMI SIF values and the dominant PFT, in order to ascertain whether it learns features relating to vegetation distribution and phenology. Table <xref ref-type="table" rid="T2"/> shows the results for both the training and test datasets to detect any overfitting.</p>

<table-wrap id="T2" specific-use="star"><label>Table 2</label><caption><p id="d2e2162">Evaluation of the neural network over the train and the test dataset. The percentiles are taken from the range of TROPOMI SIF values. RMSE, ubRMSE, Bias, Median, Minimal and maximal values of the differences between estimated SIF and observation are given in mW m<sup>−2</sup> sr<sup>−1</sup> nm<sup>−1</sup>. Bold values are when the sample is equal to the full dataset (Train or Test).</p></caption><oasis:table frame="topbot"><oasis:tgroup cols="17">
     <oasis:colspec colnum="1" colname="col1" align="left"/>
     <oasis:colspec colnum="2" colname="col2" align="right"/>
     <oasis:colspec colnum="3" colname="col3" align="right" colsep="1"/>
     <oasis:colspec colnum="4" colname="col4" align="right"/>
     <oasis:colspec colnum="5" colname="col5" align="right" colsep="1"/>
     <oasis:colspec colnum="6" colname="col6" align="right"/>
     <oasis:colspec colnum="7" colname="col7" align="right" colsep="1"/>
     <oasis:colspec colnum="8" colname="col8" align="right"/>
     <oasis:colspec colnum="9" colname="col9" align="right" colsep="1"/>
     <oasis:colspec colnum="10" colname="col10" align="right"/>
     <oasis:colspec colnum="11" colname="col11" align="right" colsep="1"/>
     <oasis:colspec colnum="12" colname="col12" align="right"/>
     <oasis:colspec colnum="13" colname="col13" align="right" colsep="1"/>
     <oasis:colspec colnum="14" colname="col14" align="right"/>
     <oasis:colspec colnum="15" colname="col15" align="right" colsep="1"/>
     <oasis:colspec colnum="16" colname="col16" align="right"/>
     <oasis:colspec colnum="17" colname="col17" align="right"/>
     <oasis:thead>
       <oasis:row>
         <oasis:entry colname="col1">Skill Scores</oasis:entry>
         <oasis:entry namest="col2" nameend="col3" align="center" colsep="1">RMSE </oasis:entry>
         <oasis:entry namest="col4" nameend="col5" align="center" colsep="1"><inline-formula><mml:math id="M75" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">p</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry namest="col6" nameend="col7" align="center" colsep="1">Bias  </oasis:entry>
         <oasis:entry namest="col8" nameend="col9" align="center" colsep="1">ubRMSE </oasis:entry>
         <oasis:entry namest="col10" nameend="col11" align="center" colsep="1">Median  </oasis:entry>
         <oasis:entry namest="col12" nameend="col13" align="center" colsep="1">Min </oasis:entry>
         <oasis:entry namest="col14" nameend="col15" align="center" colsep="1">Max </oasis:entry>
         <oasis:entry namest="col16" nameend="col17" align="center">Sample size </oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1"/>
         <oasis:entry namest="col2" nameend="col3" align="center" colsep="1"/>
         <oasis:entry namest="col4" nameend="col5" align="center" colsep="1"/>
         <oasis:entry namest="col6" nameend="col7" align="center" colsep="1">(NN-Obs) </oasis:entry>
         <oasis:entry namest="col8" nameend="col9" align="center" colsep="1"/>
         <oasis:entry namest="col10" nameend="col11" align="center" colsep="1">(NN-Obs) </oasis:entry>
         <oasis:entry namest="col12" nameend="col13" align="center" colsep="1">(NN-Obs) </oasis:entry>
         <oasis:entry namest="col14" nameend="col15" align="center" colsep="1">(NN-Obs) </oasis:entry>
         <oasis:entry namest="col16" nameend="col17" align="center"/>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">Samples</oasis:entry>
         <oasis:entry colname="col2">Train</oasis:entry>
         <oasis:entry colname="col3">Test</oasis:entry>
         <oasis:entry colname="col4">Train</oasis:entry>
         <oasis:entry colname="col5">Test</oasis:entry>
         <oasis:entry colname="col6">Train</oasis:entry>
         <oasis:entry colname="col7">Test</oasis:entry>
         <oasis:entry colname="col8">Train</oasis:entry>
         <oasis:entry colname="col9">Test</oasis:entry>
         <oasis:entry colname="col10">Train</oasis:entry>
         <oasis:entry colname="col11">Test</oasis:entry>
         <oasis:entry colname="col12">Train</oasis:entry>
         <oasis:entry colname="col13">Test</oasis:entry>
         <oasis:entry colname="col14">Train</oasis:entry>
         <oasis:entry colname="col15">Test</oasis:entry>
         <oasis:entry colname="col16">Train</oasis:entry>
         <oasis:entry colname="col17">Test</oasis:entry>
       </oasis:row>
     </oasis:thead>
     <oasis:tbody>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">Percentile [25,75]</oasis:entry>
         <oasis:entry colname="col2">0.11</oasis:entry>
         <oasis:entry colname="col3">0.12</oasis:entry>
         <oasis:entry colname="col4">0.56</oasis:entry>
         <oasis:entry colname="col5">0.53</oasis:entry>
         <oasis:entry colname="col6"><inline-formula><mml:math id="M76" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.00</oasis:entry>
         <oasis:entry colname="col7"><inline-formula><mml:math id="M77" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.01</oasis:entry>
         <oasis:entry colname="col8">0.11</oasis:entry>
         <oasis:entry colname="col9">0.12</oasis:entry>
         <oasis:entry colname="col10"><inline-formula><mml:math id="M78" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.02</oasis:entry>
         <oasis:entry colname="col11"><inline-formula><mml:math id="M79" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.02</oasis:entry>
         <oasis:entry colname="col12"><inline-formula><mml:math id="M80" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.40</oasis:entry>
         <oasis:entry colname="col13"><inline-formula><mml:math id="M81" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.43</oasis:entry>
         <oasis:entry colname="col14">1.82</oasis:entry>
         <oasis:entry colname="col15">1.79</oasis:entry>
         <oasis:entry colname="col16">7 156 450</oasis:entry>
         <oasis:entry colname="col17">7 011 925</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">Percentile [10,90]</oasis:entry>
         <oasis:entry colname="col2">0.12</oasis:entry>
         <oasis:entry colname="col3">0.12</oasis:entry>
         <oasis:entry colname="col4">0.74</oasis:entry>
         <oasis:entry colname="col5">0.73</oasis:entry>
         <oasis:entry colname="col6"><inline-formula><mml:math id="M82" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.01</oasis:entry>
         <oasis:entry colname="col7"><inline-formula><mml:math id="M83" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.02</oasis:entry>
         <oasis:entry colname="col8">0.11</oasis:entry>
         <oasis:entry colname="col9">0.12</oasis:entry>
         <oasis:entry colname="col10"><inline-formula><mml:math id="M84" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.02</oasis:entry>
         <oasis:entry colname="col11"><inline-formula><mml:math id="M85" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.02</oasis:entry>
         <oasis:entry colname="col12"><inline-formula><mml:math id="M86" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.60</oasis:entry>
         <oasis:entry colname="col13"><inline-formula><mml:math id="M87" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.64</oasis:entry>
         <oasis:entry colname="col14">1.82</oasis:entry>
         <oasis:entry colname="col15">1.79</oasis:entry>
         <oasis:entry colname="col16">11 450 319</oasis:entry>
         <oasis:entry colname="col17">11 219 078</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">Percentile [5,95]</oasis:entry>
         <oasis:entry colname="col2">0.12</oasis:entry>
         <oasis:entry colname="col3">0.13</oasis:entry>
         <oasis:entry colname="col4">0.79</oasis:entry>
         <oasis:entry colname="col5">0.77</oasis:entry>
         <oasis:entry colname="col6"><inline-formula><mml:math id="M88" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.02</oasis:entry>
         <oasis:entry colname="col7"><inline-formula><mml:math id="M89" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.02</oasis:entry>
         <oasis:entry colname="col8">0.12</oasis:entry>
         <oasis:entry colname="col9">0.13</oasis:entry>
         <oasis:entry colname="col10"><inline-formula><mml:math id="M90" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.01</oasis:entry>
         <oasis:entry colname="col11"><inline-formula><mml:math id="M91" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.02</oasis:entry>
         <oasis:entry colname="col12"><inline-formula><mml:math id="M92" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.71</oasis:entry>
         <oasis:entry colname="col13"><inline-formula><mml:math id="M93" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.76</oasis:entry>
         <oasis:entry colname="col14">1.82</oasis:entry>
         <oasis:entry colname="col15">1.79</oasis:entry>
         <oasis:entry colname="col16">12 881 610</oasis:entry>
         <oasis:entry colname="col17">12 621 462</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">Percentile [1,99]</oasis:entry>
         <oasis:entry colname="col2">0.13</oasis:entry>
         <oasis:entry colname="col3">0.14</oasis:entry>
         <oasis:entry colname="col4">0.81</oasis:entry>
         <oasis:entry colname="col5">0.81</oasis:entry>
         <oasis:entry colname="col6"><inline-formula><mml:math id="M94" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.02</oasis:entry>
         <oasis:entry colname="col7"><inline-formula><mml:math id="M95" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.03</oasis:entry>
         <oasis:entry colname="col8">0.13</oasis:entry>
         <oasis:entry colname="col9">0.14</oasis:entry>
         <oasis:entry colname="col10"><inline-formula><mml:math id="M96" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.01</oasis:entry>
         <oasis:entry colname="col11"><inline-formula><mml:math id="M97" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.02</oasis:entry>
         <oasis:entry colname="col12"><inline-formula><mml:math id="M98" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.94</oasis:entry>
         <oasis:entry colname="col13"><inline-formula><mml:math id="M99" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.98</oasis:entry>
         <oasis:entry colname="col14">1.82</oasis:entry>
         <oasis:entry colname="col15">1.79</oasis:entry>
         <oasis:entry colname="col16">14 026 641</oasis:entry>
         <oasis:entry colname="col17">13 743 370</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1"><bold>All</bold></oasis:entry>
         <oasis:entry colname="col2"><bold>0.14</bold></oasis:entry>
         <oasis:entry colname="col3"><bold>0.15</bold></oasis:entry>
         <oasis:entry colname="col4"><bold>0.82</bold></oasis:entry>
         <oasis:entry colname="col5"><bold>0.81</bold></oasis:entry>
         <oasis:entry colname="col6"><inline-formula><mml:math id="M100" display="inline"><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="bold">0.03</mml:mn></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col7"><inline-formula><mml:math id="M101" display="inline"><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="bold">0.03</mml:mn></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col8"><bold>0.14</bold></oasis:entry>
         <oasis:entry colname="col9"><bold>0.15</bold></oasis:entry>
         <oasis:entry colname="col10"><inline-formula><mml:math id="M102" display="inline"><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="bold">0.01</mml:mn></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col11"><inline-formula><mml:math id="M103" display="inline"><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="bold">0.02</mml:mn></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col12"><inline-formula><mml:math id="M104" display="inline"><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="bold">3.59</mml:mn></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col13"><inline-formula><mml:math id="M105" display="inline"><mml:mrow><mml:mo>-</mml:mo><mml:mn mathvariant="bold">3.82</mml:mn></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col14"><bold>1.82</bold></oasis:entry>
         <oasis:entry colname="col15"><bold>1.79</bold></oasis:entry>
         <oasis:entry colname="col16"><bold>14 312 899</bold></oasis:entry>
         <oasis:entry colname="col17"><bold>14 023 848</bold></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Deciduous Broad-</oasis:entry>
         <oasis:entry colname="col2">0.15</oasis:entry>
         <oasis:entry colname="col3">0.17</oasis:entry>
         <oasis:entry colname="col4">0.82</oasis:entry>
         <oasis:entry colname="col5">0.80</oasis:entry>
         <oasis:entry colname="col6"><inline-formula><mml:math id="M106" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.03</oasis:entry>
         <oasis:entry colname="col7"><inline-formula><mml:math id="M107" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.03</oasis:entry>
         <oasis:entry colname="col8">0.15</oasis:entry>
         <oasis:entry colname="col9">0.16</oasis:entry>
         <oasis:entry colname="col10"><inline-formula><mml:math id="M108" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.02</oasis:entry>
         <oasis:entry colname="col11"><inline-formula><mml:math id="M109" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.02</oasis:entry>
         <oasis:entry colname="col12"><inline-formula><mml:math id="M110" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>3.44</oasis:entry>
         <oasis:entry colname="col13"><inline-formula><mml:math id="M111" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>3.82</oasis:entry>
         <oasis:entry colname="col14">1.57</oasis:entry>
         <oasis:entry colname="col15">1.74</oasis:entry>
         <oasis:entry colname="col16">1 542 837</oasis:entry>
         <oasis:entry colname="col17">1 319 870</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">leaves Forest</oasis:entry>
         <oasis:entry colname="col2"/>
         <oasis:entry colname="col3"/>
         <oasis:entry colname="col4"/>
         <oasis:entry colname="col5"/>
         <oasis:entry colname="col6"/>
         <oasis:entry colname="col7"/>
         <oasis:entry colname="col8"/>
         <oasis:entry colname="col9"/>
         <oasis:entry colname="col10"/>
         <oasis:entry colname="col11"/>
         <oasis:entry colname="col12"/>
         <oasis:entry colname="col13"/>
         <oasis:entry colname="col14"/>
         <oasis:entry colname="col15"/>
         <oasis:entry colname="col16"/>
         <oasis:entry colname="col17"/>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Coniferous Needle-</oasis:entry>
         <oasis:entry colname="col2">0.13</oasis:entry>
         <oasis:entry colname="col3">0.15</oasis:entry>
         <oasis:entry colname="col4">0.71</oasis:entry>
         <oasis:entry colname="col5">0.65</oasis:entry>
         <oasis:entry colname="col6"><inline-formula><mml:math id="M112" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.01</oasis:entry>
         <oasis:entry colname="col7"><inline-formula><mml:math id="M113" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.01</oasis:entry>
         <oasis:entry colname="col8">0.13</oasis:entry>
         <oasis:entry colname="col9">0.15</oasis:entry>
         <oasis:entry colname="col10">0.0</oasis:entry>
         <oasis:entry colname="col11">0.0</oasis:entry>
         <oasis:entry colname="col12"><inline-formula><mml:math id="M114" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>3.59</oasis:entry>
         <oasis:entry colname="col13"><inline-formula><mml:math id="M115" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>3.64</oasis:entry>
         <oasis:entry colname="col14">1.82</oasis:entry>
         <oasis:entry colname="col15">1.79</oasis:entry>
         <oasis:entry colname="col16">1 897 962</oasis:entry>
         <oasis:entry colname="col17">1 667 631</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">leaves Forest</oasis:entry>
         <oasis:entry colname="col2"/>
         <oasis:entry colname="col3"/>
         <oasis:entry colname="col4"/>
         <oasis:entry colname="col5"/>
         <oasis:entry colname="col6"/>
         <oasis:entry colname="col7"/>
         <oasis:entry colname="col8"/>
         <oasis:entry colname="col9"/>
         <oasis:entry colname="col10"/>
         <oasis:entry colname="col11"/>
         <oasis:entry colname="col12"/>
         <oasis:entry colname="col13"/>
         <oasis:entry colname="col14"/>
         <oasis:entry colname="col15"/>
         <oasis:entry colname="col16"/>
         <oasis:entry colname="col17"/>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">C3 crops</oasis:entry>
         <oasis:entry colname="col2">0.14</oasis:entry>
         <oasis:entry colname="col3">0.15</oasis:entry>
         <oasis:entry colname="col4">0.84</oasis:entry>
         <oasis:entry colname="col5">0.83</oasis:entry>
         <oasis:entry colname="col6"><inline-formula><mml:math id="M116" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.04</oasis:entry>
         <oasis:entry colname="col7"><inline-formula><mml:math id="M117" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.05</oasis:entry>
         <oasis:entry colname="col8">0.13</oasis:entry>
         <oasis:entry colname="col9">0.14</oasis:entry>
         <oasis:entry colname="col10"><inline-formula><mml:math id="M118" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.02</oasis:entry>
         <oasis:entry colname="col11"><inline-formula><mml:math id="M119" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.03</oasis:entry>
         <oasis:entry colname="col12"><inline-formula><mml:math id="M120" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>3.55</oasis:entry>
         <oasis:entry colname="col13"><inline-formula><mml:math id="M121" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>3.45</oasis:entry>
         <oasis:entry colname="col14">1.28</oasis:entry>
         <oasis:entry colname="col15">1.55</oasis:entry>
         <oasis:entry colname="col16">5 730 186</oasis:entry>
         <oasis:entry colname="col17">5 804 070</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">C4 crops</oasis:entry>
         <oasis:entry colname="col2">0.14</oasis:entry>
         <oasis:entry colname="col3">0.14</oasis:entry>
         <oasis:entry colname="col4">0.87</oasis:entry>
         <oasis:entry colname="col5">0.87</oasis:entry>
         <oasis:entry colname="col6"><inline-formula><mml:math id="M122" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.04</oasis:entry>
         <oasis:entry colname="col7"><inline-formula><mml:math id="M123" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.05</oasis:entry>
         <oasis:entry colname="col8">0.13</oasis:entry>
         <oasis:entry colname="col9">0.13</oasis:entry>
         <oasis:entry colname="col10"><inline-formula><mml:math id="M124" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.02</oasis:entry>
         <oasis:entry colname="col11"><inline-formula><mml:math id="M125" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.03</oasis:entry>
         <oasis:entry colname="col12"><inline-formula><mml:math id="M126" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>2.81</oasis:entry>
         <oasis:entry colname="col13"><inline-formula><mml:math id="M127" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>2.78</oasis:entry>
         <oasis:entry colname="col14">0.67</oasis:entry>
         <oasis:entry colname="col15">0.75</oasis:entry>
         <oasis:entry colname="col16">207 755</oasis:entry>
         <oasis:entry colname="col17">222 823</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">Irrigated Crops</oasis:entry>
         <oasis:entry colname="col2">0.15</oasis:entry>
         <oasis:entry colname="col3">0.14</oasis:entry>
         <oasis:entry colname="col4">0.77</oasis:entry>
         <oasis:entry colname="col5">0.77</oasis:entry>
         <oasis:entry colname="col6"><inline-formula><mml:math id="M128" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.05</oasis:entry>
         <oasis:entry colname="col7"><inline-formula><mml:math id="M129" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.04</oasis:entry>
         <oasis:entry colname="col8">0.14</oasis:entry>
         <oasis:entry colname="col9">0.14</oasis:entry>
         <oasis:entry colname="col10"><inline-formula><mml:math id="M130" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.03</oasis:entry>
         <oasis:entry colname="col11"><inline-formula><mml:math id="M131" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.03</oasis:entry>
         <oasis:entry colname="col12"><inline-formula><mml:math id="M132" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>1.25</oasis:entry>
         <oasis:entry colname="col13"><inline-formula><mml:math id="M133" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>1.41</oasis:entry>
         <oasis:entry colname="col14">0.60</oasis:entry>
         <oasis:entry colname="col15">0.67</oasis:entry>
         <oasis:entry colname="col16">102 170</oasis:entry>
         <oasis:entry colname="col17">107 158</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">C3 grasslands</oasis:entry>
         <oasis:entry colname="col2">0.14</oasis:entry>
         <oasis:entry colname="col3">0.15</oasis:entry>
         <oasis:entry colname="col4">0.82</oasis:entry>
         <oasis:entry colname="col5">0.82</oasis:entry>
         <oasis:entry colname="col6"><inline-formula><mml:math id="M134" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.02</oasis:entry>
         <oasis:entry colname="col7"><inline-formula><mml:math id="M135" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.03</oasis:entry>
         <oasis:entry colname="col8">0.14</oasis:entry>
         <oasis:entry colname="col9">0.15</oasis:entry>
         <oasis:entry colname="col10"><inline-formula><mml:math id="M136" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.01</oasis:entry>
         <oasis:entry colname="col11"><inline-formula><mml:math id="M137" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.01</oasis:entry>
         <oasis:entry colname="col12"><inline-formula><mml:math id="M138" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>3.34</oasis:entry>
         <oasis:entry colname="col13"><inline-formula><mml:math id="M139" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>3.26</oasis:entry>
         <oasis:entry colname="col14">1.23</oasis:entry>
         <oasis:entry colname="col15">1.33</oasis:entry>
         <oasis:entry colname="col16">4 617 598</oasis:entry>
         <oasis:entry colname="col17">4 716 617</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Peats and Wetlands</oasis:entry>
         <oasis:entry colname="col2">0.16</oasis:entry>
         <oasis:entry colname="col3">0.17</oasis:entry>
         <oasis:entry colname="col4">0.82</oasis:entry>
         <oasis:entry colname="col5">0.78</oasis:entry>
         <oasis:entry colname="col6"><inline-formula><mml:math id="M140" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.04</oasis:entry>
         <oasis:entry colname="col7"><inline-formula><mml:math id="M141" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.04</oasis:entry>
         <oasis:entry colname="col8">0.16</oasis:entry>
         <oasis:entry colname="col9">0.17</oasis:entry>
         <oasis:entry colname="col10"><inline-formula><mml:math id="M142" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.02</oasis:entry>
         <oasis:entry colname="col11"><inline-formula><mml:math id="M143" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>0.02</oasis:entry>
         <oasis:entry colname="col12"><inline-formula><mml:math id="M144" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>2.67</oasis:entry>
         <oasis:entry colname="col13"><inline-formula><mml:math id="M145" display="inline"><mml:mo>-</mml:mo></mml:math></inline-formula>2.89</oasis:entry>
         <oasis:entry colname="col14">1.53</oasis:entry>
         <oasis:entry colname="col15">1.37</oasis:entry>
         <oasis:entry colname="col16">214 391</oasis:entry>
         <oasis:entry colname="col17">185 679</oasis:entry>
       </oasis:row>
     </oasis:tbody>
   </oasis:tgroup></oasis:table></table-wrap>

      <p id="d2e3568">As can be seen in bold in Table <xref ref-type="table" rid="T2"/>, the overall skill score values obtained for the training and test datasets are similar. This suggests that if there is any overfitting, it is limited. Comparing the skill scores for the different percentiles of TROPOMI SIF values, we can see that, as we consider more data from the edges of the distribution, the bias (i.e. the mean of the NN estimates minus their observed counterparts), the RMSE and the correlation all increase. This indicates that the neural network disregards minor variations around the mean TROPOMI SIF value. This behaviour is to be expected due to the use of Huber loss during training, which focuses on minimising the RMSE. Furthermore, when the values are farther from the mean, the neural network follows this change, thereby increasing the correlation. However, the bias also increases with the SIF value, thus increasing the RMSE.</p>
      <p id="d2e3574">A negative bias means that the neural network is underestimating the TROPOMI SIF value. This is particularly evident for the largest SIF values. The neural network performs better over areas where crops, grasslands and deciduous broadleaf forests are most prevalent. Coniferous needle-leaf forests (CNF) and flooded areas (irrigated crops) seem to present more difficulty, as Table <xref ref-type="table" rid="T2"/> shows.</p>
      <p id="d2e3579">The differences in behaviour depending on the SIF values and the PFT motivate us to verify whether some spatial properties need to be considered in our evaluation. Figure <xref ref-type="fig" rid="F4"/> shows the local values of the root mean square error (RMSE), correlation, number of available comparisons and normalised root mean square error (NRMSE) for the entire test dataset. The normalisation is done by the TROPOMI SIF value. The correlation is above 0.8 in the area south of 60° N. This is due to the number of observations available in the region. The northern area is not observed for more than half of the year by TROPOMI around the winter period and less LAI observation are available.</p>

      <fig id="F4"><label>Figure 4</label><caption><p id="d2e3586">Grid-pointwise evaluation of the performance of the match between NN and observations for the test dataset. From top to bottom: the RMSE, the number of comparison per pixels, the NRMSE, and the Pearson correlation. Units are indicated where applicable.</p></caption>
          <graphic xlink:href="https://bg.copernicus.org/articles/23/6687/2026/bg-23-6687-2026-f04.png"/>

        </fig>

      <p id="d2e3595">Based on the spatial distribution of observations over the course of a year, we can expect approximately one observation every 2 d. The RMSE is heterogeneously distributed, highlighting the known areas of C3 crops, and ranges between 0.1 and 0.2 mW m<sup>2</sup> sr<sup>−1</sup> nm<sup>−1</sup>. The NRMSE distribution is more uniform than the RMSE distribution, averaging at around 20 %. Underestimation appears to be proportional to the values of TROPOMI SIF, so considering a fixed value for the observation error seems inaccurate. That is why, we tested different observation errors.</p>
</sec>
<sec id="Ch1.S4.SS2">
  <label>4.2</label><title>Assimilation in the Ebro basin</title>
      <p id="d2e3639">In the first phase, the NN was trained across the entire European domain. In this section, the saved weights of the neural network are implemented in the assimilation scheme as an observation operator for the SIF. The LAI is simulated and the results are used as input data. In this case, the NN is only used in forward mode, so the weights will not change. Since the observation operator only considers the LAI among the control variables, the accuracy of the LAI representation is the main criterion for evaluating the assimilation results. All assimilation experiments are restricted to the Ebro basin domain.</p>
      <p id="d2e3642">The LAI increments are substantial on the full Ebro basin domain, reaching up to 0.2 m<sup>2</sup> m<sup>−2</sup>, as shown by the monthly-averaged maps over the C3 crops (Fig. <xref ref-type="fig" rid="F5"/>). The residuals, the remaining differences between the analysed SIF and the observed SIF, are smaller than the innovation vectors, the difference between modeled SIF and observed SIF. This illustrates that the assimilation of SIF leads to the identification of such observed agricultural practices in the model, even when they are not simulated. Here, we observe a confined area around the Ebro basin where LAI increased during the summer of 2021. This pattern was also evident in the other years of the experiment. This confined area is artificially irrigated <xref ref-type="bibr" rid="bib1.bibx48" id="paren.47"/>. Irrigation sustains the LAI and SIF, while other areas become drier. This illustrates that the assimilation of SIF leads to the identification of such observed agricultural practices in the model, even when they are not simulated.</p><fig id="F5"><label>Figure 5</label><caption><p id="d2e3673">Mean monthly analysis increments of the SIF<sub>20</sub> experiment on the LAI of C3 crops over the Ebro domain due to SIF assimilation during the year 2021. Grid cells displaying less than 10 % C3 crops are masked.</p></caption>
          <graphic xlink:href="https://bg.copernicus.org/articles/23/6687/2026/bg-23-6687-2026-f05.png"/>

        </fig>

      <p id="d2e3692">This positively impacts on the representation of LAI, as demonstrated by the map of the differences between the RMSEs in Fig. <xref ref-type="fig" rid="F6"/>a, b, c. During the same period, we calculated the difference in correlation with respect to the 10 d LAI-V1 product between the open-loop experiment and the analysis. The resulting map is displayed in Fig. <xref ref-type="fig" rid="F6"/>d, e, f. As only one observation is available every 10 d for calculating the correlation, all cells with a <inline-formula><mml:math id="M152" display="inline"><mml:mi>F</mml:mi></mml:math></inline-formula>-test <inline-formula><mml:math id="M153" display="inline"><mml:mi>p</mml:mi></mml:math></inline-formula>-value higher than 0.01 were removed. This ensures that significant correlation and RMSE values are displayed. This mask is used for the RMSE displayed in Fig. <xref ref-type="fig" rid="F6"/>a, b, c and the correlation panels displayed in Fig. <xref ref-type="fig" rid="F6"/>d, e, f. Figure <xref ref-type="fig" rid="F6"/>a, d show the results averaged over the first full year of TROPOMI SIF assimilation. Due to the small number of days on which both the 10 d LAI-V1 observation (with no temporal interpolation) and TROPOMI SIF were available, many grid cells were discarded by the <inline-formula><mml:math id="M154" display="inline"><mml:mi>F</mml:mi></mml:math></inline-formula>-test. An extensive decrease in RMSE is observed for the remaining grid cells, except in some areas of the French Pyrenees. The improvement in correlation is more heterogeneous, with more areas showing a loss of correlation or no change. The results obtained during the second year of assimilation, displayed in Fig. <xref ref-type="fig" rid="F6"/>b–e, cover a larger area and confirm the results of the first year. Overall, the RMSE decreased except in some specific areas, while the correlation with respect to LAI observations was only around the Ebro river basin. Once the results from both years are combined, Fig. <xref ref-type="fig" rid="F6"/>c, f provides a clearer picture of the improvements brought about by the assimilation of TROPOMI SIF. Overall, the RMSE decreased except in some areas of the Pyrenees mountains, and the correlation increased over the crop areas of the Ebro basin, as shown in Fig. <xref ref-type="fig" rid="F1"/>c, but not in forest and bare soil areas. There were no significant discrepancies between the results of the first and second years of assimilation. Furthermore, a greater number of grid cells passed the <inline-formula><mml:math id="M155" display="inline"><mml:mi>F</mml:mi></mml:math></inline-formula>-test in the second year, confirming that the first year of assimilation matching the year of the data used to train the observation operator is not problematic.</p>

      <fig id="F6" specific-use="star"><label>Figure 6</label><caption><p id="d2e3743">Differences (SIF<sub>20</sub>-OL) over the Ebro domain in <bold>(a, b, c)</bold> RMSE  and in <bold>(d, e, f)</bold> <inline-formula><mml:math id="M157" display="inline"><mml:mrow><mml:msub><mml:mi mathvariant="italic">ρ</mml:mi><mml:mi mathvariant="normal">p</mml:mi></mml:msub></mml:mrow></mml:math></inline-formula> computed with respect to LAI observations after and before assimilation of SIF averaged from: June 2018 to May 2019 <bold>(a–d)</bold>; from June 2019 to May 2020 <bold>(b–e)</bold> and from June 2018 to May 2020 <bold>(c–f)</bold>.</p></caption>
          <graphic xlink:href="https://bg.copernicus.org/articles/23/6687/2026/bg-23-6687-2026-f06.png"/>

        </fig>

      <p id="d2e3788">For the irrigated pixels marked with a cross in Fig. <xref ref-type="fig" rid="F6"/>, the SIF and LAI estimates from the analysis are closer to the observed values in terms of RMSE and correlation. This is evident in the time series of both OL and SIF<sub>20</sub>, which are displayed in Fig. <xref ref-type="fig" rid="F7"/> and were extracted over this specific irrigated grid cell. The LAI and TROPOMI SIF observations displayed in Fig. <xref ref-type="fig" rid="F7"/> all show regrowth of the vegetation in late summer each year, which is not seen by the OL experiment. However, assimilation of TROPOMI SIF (i.e. the SIF<sub>20</sub> analysis results shown in Fig. <xref ref-type="fig" rid="F7"/>) tends to be more accurate. There has been an improvement in terms of the agreement between TROPOMI SIF and LAI. The SIF estimated from the analysis remains smaller than the observed TROPOMI SIF, but captures the temporal trends. As the estimated SIF does not necessarily represent the true SIF, but only the aggregated TROPOMI SIF measurements used, we cannot comment on the physicality of this underestimation in our SIF representation. As highlighted in Sect. S2 of the Supplement, the neural network is not a substitute for a SIF physical model, as demonstrated by the comparison with airborne measurements of SIF over the same area from the HyPLant campaign <xref ref-type="bibr" rid="bib1.bibx47" id="paren.48"/>.</p>

      <fig id="F7" specific-use="star"><label>Figure 7</label><caption><p id="d2e3823">Timeseries extract on the cell locate in 1.05° E, 41.65° N for the SIF <bold>(a, b)</bold>, the LAI <bold>(c, d)</bold> and the the total soil moisture (WGTOT) in the column <bold>(e, f)</bold>. The open loop (OL in Table <xref ref-type="table" rid="T1"/>) indicated with dashed blue lines, SIF<sub>20</sub> (<bold>a, c, e</bold>) and SL<sub>20</sub> (<bold>b, d, f</bold>) analyses with the plain red line in their respective panels and observations with green dots. The observations for the SIF, and LAI are respectively extracted from TROPOMI SIF and LAI-300 described in Sect. 2.2.3.</p></caption>
          <graphic xlink:href="https://bg.copernicus.org/articles/23/6687/2026/bg-23-6687-2026-f07.png"/>

        </fig>

      <p id="d2e3868">Assimilating SIF improves the correlation with FLUXCOM-GPP over the full Ebro domain, as shown in the boxplots in Fig. <xref ref-type="fig" rid="F8"/>. This demonstrates that, on average, the SIF<sub>20</sub> experiment provides an equally accurate representation of GPP as the LAI<sub>20</sub> experiment, and a more accurate representation than the OL experiment. Regarding LAI, Fig. <xref ref-type="fig" rid="F9"/> shows that SIF<sub>20</sub> provides a better RMSE than the OL experiment, as indicated by Fig. <xref ref-type="fig" rid="F6"/>, but this RMSE remains slightly higher on average than the LAI<sub>20</sub> experiment over the Ebro domain.</p>

      <fig id="F8"><label>Figure 8</label><caption><p id="d2e3917">Box plot of the correlation between analysis, open-loop simulated GPP and the FLUXCOM dataset over the Ebro domain. From left to right: Open-loop, SIF<sub>20</sub>, SIF<sub>FR</sub>, Coassimilation SL<sub>20</sub> and SL<sub>FR</sub> configurations and assimilation of LAI-V1. The boxes represent the 25th and the 75th percentiles, the whiskers the 5th and 95th percentiles. The red line represents the 50th percentile and the green triangle the mean.</p></caption>
          <graphic xlink:href="https://bg.copernicus.org/articles/23/6687/2026/bg-23-6687-2026-f08.png"/>

        </fig>

      <fig id="F9"><label>Figure 9</label><caption><p id="d2e3964">Boxplot of the RMSE for the different LAI predictions taken between June 2018 and June 2020 over the Ebro domain. Box represents the 25th and the 75th percentiles, the whiskers the 5th and 95th percentiles. The red line represents the 50th percentile and the green triangle the mean.</p></caption>
          <graphic xlink:href="https://bg.copernicus.org/articles/23/6687/2026/bg-23-6687-2026-f09.png"/>

        </fig>

      <p id="d2e3973">In addition to the experiment with 20 % (SIF<sub>20</sub>), we also tested assimilation with two different error scenarios, namely SIF<sub>01</sub> and SIF<sub>FR</sub>, which is a combination of the two first scenarios.</p>
      <p id="d2e4003">The time series displayed in Fig. <xref ref-type="fig" rid="F10"/> are showing the differences between the Ebro domain averaged RMSE with respect to LAI-V1 10 d observations from the different analyses and the one from the open-loop RMSE. The RMSE for SIF<sub>20</sub> analysis tends to match the RMSE from the reference LAI<sub>20</sub> experiment and is sometimes even lower. In the case of SIF<sub>01</sub> and SIF<sub>FR</sub>, the RMSE is close to that of the open-loop experiment, even though the SIF<sub>FR</sub> configuration provide a better RMSE on LAI than the SIF<sub>01</sub> configuration. This suggests that our first guess (i.e. 20 % of relative error) in terms of observation error is close to the optimal situation for minimising the RMSE, and that higher values tend to render assimilation ineffective. For the SIF<sub>FR</sub>, this skill loss is also visible in terms of correlation with FLUXCOM GPP according to Fig. <xref ref-type="fig" rid="F8"/>. Experiment SIF<sub>20</sub> and LAI<sub>20</sub> achieve similar accuracy in terms of representation of LAI and GPP, yet SIF<sub>20</sub> performances have more variances, but led to five time more observation to be assimilated.</p>

      <fig id="F10"><label>Figure 10</label><caption><p id="d2e4104"><inline-formula><mml:math id="M183" display="inline"><mml:mi mathvariant="normal">Δ</mml:mi></mml:math></inline-formula>RMSE between the Open-Loop and the LAI analysis of the assimilation experiments from January 2018 to January 2020 over the Ebro domain.</p></caption>
          <graphic xlink:href="https://bg.copernicus.org/articles/23/6687/2026/bg-23-6687-2026-f10.png"/>

        </fig>

</sec>
</sec>
<sec id="Ch1.S5">
  <label>5</label><title>Discussion</title>
<sec id="Ch1.S5.SS1">
  <label>5.1</label><title>Is the Co-assimilation of LAI and SIF provide better results than assimilating one of the two?</title>
      <p id="d2e4136">When it comes to data assimilation, the question of redundancy in observations is important, since disagreement between observations can degrade the analysis, while agreement can improve it further. Two configurations of co-assimilation, SL<sub>20</sub> and SL<sub>FR</sub> of the LAI-300 from CLMS and the TROPOMI SIF are run on the Ebro basin domain. We compare the RMSE on LAI for each cell of the assimilation domain averaged from June 2018 to May 2020 for all assimilation experiments and display the results in Fig. <xref ref-type="fig" rid="F9"/>. This co-assimilation configuration achieves an RMSE on LAI that is at least as good as that achieved by assimilating LAI or SIF alone and highlights some synergistic behaviour in the assimilation of both observations. This result is also evident in the correlation with FLUXCOM GPP, as shown in Fig. <xref ref-type="fig" rid="F8"/>. In terms of local behaviour, comparing the SL<inline-formula><mml:math id="M186" display="inline"><mml:mn mathvariant="normal">20</mml:mn></mml:math></inline-formula> time series with the SIF<sub>20</sub> time series in Fig. <xref ref-type="fig" rid="F7"/> shows that co-assimilation leads to much better agreement with observed LAI while keeping SIF close to TROPOMI SIF. This is especially true during spring where the assimilation of SIF alone led to too high LAI. The SL<inline-formula><mml:math id="M188" display="inline"><mml:mn mathvariant="normal">20</mml:mn></mml:math></inline-formula> analysis improves also the representation of the second SIF and LAI peak due to irrigation in summer 2020.</p>
      <p id="d2e4187">While assimilating TROPOMI SIF as the only observation should achieve results similar to the LAI<sub>20</sub> experiment with a relative error of 20 %, both cases of co-assimilation (co-assimilating SIF alongside LAI) achieve better results than assimilating only LAI-V1 or TROPOMI SIF, regardless of the observation error on TROPOMI SIF. Furthermore, the FR configuration in co-assimilation does not degrade the results as much  as when only TROPOMI SIF is assimilated, providing a more consistent improvement than the SL<sub>20</sub> analysis. This demonstrates that co-assimilation of TROPOMI SIF with a 10 d LAI product improves the LAI representation further. The key is that the assimilation of TROPOMI SIF leads to changes in day-to-day monitoring of fluxes and LAI, albeit with some variance. In contrast, the assimilation of unbiased LAI reduces variability around LAI observations. This results in more accurate trends and better agreement with LAI observations and, to a lesser extent, GPP observations than the assimilation of the LAI product alone, as illustrated in Fig. <xref ref-type="fig" rid="F8"/>.</p>

      <fig id="F11" specific-use="star"><label>Figure 11</label><caption><p id="d2e4212">Same as for Fig. <xref ref-type="fig" rid="F7"/> but for the daily mean GPP <bold>(a, b)</bold> and the daily cumulated evapotranspiration (EVAPC) in panels <bold>(c, d)</bold>. The open loop (OL in Table <xref ref-type="table" rid="T1"/>) indicated with dashed blue lines, SIF<sub>20</sub> (<bold>a, c</bold>) and SL<sub>20</sub> (<bold>b, d</bold>) analyses with the plain red line in their respective panels and observations with green dots. The observation of GPP are extracted from FLUXCOM database.</p></caption>
          <graphic xlink:href="https://bg.copernicus.org/articles/23/6687/2026/bg-23-6687-2026-f11.png"/>

        </fig>

</sec>
<sec id="Ch1.S5.SS2">
  <label>5.2</label><title>Does the GPP require additional observations to be better constrained?</title>
      <p id="d2e4264">Unexpectedly, the GPP seems to benefit less from TROPOMI SIF assimilation, even though SIF is known to be well correlated with the GPP. While the GPP is improved on average across the domain by assimilating TROPOMI SIF, the local comparison displayed in Fig. <xref ref-type="fig" rid="F7"/> shows that it is difficult to substantially improve the analysed GPP. It is slightly better in case of coassimilation without solving the inaccuracy in the GPP representation. This result stems from the way the GPP is modelled by the ISBA LSM. The modelled GPP depends not only on the vegetation parameter, but also on the soil water content. Assimilating SIF often results in a more accurate representation of LAI, and in our local case, it adds more LAI in summer than the open-loop model forecasts. This increase in LAI causes the model soil to dry out, as shown by the total soil moisture (WGTOT) in the soil column in Figs. <xref ref-type="fig" rid="F7"/>, <xref ref-type="fig" rid="F11"/>, and increases the evapotranspiration accumulated over the day (EVAPC in Fig. <xref ref-type="fig" rid="F11"/>). Consequently, the computed GPP cannot increase, whereas the observed counterpart (i.e. FLUXCOM) captures this second increase in late summer. FLUXCOM is an observation product based mainly on atmospheric variables, but it also incorporates vegetation and temperature data from MODIS satellites. This remote sensing data helps FLUXCOM to identify these recurrent increases in GPP due to artificial irrigation (see figures in the Supplement, Sect. S3). Assimilating SIF yields the same results as assimilating LAI directly from the GPP perspective. As such, assimilating SIF corrects the LAI and updates the soil water content to balance the change in LAI. However, it cannot infer an increase in soil water content, which would make the model's GPP more similar to the FLUXCOM estimate.</p>
</sec>
</sec>
<sec id="Ch1.S6" sec-type="conclusions">
  <label>6</label><title>Conclusion</title>
      <p id="d2e4284">In this article, we examined the use of a deep learning emulator of TROPOMI SIF as an observation operator, and explored the advantages of incorporating TROPOMI SIF into a land surface model with this observation operator. The neural network operator takes the LAI and three metadata as inputs: the two geographical coordinates and the day of the year. Using this simple setup, we achieved an average RMSE of 0.15 mW m<sup>2</sup> sr<sup>−1</sup> nm<sup>−1</sup> on the TROPOMI SIF averaged estimate, with no evident overfitting. The Pearson correlation coefficient between the neural network estimate and TROPOSIF is greater than 0.8, with higher correlations observed in cropland areas than in coniferous areas. However, the neural network struggles with statistically extreme values, which increases the averaged RMSE and introduces bias. It is also unable to capture small variations close to the expected mean value. Nevertheless, it does capture major changes in the TROPOMI SIF, such as those due to the changing seasons.</p>
      <p id="d2e4320">The neural network was then successfully employed as an observation operator, taking into account the observation error relative to the average TROPOMI SIF value. Several assimilation experiments were conducted over an irrigated cropland area in the Ebro basin. TROPOMI SIF assimilation had a similarly positive impact on LAI as assimilation of the 10 d LAI observation product. However, it should be noted that daily assimilation of TROPOMI SIF yields a higher update rate than assimilation of 10 d LAI observations. Despite not being predicted by the LSM or provided as inputs to the neural network, irrigated areas were well represented. The analysis also improved the representation of both TROPOMI SIF and changes in GPP, which are both important for monitoring the carbon cycle.</p>
      <p id="d2e4323">This study has shown that, when assimilating SIF alone, an observation error of 20 % leads to the best assimilation performance. In a co-assimilation framework incorporating LAI, the representation of vegetation variables (i.e. LAI and GPP) improves further, demonstrating the value of redundancy in observations used in a data assimilation system. However, the best performance is obtained using a more complex error description (FR). Further research is required to quantify observation errors accurately. Unlike the assimilation of LAI, assimilation of TROPOMI SIF leads to a small improvement in GPP representation over unmodelled irrigated areas, since additional water supply is not represented.</p>
      <p id="d2e4326">We demonstrated the advantages of integrating TROPOMI SIF products into a land data assimilation system by applying a deep learning operator. This opens up new possibilities for assimilating new SIF products. Although the neural network operator was designed for TROPOMI SIF products, it can be adapted for other SIF products. We also addressed the co-assimilation of SIF with LAI, a process that could be extended to other satellite products, such as land surface temperature.</p>
</sec>

      
      </body>
    <back><notes notes-type="codedataavailability"><title>Code and data availability</title>

      <p id="d2e4333">The final weights computed for the neural-network, the database used for training are stored in Zenodo repository at <ext-link xlink:href="https://doi.org/10.5281/zenodo.18669437" ext-link-type="DOI">10.5281/zenodo.18669437</ext-link> <xref ref-type="bibr" rid="bib1.bibx54" id="paren.49"/>. The simulations and different analysis resulting from SIF assimilation are available in Zenodo repository at <ext-link xlink:href="https://doi.org/10.5281/zenodo.18668100" ext-link-type="DOI">10.5281/zenodo.18668100</ext-link> <xref ref-type="bibr" rid="bib1.bibx53" id="paren.50"/>. The Hyplant raw measurements are available through LIAISE data portal <uri>https://liaise.aeris-data.fr/about/</uri> (last access: 21 September 2026).</p>
  </notes><app-group>
        <supplementary-material position="anchor"><p id="d2e4351">The supplement related to this article is available online at <inline-supplementary-material xlink:href="https://doi.org/10.5194/bg-23-6687-2026-supplement" xlink:title="pdf">https://doi.org/10.5194/bg-23-6687-2026-supplement</inline-supplementary-material>.</p></supplementary-material>
        </app-group><notes notes-type="authorcontribution"><title>Author contributions</title>

      <p id="d2e4360">PV conducted the investigation and analysed the data, writing the article under the supervision of JCC. JV, BB and ORM helped with the development of the methodology and software. JV, BB, ORM, SG also helped PV validate the results. JCC and PdR administered the project, while CB provided useful advice on the TROPOSIF dataset and UR and BS provided the Hyplant dataset. All co-authors reviewed the article.</p>
  </notes><notes notes-type="competinginterests"><title>Competing interests</title>

      <p id="d2e4366">The contact author has declared that none of the authors has any competing interests.</p>
  </notes><notes notes-type="disclaimer"><title>Disclaimer</title>

      <p id="d2e4372">Publisher's note: Copernicus Publications remains neutral with regard to jurisdictional claims made in the text, published maps, institutional affiliations, or any other geographical representation in this paper. The authors bear the ultimate responsibility for providing appropriate place names. Views expressed in the text are those of the authors and do not necessarily reflect the views of the publisher.</p>
  </notes><ack><title>Acknowledgements</title><p id="d2e4378">This research received funding from European Union Horizon Europe through CORSO project (grant no. 101082194) and CERISE project (grant no. 101082139). Views and opinions expressed are those of the authors only and do not necessarily reflect those of the European Union or the Comission. Neither the European Union nor the granting authority can be held responsible for them. Thanks to the HyPlant airborne campaign funded by CNES, it was possible to make a comparison with independent SIF observations. The authors would like to thank Dr Alban Farchi and the two anonymous reviewers for their helpful comments.</p></ack><notes notes-type="financialsupport"><title>Financial support</title>

      <p id="d2e4383">This research has been supported by the HORIZON EUROPE Digital, Industry and Space (grant nos. 101082194 and 101082139).</p>
  </notes><notes notes-type="reviewstatement"><title>Review statement</title>

      <p id="d2e4390">This paper was edited by Andrew Feldman and reviewed by Alban Farchi and two anonymous referees.</p>
  </notes><ref-list>
    <title>References</title>

      <ref id="bib1.bibx1"><label>Agustí-Panareda et al.(2022)Agustí-Panareda, McNorton, Balsamo, Baier, Bousserez, Boussetta, Brunner, Chevallier, Choulga, Diamantakis et al.</label><mixed-citation>Agustí-Panareda, A., McNorton, J., Balsamo, G., Baier, B. C., Bousserez,  N., Boussetta, S., Brunner, D., Chevallier, F., Choulga, M., Diamantakis, M., Engelen, R., Flemming, J., Granier, C., Guevara, M., Denier van der Gon, H., Elguindi, N., Haussaire, J.-M., Jung, M., Janssens-Maenhout, G., Kivi, R., Massart, S., Papale, D., Parrington, M., Razinger, M., Sweeney, C., Vermeulen, A., and Walther, S.: Global nature run data with realistic high-resolution carbon weather for the year of the Paris agreement, Scientific Data, 9, 160, <ext-link xlink:href="https://doi.org/10.1038/s41597-022-01228-2" ext-link-type="DOI">10.1038/s41597-022-01228-2</ext-link>, 2022.</mixed-citation></ref>
      <ref id="bib1.bibx2"><label>Albergel et al.(2010)Albergel, Calvet, Mahfouf, Rüdiger, Barbu, Lafont, Roujean, Walker, Crapeau, and Wigneron</label><mixed-citation>Albergel, C., Calvet, J.-C., Mahfouf, J.-F., Rüdiger, C., Barbu, A. L., Lafont, S., Roujean, J.-L., Walker, J. P., Crapeau, M., and Wigneron, J.-P.: Monitoring of water and carbon fluxes using a land data assimilation system: a case study for southwestern France, Hydrol. Earth Syst. Sci., 14, 1109–1124, <ext-link xlink:href="https://doi.org/10.5194/hess-14-1109-2010" ext-link-type="DOI">10.5194/hess-14-1109-2010</ext-link>, 2010.</mixed-citation></ref>
      <ref id="bib1.bibx3"><label>Albergel et al.(2017)Albergel, Munier, Leroux, Dewaele, Fairbairn, Barbu, Gelati, Dorigo, Faroux, Meurey, Le Moigne, Decharme, Mahfouf, and Calvet</label><mixed-citation>Albergel, C., Munier, S., Leroux, D. J., Dewaele, H., Fairbairn, D., Barbu, A. L., Gelati, E., Dorigo, W., Faroux, S., Meurey, C., Le Moigne, P., Decharme, B., Mahfouf, J.-F., and Calvet, J.-C.: Sequential assimilation of satellite-derived vegetation and soil moisture products using SURFEX_v8.0: LDAS-Monde assessment over the Euro-Mediterranean area, Geosci. Model Dev., 10, 3889–3912, <ext-link xlink:href="https://doi.org/10.5194/gmd-10-3889-2017" ext-link-type="DOI">10.5194/gmd-10-3889-2017</ext-link>, 2017.</mixed-citation></ref>
      <ref id="bib1.bibx4"><label>Albergel et al.(2020)Albergel, Zheng, Bonan, Dutra, Rodríguez-Fernández, Munier, Draper, de Rosnay, Muñoz Sabater, Balsamo, Fairbairn, Meurey, and Calvet</label><mixed-citation>Albergel, C., Zheng, Y., Bonan, B., Dutra, E., Rodríguez-Fernández, N., Munier, S., Draper, C., de Rosnay, P., Muñoz-Sabater, J., Balsamo, G., Fairbairn, D., Meurey, C., and Calvet, J.-C.: Data assimilation for continuous global assessment of severe conditions over terrestrial surfaces, Hydrol. Earth Syst. Sci., 24, 4291–4316, <ext-link xlink:href="https://doi.org/10.5194/hess-24-4291-2020" ext-link-type="DOI">10.5194/hess-24-4291-2020</ext-link>, 2020.</mixed-citation></ref>
      <ref id="bib1.bibx5"><label>Bacour et al.(2019)Bacour, Maignan, MacBean, Porcar-Castell, Flexas, Frankenberg, Peylin, Chevallier, Vuichard, and Bastrikov</label><mixed-citation>Bacour, C., Maignan, F., MacBean, N., Porcar-Castell, A., Flexas, J., Frankenberg, C., Peylin, P., Chevallier, F., Vuichard, N., and Bastrikov, V.: Improving Estimates of Gross Primary Productivity by Assimilating Solar-Induced Fluorescence Satellite Retrievals in a Terrestrial Biosphere Model Using a Process-Based SIF Model, J. Geophys. Res.- Biogeo., 124, 3281–3306, <ext-link xlink:href="https://doi.org/10.1029/2019JG005040" ext-link-type="DOI">10.1029/2019JG005040</ext-link>, 2019.</mixed-citation></ref>
      <ref id="bib1.bibx6"><label>Barbu et al.(2011)Barbu, Calvet, Mahfouf, Albergel, and Lafont</label><mixed-citation>Barbu, A. L., Calvet, J.-C., Mahfouf, J.-F., Albergel, C., and Lafont, S.: Assimilation of Soil Wetness Index and Leaf Area Index into the ISBA-A-gs land surface model: grassland case study, Biogeosciences, 8, 1971–1986, <ext-link xlink:href="https://doi.org/10.5194/bg-8-1971-2011" ext-link-type="DOI">10.5194/bg-8-1971-2011</ext-link>, 2011.</mixed-citation></ref>
      <ref id="bib1.bibx7"><label>Baret et al.(2013)Baret, Weiss, Lacaze, Camacho, Makhmara, Pacholcyzk, and Smets</label><mixed-citation>Baret, F., Weiss, M., Lacaze, R., Camacho, F., Makhmara, H., Pacholcyzk, P., and Smets, B.: GEOV1: LAI and FAPAR essential climate variables and FCOVER global time series capitalizing over existing products. Part1: Principles of development and production, Remote Sens. Environ., 137, 299–309, <ext-link xlink:href="https://doi.org/10.1016/j.rse.2012.12.027" ext-link-type="DOI">10.1016/j.rse.2012.12.027</ext-link>, 2013.</mixed-citation></ref>
      <ref id="bib1.bibx8"><label>Berry(2018)</label><mixed-citation> Berry, J.: 3.10 solar induced chlorophyll fluorescence: Origins, relation to photosynthesis and retrieval, Compr. Remote Sens, 3, 143–162, 2018.</mixed-citation></ref>
      <ref id="bib1.bibx9"><label>Bolhar-Nordenkampf et al.(1989)Bolhar-Nordenkampf, Long, Baker, Oquist, Schreiber, and Lechner</label><mixed-citation>Bolhar-Nordenkampf, H. R., Long, S. P., Baker, N. R., Oquist, G., Schreiber,  U., and Lechner, E. G.: Chlorophyll Fluorescence as a Probe of the  Photosynthetic Competence of Leaves in the Field: A Review of Current  Instrumentation, Funct. Ecol., 3, 497–514, <ext-link xlink:href="https://doi.org/10.2307/2389624" ext-link-type="DOI">10.2307/2389624</ext-link>, 1989.</mixed-citation></ref>
      <ref id="bib1.bibx10"><label>Bonan et al.(2020)Bonan, Albergel, Zheng, Barbu, Fairbairn, Munier, and Calvet</label><mixed-citation>Bonan, B., Albergel, C., Zheng, Y., Barbu, A. L., Fairbairn, D., Munier, S., and Calvet, J.-C.: An ensemble square root filter for the joint assimilation of surface soil moisture and leaf area index within the Land Data Assimilation System LDAS-Monde: application over the Euro-Mediterranean region, Hydrol. Earth Syst. Sci., 24, 325–347, <ext-link xlink:href="https://doi.org/10.5194/hess-24-325-2020" ext-link-type="DOI">10.5194/hess-24-325-2020</ext-link>, 2020.</mixed-citation></ref>
      <ref id="bib1.bibx11"><label>Butterfield et al.(2023)</label><mixed-citation>Butterfield, Z., Magney, T., Grossmann, K., Bohrer, G., Vogel, C., Barr, S.,  and Keppel-Aleks, G.: Accounting for changes in radiation improves the  ability of SIF to track water stress-induced losses in summer GPP in a  temperate deciduous forest, J. Geophys. Res.-Biogeo., 128, e2022JG007352, <ext-link xlink:href="https://doi.org/10.1029/2022JG007352" ext-link-type="DOI">10.1029/2022JG007352</ext-link>, 2023.</mixed-citation></ref>
      <ref id="bib1.bibx12"><label>Calvet(2000)</label><mixed-citation> Calvet, J.-C.: Investigating soil and atmospheric plant water stress using physiological and micrometeorological data, Agr. Forest Meteorol., 103, 229–247, 2000.</mixed-citation></ref>
      <ref id="bib1.bibx13"><label>Calvet and Champeaux(2020)</label><mixed-citation>Calvet, J.-C. and Champeaux, J.-L.: L'apport de la télédétection spatiale à la modélisation des surfaces continentales, La Météorologie, 108, 52–58, <ext-link xlink:href="https://doi.org/10.37053/lameteorologie-2020-0016" ext-link-type="DOI">10.37053/lameteorologie-2020-0016</ext-link>, 2020.</mixed-citation></ref>
      <ref id="bib1.bibx14"><label>Calvet et al.(1998)Calvet, Noilhan, Roujean, Bessemoulin, Cabelguenne, Olioso, and Wigneron</label><mixed-citation> Calvet, J.-C., Noilhan, J., Roujean, J.-L., Bessemoulin, P., Cabelguenne, M., Olioso, A., and Wigneron, J.-P.: An interactive vegetation SVAT model tested against data from six contrasting sites, Agr. Forest Meteorol., 92, 73–95, 1998.</mixed-citation></ref>
      <ref id="bib1.bibx15"><label>Calvet et al.(2004)Calvet, Rivalland, Picon-Cochard, and Guehl</label><mixed-citation>Calvet, J.-C., Rivalland, V., Picon-Cochard, C., and Guehl, J.-M.: Modelling forest transpiration and CO<sub>2</sub> fluxes – Response to soil moisture stress, Agr. Forest Meteorol., 124, 143–156, 2004.</mixed-citation></ref>
      <ref id="bib1.bibx16"><label>Corchia et al.(2023)Corchia, Bonan, Rodríguez-Fernández, Colas, and Calvet</label><mixed-citation>Corchia, T., Bonan, B., Rodríguez-Fernández, N., Colas, G., and  Calvet, J.-C.: Assimilation of ASCAT Radar Backscatter Coefficients over  Southwestern France, Remote Sens., 15, 4258, <ext-link xlink:href="https://doi.org/10.3390/rs15174258" ext-link-type="DOI">10.3390/rs15174258</ext-link>, 2023.</mixed-citation></ref>
      <ref id="bib1.bibx17"><label>De Rosnay et al.(2013)De Rosnay, Drusch, Vasiljevic, Balsamo, Albergel, and Isaksen</label><mixed-citation> De Rosnay, P., Drusch, M., Vasiljevic, D., Balsamo, G., Albergel, C., and Isaksen, L.: A simplified extended Kalman filter for the global operational soil moisture analysis at ECMWF, Q. J. Roy. Meteor. Soc., 139, 1199–1213, 2013.</mixed-citation></ref>
      <ref id="bib1.bibx18"><label>Decharme et al.(2011)Decharme, Boone, Delire, and Noilhan</label><mixed-citation>Decharme, B., Boone, A., Delire, C., and Noilhan, J.: Local evaluation of the Interaction between Soil Biosphere Atmosphere soil multilayer diffusion scheme using four pedotransfer functions, J. Geophys. Res.-Atmos., 116, <ext-link xlink:href="https://doi.org/10.1029/2011JD016002" ext-link-type="DOI">10.1029/2011JD016002</ext-link>, 2011.</mixed-citation></ref>
      <ref id="bib1.bibx19"><label>Decharme et al.(2013)Decharme, Martin, and Faroux</label><mixed-citation> Decharme, B., Martin, E., and Faroux, S.: Reconciling soil thermal and hydrological lower boundary conditions in land surface models, J. Geophys. Res.-Atmos., 118, 7819–7834, 2013.</mixed-citation></ref>
      <ref id="bib1.bibx20"><label>Fairbairn et al.(2017)Fairbairn, Barbu, Napoly, Albergel, Mahfouf, and Calvet</label><mixed-citation>Fairbairn, D., Barbu, A. L., Napoly, A., Albergel, C., Mahfouf, J.-F., and Calvet, J.-C.: The effect of satellite-derived surface soil moisture and leaf area index land data assimilation on streamflow simulations over France, Hydrol. Earth Syst. Sci., 21, 2015–2033, <ext-link xlink:href="https://doi.org/10.5194/hess-21-2015-2017" ext-link-type="DOI">10.5194/hess-21-2015-2017</ext-link>, 2017.</mixed-citation></ref>
      <ref id="bib1.bibx21"><label>Faroux et al.(2013)Faroux, Kaptué Tchuenté, Roujean, Masson, Martin, and Le Moigne</label><mixed-citation>Faroux, S., Kaptué Tchuenté, A. T., Roujean, J.-L., Masson, V., Martin, E., and Le Moigne, P.: ECOCLIMAP-II/Europe: a twofold database of ecosystems and surface parameters at 1 km resolution based on satellite information for use in land surface, meteorological and climate models, Geosci. Model Dev., 6, 563–582, <ext-link xlink:href="https://doi.org/10.5194/gmd-6-563-2013" ext-link-type="DOI">10.5194/gmd-6-563-2013</ext-link>, 2013.</mixed-citation></ref>
      <ref id="bib1.bibx22"><label>Friedl and Sulla-Menashe(2015)</label><mixed-citation>Friedl, M. and Sulla-Menashe, D.: MODIS/Terra+Aqua Land Cover Type Yearly L3 Global 0.05Deg CMG. NASA LP DAAC, Tech. rep., Boston University and MODAPS SIPS, NASA, <ext-link xlink:href="https://doi.org/10.5067/MODIS/MCD12C1.006" ext-link-type="DOI">10.5067/MODIS/MCD12C1.006</ext-link>, 2015.</mixed-citation></ref>
      <ref id="bib1.bibx23"><label>Friedlingstein et al.(2025)Friedlingstein, O'Sullivan, Jones, Andrew, Hauck, Landschützer, Le Quéré, Li, Luijkx, Olsen, Peters, Peters, Pongratz, Schwingshackl, Sitch, Canadell, Ciais, Jackson, Alin, Arneth, Arora, Bates, Becker, Bellouin, Berghoff, Bittig, Bopp, Cadule, Campbell, Chamberlain, Chandra, Chevallier, Chini, Colligan, Decayeux, Djeutchouang, Dou, Duran Rojas, Enyo, Evans, Fay, Feely, Ford, Foster, Gasser, Gehlen, Gkritzalis, Grassi, Gregor, Gruber, Gürses, Harris, Hefner, Heinke, Hurtt, Iida, Ilyina, Jacobson, Jain, Jarníková, Jersild, Jiang, Jin, Kato, Keeling, Klein Goldewijk, Knauer, Korsbakken, Lan, Lauvset, Lefèvre, Liu, Liu, Ma, Maksyutov, Marland, Mayot, McGuire, Metzl, Monacci, Morgan, Nakaoka, Neill, Niwa, Nützel, Olivier, Ono, Palmer, Pierrot, Qin, Resplandy, Roobaert, Rosan, Rödenbeck, Schwinger, Smallman, Smith, Sospedra-Alfonso, Steinhoff, Sun, Sutton, Séférian, Takao, Tatebe, Tian, Tilbrook, Torres, Tourigny, Tsujino, Tubiello, van der Werf, Wanninkhof, Wang, Yang, Yang, Yu, Yuan, Yue, Zaehle, Zeng, and Zeng</label><mixed-citation>Friedlingstein, P., O'Sullivan, M., Jones, M. W., Andrew, R. M., Hauck, J., Landschützer, P., Le Quéré, C., Li, H., Luijkx, I. T., Olsen, A., Peters, G. P., Peters, W., Pongratz, J., Schwingshackl, C., Sitch, S., Canadell, J. G., Ciais, P., Jackson, R. B., Alin, S. R., Arneth, A., Arora, V., Bates, N. R., Becker, M., Bellouin, N., Berghoff, C. F., Bittig, H. C., Bopp, L., Cadule, P., Campbell, K., Chamberlain, M. A., Chandra, N., Chevallier, F., Chini, L. P., Colligan, T., Decayeux, J., Djeutchouang, L. M., Dou, X., Duran Rojas, C., Enyo, K., Evans, W., Fay, A. R., Feely, R. A., Ford, D. J., Foster, A., Gasser, T., Gehlen, M., Gkritzalis, T., Grassi, G., Gregor, L., Gruber, N., Gürses, Ö., Harris, I., Hefner, M., Heinke, J., Hurtt, G. C., Iida, Y., Ilyina, T., Jacobson, A. R., Jain, A. K., Jarníková, T., Jersild, A., Jiang, F., Jin, Z., Kato, E., Keeling, R. F., Klein Goldewijk, K., Knauer, J., Korsbakken, J. I., Lan, X., Lauvset, S. K., Lefèvre, N., Liu, Z., Liu, J., Ma, L., Maksyutov, S., Marland, G., Mayot, N., McGuire, P. C., Metzl, N., Monacci, N. M., Morgan, E. J., Nakaoka, S.-I., Neill, C., Niwa, Y., Nützel, T., Olivier, L., Ono, T., Palmer, P. I., Pierrot, D., Qin, Z., Resplandy, L., Roobaert, A., Rosan, T. M., Rödenbeck, C., Schwinger, J., Smallman, T. L., Smith, S. M., Sospedra-Alfonso, R., Steinhoff, T., Sun, Q., Sutton, A. J., Séférian, R., Takao, S., Tatebe, H., Tian, H., Tilbrook, B., Torres, O., Tourigny, E., Tsujino, H., Tubiello, F., van der Werf, G., Wanninkhof, R., Wang, X., Yang, D., Yang, X., Yu, Z., Yuan, W., Yue, X., Zaehle, S., Zeng, N., and Zeng, J.: Global Carbon Budget 2024, Earth Syst. Sci. Data, 17, 965–1039, <ext-link xlink:href="https://doi.org/10.5194/essd-17-965-2025" ext-link-type="DOI">10.5194/essd-17-965-2025</ext-link>, 2025.</mixed-citation></ref>
      <ref id="bib1.bibx24"><label>Garrigues et al.(2026)Garrigues, de Rosnay, Weston, Rüdiger, Bacour, Fairbairn, Vanderbecken, Rojas-Munoz, Pinnington, Agusti-Panareda, Boussetta, Calvet, and Engelen</label><mixed-citation>Garrigues, S., de Rosnay, P., Weston, P., Rüdiger, C., Bacour, C., Fairbairn, D., Vanderbecken, P., Rojas-Munoz, O., Pinnington, E., Agusti-Panareda, A., Boussetta, S., Calvet, J.-C., and Engelen, R.: Assimilation of solar-induced fluorescence satellite observations in the European Centre for Medium-Range Weather Forecasts integrated forecast system, Q. J. Roy. Meteor. Soc., 152, e70111, <ext-link xlink:href="https://doi.org/10.1002/qj.70111" ext-link-type="DOI">10.1002/qj.70111</ext-link>, 2026.</mixed-citation></ref>
      <ref id="bib1.bibx25"><label>Gibelin et al.(2006)</label><mixed-citation>Gibelin, A.-L., Calvet, J.-C., Roujean, J.-L., Jarlan, L., and Los, S. O.: Ability of the land surface model ISBA-A-gs to simulate leaf area index at the global scale: Comparison with satellites products, J. Geophys. Res.-Atmos., 111, <ext-link xlink:href="https://doi.org/10.1029/2005JD006691" ext-link-type="DOI">10.1029/2005JD006691</ext-link>, 2006.</mixed-citation></ref>
      <ref id="bib1.bibx26"><label>Guanter et al.(2015)Guanter, Aben, Tol, Krijger, Hollstein, Köhler, Damm, Joiner, Frankenberg, and Landgraf</label><mixed-citation>Guanter, L., Aben, I., Tol, P., Krijger, J. M., Hollstein, A., Köhler, P., Damm, A., Joiner, J., Frankenberg, C., and Landgraf, J.: Potential of the TROPOspheric Monitoring Instrument (TROPOMI) onboard the Sentinel-5 Precursor for the monitoring of terrestrial chlorophyll fluorescence, Atmos. Meas. Tech., 8, 1337–1352, <ext-link xlink:href="https://doi.org/10.5194/amt-8-1337-2015" ext-link-type="DOI">10.5194/amt-8-1337-2015</ext-link>, 2015.</mixed-citation></ref>
      <ref id="bib1.bibx27"><label>Guanter et al.(2021)Guanter, Bacour, Schneider, Aben, van Kempen, Maignan, Retscher, Köhler, Frankenberg, Joiner, and Zhang</label><mixed-citation>Guanter, L., Bacour, C., Schneider, A., Aben, I., van Kempen, T. A., Maignan, F., Retscher, C., Köhler, P., Frankenberg, C., Joiner, J., and Zhang, Y.: The TROPOSIF global sun-induced fluorescence dataset from the Sentinel-5P TROPOMI mission, Earth Syst. Sci. Data, 13, 5423–5440, <ext-link xlink:href="https://doi.org/10.5194/essd-13-5423-2021" ext-link-type="DOI">10.5194/essd-13-5423-2021</ext-link>, 2021.</mixed-citation></ref>
      <ref id="bib1.bibx28"><label>Hamer et al.(2026)Hamer, Markelj, Rojas-Munoz, Bonan, Calvet, Marécal, Guenther, Trimmel, Vallejo, Eckhardt, Sousa Santos, Sindelarova, Simpson, Schmidbauer, and Tarrasón</label><mixed-citation>Hamer, P. D., Markelj, M., Rojas-Munoz, O., Bonan, B., Calvet, J.-C., Marécal, V., Guenther, A., Trimmel, H., Vallejo, I., Eckhardt, S., Sousa Santos, G., Sindelarova, K., Simpson, D., Schmidbauer, N., Hellén, H., Rubli, P., Reimann, S., Claude, A., Kubistin, D., Cozic, J., Dernie, J., and Tarrasón, L.: Two biogenic volatile organic compound emission datasets over Europe based on land surface modelling and satellite data assimilation, Earth Syst. Sci. Data, 18, 3635–3669, <ext-link xlink:href="https://doi.org/10.5194/essd-18-3635-2026" ext-link-type="DOI">10.5194/essd-18-3635-2026</ext-link>, 2026.</mixed-citation></ref>
      <ref id="bib1.bibx29"><label>Horowitz(2016)</label><mixed-citation>Horowitz, C. A.: Paris Agreement, International Legal Materials, Cambridge University Press, 55, 740–755, <ext-link xlink:href="https://doi.org/10.1017/S0020782900004253" ext-link-type="DOI">10.1017/S0020782900004253</ext-link>, 2016.</mixed-citation></ref>
      <ref id="bib1.bibx30"><label>Jacobs(1994)</label><mixed-citation>Jacobs, C.: Direct impact of atmospheric CO2 enrichment on regional transpiration, internal phd, wU thesis 1772 Proefschrift Wageningen, Netherlands, ISBN 9789054852506, <ext-link xlink:href="https://doi.org/10.18174/206972" ext-link-type="DOI">10.18174/206972</ext-link>, 1994.</mixed-citation></ref>
      <ref id="bib1.bibx31"><label>Jacobs et al.(1996)Jacobs, Van den Hurk, and De Bruin</label><mixed-citation> Jacobs, C., Van den Hurk, B., and De Bruin, H.: Stomatal behaviour and photosynthetic rate of unstressed grapevines in semi-arid conditions, Agr. Forest Meteorol., 80, 111–134, 1996.</mixed-citation></ref>
      <ref id="bib1.bibx32"><label>Jarlan et al.(2023)</label><mixed-citation>Jarlan, L., Albergel, C., Bonan, B., Calvet, J.-C., De Rosnay, P., Ottlé, C., and Peylin, P.: Assimilation de données de télédétection pour le suivi des surfaces continentales, in: Inversion et assimilation de données de télédétection: Estimation des paramètres géophysiques, edited by: Yan, Y., ISTE, 45–95, <ext-link xlink:href="https://doi.org/10.51926/ISTE.9142.ch2" ext-link-type="DOI">10.51926/ISTE.9142.ch2</ext-link>, 2023.</mixed-citation></ref>
      <ref id="bib1.bibx33"><label>Joiner et al.(2016)Joiner, Yoshida, Guanter, and Middleton</label><mixed-citation>Joiner, J., Yoshida, Y., Guanter, L., and Middleton, E. M.: New methods for the retrieval of chlorophyll red fluorescence from hyperspectral satellite instruments: simulations and application to GOME-2 and SCIAMACHY, Atmos. Meas. Tech., 9, 3939–3967, <ext-link xlink:href="https://doi.org/10.5194/amt-9-3939-2016" ext-link-type="DOI">10.5194/amt-9-3939-2016</ext-link>, 2016.</mixed-citation></ref>
      <ref id="bib1.bibx34"><label>Jung et al.(2020)Jung, Schwalm, Migliavacca, Walther, Camps-Valls, Koirala, Anthoni, Besnard, Bodesheim, Carvalhais, Chevallier, Gans, Goll, Haverd, Köhler, Ichii, Jain, Liu, Lombardozzi, Nabel, Nelson, O'Sullivan, Pallandt, Papale, Peters, Pongratz, Rödenbeck, Sitch, Tramontana, Walker, Weber, and Reichstein</label><mixed-citation>Jung, M., Schwalm, C., Migliavacca, M., Walther, S., Camps-Valls, G., Koirala, S., Anthoni, P., Besnard, S., Bodesheim, P., Carvalhais, N., Chevallier, F., Gans, F., Goll, D. S., Haverd, V., Köhler, P., Ichii, K., Jain, A. K., Liu, J., Lombardozzi, D., Nabel, J. E. M. S., Nelson, J. A., O'Sullivan, M., Pallandt, M., Papale, D., Peters, W., Pongratz, J., Rödenbeck, C., Sitch, S., Tramontana, G., Walker, A., Weber, U., and Reichstein, M.: Scaling carbon fluxes from eddy covariance sites to globe: synthesis and evaluation of the FLUXCOM approach, Biogeosciences, 17, 1343–1365, <ext-link xlink:href="https://doi.org/10.5194/bg-17-1343-2020" ext-link-type="DOI">10.5194/bg-17-1343-2020</ext-link>, 2020.</mixed-citation></ref>
      <ref id="bib1.bibx35"><label>Köhler et al.(2015)Köhler, Guanter, and Joiner</label><mixed-citation>Köhler, P., Guanter, L., and Joiner, J.: A linear method for the retrieval of sun-induced chlorophyll fluorescence from GOME-2 and SCIAMACHY data, Atmos. Meas. Tech., 8, 2589–2608, <ext-link xlink:href="https://doi.org/10.5194/amt-8-2589-2015" ext-link-type="DOI">10.5194/amt-8-2589-2015</ext-link>, 2015.</mixed-citation></ref>
      <ref id="bib1.bibx36"><label>Köhler et al.(2018)Köhler, Frankenberg, Magney, Guanter, Joiner, and Landgraf</label><mixed-citation>Köhler, P., Frankenberg, C., Magney, T. S., Guanter, L., Joiner, J., and Landgraf, J.: Global Retrievals of Solar-Induced Chlorophyll Fluorescence With TROPOMI: First Results and Intersensor Comparison to OCO-2, Geophys. Res. Lett., 45, 10456–10463, <ext-link xlink:href="https://doi.org/10.1029/2018GL079031" ext-link-type="DOI">10.1029/2018GL079031</ext-link>, 2018.</mixed-citation></ref>
      <ref id="bib1.bibx37"><label>Leroux et al.(2018)Leroux, Calvet, Munier, and Albergel</label><mixed-citation>Leroux, D. J., Calvet, J.-C., Munier, S., and Albergel, C.: Using Satellite-Derived Vegetation Products to Evaluate LDAS-Monde over the Euro-Mediterranean Area, Remote Sens., 10, <ext-link xlink:href="https://doi.org/10.3390/rs10081199" ext-link-type="DOI">10.3390/rs10081199</ext-link>, 2018.</mixed-citation></ref>
      <ref id="bib1.bibx38"><label>LOBELIA(2024)</label><mixed-citation>LOBELIA: Revealing emission sources from space, Zenodo, <ext-link xlink:href="https://doi.org/10.5281/zenodo.10837172" ext-link-type="DOI">10.5281/zenodo.10837172</ext-link>, 2024.</mixed-citation></ref>
      <ref id="bib1.bibx39"><label>Lunel et al.(2024)Lunel, Boone, and Le Moigne</label><mixed-citation>Lunel, T., Boone, A. A., and Le Moigne, P.: Irrigation strongly influences  near-surface conditions and induces breeze circulation: Observational and  model-based evidence, Q. J. Roy. Meteor. Soc., 150, <ext-link xlink:href="https://doi.org/10.1002/qj.4736" ext-link-type="DOI">10.1002/qj.4736</ext-link>, 2024.</mixed-citation></ref>
      <ref id="bib1.bibx40"><label>Mahfouf and Bilodeau(2009)</label><mixed-citation>Mahfouf, J.-F. and Bilodeau, B.: A simple strategy for linearizing complex moist convective schemes, Q. J. Roy. Meteor. Soc., 135, 953–962, <ext-link xlink:href="https://doi.org/10.1002/qj.427" ext-link-type="DOI">10.1002/qj.427</ext-link>, 2009.</mixed-citation></ref>
      <ref id="bib1.bibx41"><label>Mahfouf et al.(2009)Mahfouf, Bergaoui, Draper, Bouyssel, Taillefer, and Taseva</label><mixed-citation>Mahfouf, J.-F., Bergaoui, K., Draper, C., Bouyssel, F., Taillefer, F., and Taseva, L.: A comparison of two off-line soil analysis schemes for assimilation of screen level observations, J. Geophys. Res.-Atmos., 114, <ext-link xlink:href="https://doi.org/10.1029/2008JD011077" ext-link-type="DOI">10.1029/2008JD011077</ext-link>, 2009.</mixed-citation></ref>
      <ref id="bib1.bibx42"><label>Masson et al.(2013)Masson, Le Moigne, Martin, Faroux, Alias, Alkama, Belamari, Barbu, Boone, Bouyssel et al.</label><mixed-citation>Masson, V., Le Moigne, P., Martin, E., Faroux, S., Alias, A., Alkama, R., Belamari, S., Barbu, A., Boone, A., Bouyssel, F., Brousseau, P., Brun, E., Calvet, J.-C., Carrer, D., Decharme, B., Delire, C., Donier, S., Essaouini, K., Gibelin, A.-L., Giordani, H., Habets, F., Jidane, M., Kerdraon, G., Kourzeneva, E., Lafaysse, M., Lafont, S., Lebeaupin Brossier, C., Lemonsu, A., Mahfouf, J.-F., Marguinaud, P., Mokhtari, M., Morin, S., Pigeon, G., Salgado, R., Seity, Y., Taillefer, F., Tanguy, G., Tulet, P., Vincendon, B., Vionnet, V., and Voldoire, A.: The SURFEXv7.2 land and ocean surface platform for coupled or offline simulation of earth surface variables and fluxes, Geosci. Model Dev., 6, 929–960, <ext-link xlink:href="https://doi.org/10.5194/gmd-6-929-2013" ext-link-type="DOI">10.5194/gmd-6-929-2013</ext-link>, 2013.</mixed-citation></ref>
      <ref id="bib1.bibx43"><label>Moya et al.(2004)Moya, Camenen, Evain, Goulas, Cerovic, Latouche, Flexas, and Ounis</label><mixed-citation>Moya, I., Camenen, L., Evain, S., Goulas, Y., Cerovic, Z., Latouche, G., Flexas, J., and Ounis, A.: A new instrument for passive remote sensing: 1. Measurements of sunlight-induced chlorophyll fluorescence, Remote Sens. Environ., 91, 186–197, <ext-link xlink:href="https://doi.org/10.1016/j.rse.2004.02.012" ext-link-type="DOI">10.1016/j.rse.2004.02.012</ext-link>, 2004.</mixed-citation></ref>
      <ref id="bib1.bibx44"><label>Nelson et al.(2024)Nelson, Walther, Gans, Kraft, Weber, Novick, Buchmann, Migliavacca, Wohlfahrt, Šigut, Ibrom, Papale, Göckede, Duveiller, Knohl, Hörtnagl, Scott, Dušek, Zhang, Hamdi, Reichstein, Aranda-Barranco, Ardö, Op de Beeck, Billesbach, Bowling, Bracho, Brümmer, Camps-Valls, Chen, Cleverly, Desai, Dong, El-Madany, Euskirchen, Feigenwinter, Galvagno, Gerosa, Gielen, Goded, Goslee, Gough, Heinesch, Ichii, Jackowicz-Korczynski, Klosterhalfen, Knox, Kobayashi, Kohonen, Korkiakoski, Mammarella, Gharun, Marzuoli, Matamala, Metzger, Montagnani, Nicolini, O'Halloran, Ourcival, Peichl, Pendall, Ruiz Reverter, Roland, Sabbatini, Sachs, Schmidt, Schwalm, Shekhar, Silberstein, Silveira, Spano, Tagesson, Tramontana, Trotta, Turco, Vesala, Vincke, Vitale, Vivoni, Wang, Woodgate, Yepez, Zhang, Zona, and Jung</label><mixed-citation>Nelson, J. A., Walther, S., Gans, F., Kraft, B., Weber, U., Novick, K., Buchmann, N., Migliavacca, M., Wohlfahrt, G., Šigut, L., Ibrom, A., Papale, D., Göckede, M., Duveiller, G., Knohl, A., Hörtnagl, L., Scott, R. L., Dušek, J., Zhang, W., Hamdi, Z. M., Reichstein, M., Aranda-Barranco, S., Ardö, J., Op de Beeck, M., Billesbach, D., Bowling, D., Bracho, R., Brümmer, C., Camps-Valls, G., Chen, S., Cleverly, J. R., Desai, A., Dong, G., El-Madany, T. S., Euskirchen, E. S., Feigenwinter, I., Galvagno, M., Gerosa, G. A., Gielen, B., Goded, I., Goslee, S., Gough, C. M., Heinesch, B., Ichii, K., Jackowicz-Korczynski, M. A., Klosterhalfen, A., Knox, S., Kobayashi, H., Kohonen, K.-M., Korkiakoski, M., Mammarella, I., Gharun, M., Marzuoli, R., Matamala, R., Metzger, S., Montagnani, L., Nicolini, G., O'Halloran, T., Ourcival, J.-M., Peichl, M., Pendall, E., Ruiz Reverter, B., Roland, M., Sabbatini, S., Sachs, T., Schmidt, M., Schwalm, C. R., Shekhar, A., Silberstein, R., Silveira, M. L., Spano, D., Tagesson, T., Tramontana, G., Trotta, C., Turco, F., Vesala, T., Vincke, C., Vitale, D., Vivoni, E. R., Wang, Y., Woodgate, W., Yepez, E. A., Zhang, J., Zona, D., and Jung, M.: X-BASE: the first terrestrial carbon and water flux products from an extended data-driven scaling framework, FLUXCOM-X, Biogeosciences, 21, 5079–5115, <ext-link xlink:href="https://doi.org/10.5194/bg-21-5079-2024" ext-link-type="DOI">10.5194/bg-21-5079-2024</ext-link>, 2024.</mixed-citation></ref>
      <ref id="bib1.bibx45"><label>Noilhan and Mahfouf(1996)</label><mixed-citation>Noilhan, J. and Mahfouf, J.-F.: The ISBA land surface parameterisation scheme, Global Planet. Change, 13, 145–159, <ext-link xlink:href="https://doi.org/10.1016/0921-8181(95)00043-7" ext-link-type="DOI">10.1016/0921-8181(95)00043-7</ext-link>, 1996.</mixed-citation></ref>
      <ref id="bib1.bibx46"><label>Noilhan and Planton(1989)</label><mixed-citation>Noilhan, J. and Planton, S.: A Simple Parameterization of Land Surface Processes for Meteorological Models, Mon. Weather Rev., 117, 536–549, <ext-link xlink:href="https://doi.org/10.1175/1520-0493(1989)117&lt;0536:ASPOLS&gt;2.0.CO;2" ext-link-type="DOI">10.1175/1520-0493(1989)117&lt;0536:ASPOLS&gt;2.0.CO;2</ext-link>, 1989.</mixed-citation></ref>
      <ref id="bib1.bibx47"><label>Rascher et al.(2015)</label><mixed-citation>Rascher, U., Alonso, L., Burkart, A., Cilia, C., Cogliati, S., Colombo, R., Damm, A., Drusch, M., Guanter, L., Hanus, J., Hyvärinen, T., Julitta, T., Jussila, J., Kataja, K., Kokkalis, P., Kraft, S., Kraska, T., Matveeva, M., Moreno, J., Muller, O., Panigada, C., Pikl, M., Pinto, F., Prey, L., Pude, R., Rossini, M., Schickling, A., Schurr, U., Schüttemeyer, D., Verrelst, J., and Zemek, F.: Sun-induced fluorescence – a new probe of photosynthesis: First maps from the imaging spectrometer HyPlant, Glob. Change Biol., 21, 4673–4684, <ext-link xlink:href="https://doi.org/10.1111/gcb.13017" ext-link-type="DOI">10.1111/gcb.13017</ext-link>, 2015. </mixed-citation></ref>
      <ref id="bib1.bibx48"><label>Ricart et al.(2016)Ricart, Ribas, and Pavón</label><mixed-citation>Ricart, S., Ribas, A., and Pavón, D.: Qualifying irrigation system sustainability by means of stakeholder perceptions and concerns: lessons from the Segarra-Garrigues Canal, Spain, Nat. Resour. Forum, 40, 77–90, <ext-link xlink:href="https://doi.org/10.1111/1477-8947.12097" ext-link-type="DOI">10.1111/1477-8947.12097</ext-link>, 2016.</mixed-citation></ref>
      <ref id="bib1.bibx49"><label>Richards(1931)</label><mixed-citation> Richards, L. A.: Capillary conduction of liquids through porous mediums, Physics, 1, 318–333, 1931.</mixed-citation></ref>
      <ref id="bib1.bibx50"><label>Rüdiger et al.(2010)Rüdiger, Albergel, Mahfouf, Calvet, and Walker</label><mixed-citation>Rüdiger, C., Albergel, C., Mahfouf, J.-F., Calvet, J.-C., and Walker, J. P.: Evaluation of the observation operator Jacobian for leaf area index data assimilation with an extended Kalman filter, J. Geophys. Res.-Atmos., 115, <ext-link xlink:href="https://doi.org/10.1029/2009JD012912" ext-link-type="DOI">10.1029/2009JD012912</ext-link>, 2010.</mixed-citation></ref>
      <ref id="bib1.bibx51"><label>Sabater et al.(2008)Sabater, Rüdiger, Calvet, Fritz, Jarlan, and Kerr</label><mixed-citation>Sabater, J. M., Rüdiger, C., Calvet, J.-C., Fritz, N., Jarlan, L., and Kerr, Y.: Joint assimilation of surface soil moisture and LAI observations into a land surface model, Agr. Forest Meteorol., 148, 1362–1373, <ext-link xlink:href="https://doi.org/10.1016/j.agrformet.2008.04.003" ext-link-type="DOI">10.1016/j.agrformet.2008.04.003</ext-link>, 2008.</mixed-citation></ref>
      <ref id="bib1.bibx52"><label>Trimmel et al.(2023)Trimmel, Hamer, Mayer, Schreier, Weihs, Eitzinger, Sandén, Fitzky, Richter, Calvet, Bonan, Meurey, Vallejo, Eckhardt, Santos, Oumami, Arteta, Marécal, Tarrasón, Karl, and Rieder</label><mixed-citation>Trimmel, H., Hamer, P., Mayer, M., Schreier, S. F., Weihs, P., Eitzinger, J., Sandén, H., Fitzky, A. C., Richter, A., Calvet, J.-C., Bonan, B., Meurey, C., Vallejo, I., Eckhardt, S., Santos, G. S., Oumami, S., Arteta, J., Marécal, V., Tarrasón, L., Karl, T., and Rieder, H. E.: The influence of vegetation drought stress on formaldehyde and ozone distributions over a central European city, Atmos. Environ., 304, 119768, <ext-link xlink:href="https://doi.org/10.1016/j.atmosenv.2023.119768" ext-link-type="DOI">10.1016/j.atmosenv.2023.119768</ext-link>, 2023.</mixed-citation></ref>
      <ref id="bib1.bibx53"><label>Vanderbecken(2026a)Vanderbecken</label><mixed-citation>Vanderbecken, P. J.: WP4 – supplementary data – Using deep learning to assimilate sun-induced fluorescence satellite observations in the ISBA land surface model: Datasets, Zenodo [data set], <ext-link xlink:href="https://doi.org/10.5281/zenodo.18668100" ext-link-type="DOI">10.5281/zenodo.18668100</ext-link>, 2026a.</mixed-citation></ref>
      <ref id="bib1.bibx54"><label>Vanderbecken(2026b)Vanderbecken</label><mixed-citation>Vanderbecken, P. J.: WP4 – supplementary data – Using deep learning to assimilate sun-induced fluorescence satellite observations in the ISBA land surface model: model, Zenodo [code], <ext-link xlink:href="https://doi.org/10.5281/zenodo.18669437" ext-link-type="DOI">10.5281/zenodo.18669437</ext-link>, 2026b.</mixed-citation></ref>
      <ref id="bib1.bibx55"><label>Veefkind et al.(2012)Veefkind, Aben, McMullan, Förster, de Vries, Otter, Claas, Eskes, de Haan, Kleipool, van Weele, Hasekamp, Hoogeveen, Landgraf, Snel, Tol, Ingmann, Voors, Kruizinga, Vink, Visser, and Levelt</label><mixed-citation>Veefkind, J., Aben, I., McMullan, K., Förster, H., de Vries, J., Otter, G., Claas, J., Eskes, H., de Haan, J., Kleipool, Q., van Weele, M., Hasekamp, O., Hoogeveen, R., Landgraf, J., Snel, R., Tol, P., Ingmann, P., Voors, R., Kruizinga, B., Vink, R., Visser, H., and Levelt, P.: TROPOMI on the ESA Sentinel-5 Precursor: A GMES mission for global observations of the atmospheric composition for climate, air quality and ozone layer applications, Remote Sens. Environ., 120, 70–83, <ext-link xlink:href="https://doi.org/10.1016/j.rse.2011.09.027" ext-link-type="DOI">10.1016/j.rse.2011.09.027</ext-link>, 2012.</mixed-citation></ref>

  </ref-list></back>
    <!--<article-title-html>Using deep learning to assimilate sun-induced fluorescence satellite observations in the ISBA land surface model</article-title-html>
<abstract-html/>
<ref-html id="bib1.bib1"><label>Agustí-Panareda et al.(2022)Agustí-Panareda, McNorton,
Balsamo, Baier, Bousserez, Boussetta, Brunner, Chevallier, Choulga,
Diamantakis et al.</label><mixed-citation>
      
Agustí-Panareda, A., McNorton, J., Balsamo, G., Baier, B. C., Bousserez,  N., Boussetta, S., Brunner, D., Chevallier, F., Choulga, M., Diamantakis, M., Engelen, R., Flemming, J., Granier, C., Guevara, M., Denier van der Gon, H., Elguindi, N., Haussaire, J.-M., Jung, M., Janssens-Maenhout, G., Kivi, R., Massart, S., Papale, D., Parrington, M., Razinger, M., Sweeney, C., Vermeulen, A., and Walther, S.: Global nature run data with realistic high-resolution carbon weather for the year of the Paris agreement, Scientific Data, 9, 160, <a href="https://doi.org/10.1038/s41597-022-01228-2" target="_blank">https://doi.org/10.1038/s41597-022-01228-2</a>, 2022.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib2"><label>Albergel et al.(2010)Albergel, Calvet, Mahfouf, Rüdiger, Barbu,
Lafont, Roujean, Walker, Crapeau, and Wigneron</label><mixed-citation>
      
Albergel, C., Calvet, J.-C., Mahfouf, J.-F., Rüdiger, C., Barbu, A. L., Lafont, S., Roujean, J.-L., Walker, J. P., Crapeau, M., and Wigneron, J.-P.: Monitoring of water and carbon fluxes using a land data assimilation system: a case study for southwestern France, Hydrol. Earth Syst. Sci., 14, 1109–1124, <a href="https://doi.org/10.5194/hess-14-1109-2010" target="_blank">https://doi.org/10.5194/hess-14-1109-2010</a>, 2010.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib3"><label>Albergel et al.(2017)Albergel, Munier, Leroux, Dewaele, Fairbairn,
Barbu, Gelati, Dorigo, Faroux, Meurey, Le Moigne, Decharme, Mahfouf, and
Calvet</label><mixed-citation>
      
Albergel, C., Munier, S., Leroux, D. J., Dewaele, H., Fairbairn, D., Barbu, A. L., Gelati, E., Dorigo, W., Faroux, S., Meurey, C., Le Moigne, P., Decharme, B., Mahfouf, J.-F., and Calvet, J.-C.: Sequential assimilation of satellite-derived vegetation and soil moisture products using SURFEX_v8.0: LDAS-Monde assessment over the Euro-Mediterranean area, Geosci. Model Dev., 10, 3889–3912, <a href="https://doi.org/10.5194/gmd-10-3889-2017" target="_blank">https://doi.org/10.5194/gmd-10-3889-2017</a>, 2017.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib4"><label>Albergel et al.(2020)Albergel, Zheng, Bonan, Dutra,
Rodríguez-Fernández, Munier, Draper, de Rosnay, Muñoz Sabater,
Balsamo, Fairbairn, Meurey, and Calvet</label><mixed-citation>
      
Albergel, C., Zheng, Y., Bonan, B., Dutra, E., Rodríguez-Fernández, N., Munier, S., Draper, C., de Rosnay, P., Muñoz-Sabater, J., Balsamo, G., Fairbairn, D., Meurey, C., and Calvet, J.-C.: Data assimilation for continuous global assessment of severe conditions over terrestrial surfaces, Hydrol. Earth Syst. Sci., 24, 4291–4316, <a href="https://doi.org/10.5194/hess-24-4291-2020" target="_blank">https://doi.org/10.5194/hess-24-4291-2020</a>, 2020.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib5"><label>Bacour et al.(2019)Bacour, Maignan, MacBean, Porcar-Castell, Flexas,
Frankenberg, Peylin, Chevallier, Vuichard, and Bastrikov</label><mixed-citation>
      
Bacour, C., Maignan, F., MacBean, N., Porcar-Castell, A., Flexas, J.,
Frankenberg, C., Peylin, P., Chevallier, F., Vuichard, N., and Bastrikov, V.:
Improving Estimates of Gross Primary Productivity by Assimilating
Solar-Induced Fluorescence Satellite Retrievals in a Terrestrial Biosphere
Model Using a Process-Based SIF Model, J. Geophys. Res.- Biogeo., 124, 3281–3306, <a href="https://doi.org/10.1029/2019JG005040" target="_blank">https://doi.org/10.1029/2019JG005040</a>,
2019.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib6"><label>Barbu et al.(2011)Barbu, Calvet, Mahfouf, Albergel, and
Lafont</label><mixed-citation>
      
Barbu, A. L., Calvet, J.-C., Mahfouf, J.-F., Albergel, C., and Lafont, S.: Assimilation of Soil Wetness Index and Leaf Area Index into the ISBA-A-gs land surface model: grassland case study, Biogeosciences, 8, 1971–1986, <a href="https://doi.org/10.5194/bg-8-1971-2011" target="_blank">https://doi.org/10.5194/bg-8-1971-2011</a>, 2011.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib7"><label>Baret et al.(2013)Baret, Weiss, Lacaze, Camacho, Makhmara,
Pacholcyzk, and Smets</label><mixed-citation>
      
Baret, F., Weiss, M., Lacaze, R., Camacho, F., Makhmara, H., Pacholcyzk, P.,
and Smets, B.: GEOV1: LAI and FAPAR essential climate variables and FCOVER
global time series capitalizing over existing products. Part1: Principles of
development and production, Remote Sens. Environ., 137, 299–309,
<a href="https://doi.org/10.1016/j.rse.2012.12.027" target="_blank">https://doi.org/10.1016/j.rse.2012.12.027</a>, 2013.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib8"><label>Berry(2018)</label><mixed-citation>
      
Berry, J.: 3.10 solar induced chlorophyll fluorescence: Origins, relation to
photosynthesis and retrieval, Compr. Remote Sens, 3, 143–162, 2018.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib9"><label>Bolhar-Nordenkampf et al.(1989)Bolhar-Nordenkampf, Long, Baker,
Oquist, Schreiber, and Lechner</label><mixed-citation>
      
Bolhar-Nordenkampf, H. R., Long, S. P., Baker, N. R., Oquist, G., Schreiber,  U., and Lechner, E. G.: Chlorophyll Fluorescence as a Probe of the  Photosynthetic Competence of Leaves in the Field: A Review of Current  Instrumentation, Funct. Ecol., 3, 497–514, <a href="https://doi.org/10.2307/2389624" target="_blank">https://doi.org/10.2307/2389624</a>, 1989.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib10"><label>Bonan et al.(2020)Bonan, Albergel, Zheng, Barbu, Fairbairn, Munier,
and Calvet</label><mixed-citation>
      
Bonan, B., Albergel, C., Zheng, Y., Barbu, A. L., Fairbairn, D., Munier, S., and Calvet, J.-C.: An ensemble square root filter for the joint assimilation of surface soil moisture and leaf area index within the Land Data Assimilation System LDAS-Monde: application over the Euro-Mediterranean region, Hydrol. Earth Syst. Sci., 24, 325–347, <a href="https://doi.org/10.5194/hess-24-325-2020" target="_blank">https://doi.org/10.5194/hess-24-325-2020</a>, 2020.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib11"><label>Butterfield et al.(2023)</label><mixed-citation>
      
Butterfield, Z., Magney, T., Grossmann, K., Bohrer, G., Vogel, C., Barr, S.,  and Keppel-Aleks, G.: Accounting for changes in radiation improves the  ability of SIF to track water stress-induced losses in summer GPP in a  temperate deciduous forest, J. Geophys. Res.-Biogeo., 128, e2022JG007352, <a href="https://doi.org/10.1029/2022JG007352" target="_blank">https://doi.org/10.1029/2022JG007352</a>, 2023.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib12"><label>Calvet(2000)</label><mixed-citation>
      
Calvet, J.-C.: Investigating soil and atmospheric plant water stress using
physiological and micrometeorological data, Agr. Forest
Meteorol., 103, 229–247, 2000.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib13"><label>Calvet and Champeaux(2020)</label><mixed-citation>
      
Calvet, J.-C. and Champeaux, J.-L.: L'apport de la télédétection spatiale à
la modélisation des surfaces continentales, La Météorologie, 108, 52–58,
<a href="https://doi.org/10.37053/lameteorologie-2020-0016" target="_blank">https://doi.org/10.37053/lameteorologie-2020-0016</a>, 2020.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib14"><label>Calvet et al.(1998)Calvet, Noilhan, Roujean, Bessemoulin,
Cabelguenne, Olioso, and Wigneron</label><mixed-citation>
      
Calvet, J.-C., Noilhan, J., Roujean, J.-L., Bessemoulin, P., Cabelguenne, M.,
Olioso, A., and Wigneron, J.-P.: An interactive vegetation SVAT model tested
against data from six contrasting sites, Agr. Forest Meteorol.,
92, 73–95, 1998.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib15"><label>Calvet et al.(2004)Calvet, Rivalland, Picon-Cochard, and
Guehl</label><mixed-citation>
      
Calvet, J.-C., Rivalland, V., Picon-Cochard, C., and Guehl, J.-M.: Modelling
forest transpiration and CO<sub>2</sub> fluxes – Response to soil moisture stress,
Agr. Forest Meteorol., 124, 143–156, 2004.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib16"><label>Corchia et al.(2023)Corchia, Bonan, Rodríguez-Fernández,
Colas, and Calvet</label><mixed-citation>
      
Corchia, T., Bonan, B., Rodríguez-Fernández, N., Colas, G., and  Calvet, J.-C.: Assimilation of ASCAT Radar Backscatter Coefficients over  Southwestern France, Remote Sens., 15, 4258, <a href="https://doi.org/10.3390/rs15174258" target="_blank">https://doi.org/10.3390/rs15174258</a>, 2023.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib17"><label>De Rosnay et al.(2013)De Rosnay, Drusch, Vasiljevic, Balsamo,
Albergel, and Isaksen</label><mixed-citation>
      
De Rosnay, P., Drusch, M., Vasiljevic, D., Balsamo, G., Albergel, C., and
Isaksen, L.: A simplified extended Kalman filter for the global operational
soil moisture analysis at ECMWF, Q. J. Roy.
Meteor. Soc., 139, 1199–1213, 2013.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib18"><label>Decharme et al.(2011)Decharme, Boone, Delire, and
Noilhan</label><mixed-citation>
      
Decharme, B., Boone, A., Delire, C., and Noilhan, J.: Local evaluation of the
Interaction between Soil Biosphere Atmosphere soil multilayer diffusion
scheme using four pedotransfer functions, J. Geophys. Res.-Atmos., 116, <a href="https://doi.org/10.1029/2011JD016002" target="_blank">https://doi.org/10.1029/2011JD016002</a>, 2011.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib19"><label>Decharme et al.(2013)Decharme, Martin, and
Faroux</label><mixed-citation>
      
Decharme, B., Martin, E., and Faroux, S.: Reconciling soil thermal and
hydrological lower boundary conditions in land surface models, J. Geophys. Res.-Atmos., 118, 7819–7834, 2013.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib20"><label>Fairbairn et al.(2017)Fairbairn, Barbu, Napoly, Albergel, Mahfouf,
and Calvet</label><mixed-citation>
      
Fairbairn, D., Barbu, A. L., Napoly, A., Albergel, C., Mahfouf, J.-F., and Calvet, J.-C.: The effect of satellite-derived surface soil moisture and leaf area index land data assimilation on streamflow simulations over France, Hydrol. Earth Syst. Sci., 21, 2015–2033, <a href="https://doi.org/10.5194/hess-21-2015-2017" target="_blank">https://doi.org/10.5194/hess-21-2015-2017</a>, 2017.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib21"><label>Faroux et al.(2013)Faroux, Kaptué Tchuenté, Roujean, Masson,
Martin, and Le Moigne</label><mixed-citation>
      
Faroux, S., Kaptué Tchuenté, A. T., Roujean, J.-L., Masson, V., Martin, E., and Le Moigne, P.: ECOCLIMAP-II/Europe: a twofold database of ecosystems and surface parameters at 1&thinsp;km resolution based on satellite information for use in land surface, meteorological and climate models, Geosci. Model Dev., 6, 563–582, <a href="https://doi.org/10.5194/gmd-6-563-2013" target="_blank">https://doi.org/10.5194/gmd-6-563-2013</a>, 2013.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib22"><label>Friedl and Sulla-Menashe(2015)</label><mixed-citation>
      
Friedl, M. and Sulla-Menashe, D.: MODIS/Terra+Aqua Land Cover Type Yearly L3
Global 0.05Deg CMG. NASA LP DAAC, Tech. rep., Boston University and MODAPS
SIPS, NASA, <a href="https://doi.org/10.5067/MODIS/MCD12C1.006" target="_blank">https://doi.org/10.5067/MODIS/MCD12C1.006</a>, 2015.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib23"><label>Friedlingstein et al.(2025)Friedlingstein, O'Sullivan, Jones, Andrew,
Hauck, Landschützer, Le Quéré, Li, Luijkx, Olsen, Peters, Peters,
Pongratz, Schwingshackl, Sitch, Canadell, Ciais, Jackson, Alin, Arneth,
Arora, Bates, Becker, Bellouin, Berghoff, Bittig, Bopp, Cadule, Campbell,
Chamberlain, Chandra, Chevallier, Chini, Colligan, Decayeux, Djeutchouang,
Dou, Duran Rojas, Enyo, Evans, Fay, Feely, Ford, Foster, Gasser, Gehlen,
Gkritzalis, Grassi, Gregor, Gruber, Gürses, Harris, Hefner, Heinke, Hurtt,
Iida, Ilyina, Jacobson, Jain, Jarníková, Jersild, Jiang, Jin, Kato,
Keeling, Klein Goldewijk, Knauer, Korsbakken, Lan, Lauvset, Lefèvre, Liu,
Liu, Ma, Maksyutov, Marland, Mayot, McGuire, Metzl, Monacci, Morgan, Nakaoka,
Neill, Niwa, Nützel, Olivier, Ono, Palmer, Pierrot, Qin, Resplandy,
Roobaert, Rosan, Rödenbeck, Schwinger, Smallman, Smith, Sospedra-Alfonso,
Steinhoff, Sun, Sutton, Séférian, Takao, Tatebe, Tian, Tilbrook, Torres,
Tourigny, Tsujino, Tubiello, van der Werf, Wanninkhof, Wang, Yang, Yang, Yu,
Yuan, Yue, Zaehle, Zeng, and Zeng</label><mixed-citation>
      
Friedlingstein, P., O'Sullivan, M., Jones, M. W., Andrew, R. M., Hauck, J., Landschützer, P., Le Quéré, C., Li, H., Luijkx, I. T., Olsen, A., Peters, G. P., Peters, W., Pongratz, J., Schwingshackl, C., Sitch, S., Canadell, J. G., Ciais, P., Jackson, R. B., Alin, S. R., Arneth, A., Arora, V., Bates, N. R., Becker, M., Bellouin, N., Berghoff, C. F., Bittig, H. C., Bopp, L., Cadule, P., Campbell, K., Chamberlain, M. A., Chandra, N., Chevallier, F., Chini, L. P., Colligan, T., Decayeux, J., Djeutchouang, L. M., Dou, X., Duran Rojas, C., Enyo, K., Evans, W., Fay, A. R., Feely, R. A., Ford, D. J., Foster, A., Gasser, T., Gehlen, M., Gkritzalis, T., Grassi, G., Gregor, L., Gruber, N., Gürses, Ö., Harris, I., Hefner, M., Heinke, J., Hurtt, G. C., Iida, Y., Ilyina, T., Jacobson, A. R., Jain, A. K., Jarníková, T., Jersild, A., Jiang, F., Jin, Z., Kato, E., Keeling, R. F., Klein Goldewijk, K., Knauer, J., Korsbakken, J. I., Lan, X., Lauvset, S. K., Lefèvre, N., Liu, Z., Liu, J., Ma, L., Maksyutov, S., Marland, G., Mayot, N., McGuire, P. C., Metzl, N., Monacci, N. M., Morgan, E. J., Nakaoka, S.-I., Neill, C., Niwa, Y., Nützel, T., Olivier, L., Ono, T., Palmer, P. I., Pierrot, D., Qin, Z., Resplandy, L., Roobaert, A., Rosan, T. M., Rödenbeck, C., Schwinger, J., Smallman, T. L., Smith, S. M., Sospedra-Alfonso, R., Steinhoff, T., Sun, Q., Sutton, A. J., Séférian, R., Takao, S., Tatebe, H., Tian, H., Tilbrook, B., Torres, O., Tourigny, E., Tsujino, H., Tubiello, F., van der Werf, G., Wanninkhof, R., Wang, X., Yang, D., Yang, X., Yu, Z., Yuan, W., Yue, X., Zaehle, S., Zeng, N., and Zeng, J.: Global Carbon Budget 2024, Earth Syst. Sci. Data, 17, 965–1039, <a href="https://doi.org/10.5194/essd-17-965-2025" target="_blank">https://doi.org/10.5194/essd-17-965-2025</a>, 2025.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib24"><label>Garrigues et al.(2026)Garrigues, de Rosnay, Weston, Rüdiger, Bacour,
Fairbairn, Vanderbecken, Rojas-Munoz, Pinnington, Agusti-Panareda, Boussetta,
Calvet, and Engelen</label><mixed-citation>
      
Garrigues, S., de Rosnay, P., Weston, P., Rüdiger, C., Bacour, C., Fairbairn,
D., Vanderbecken, P., Rojas-Munoz, O., Pinnington, E., Agusti-Panareda, A.,
Boussetta, S., Calvet, J.-C., and Engelen, R.: Assimilation of solar-induced
fluorescence satellite observations in the European Centre for Medium-Range
Weather Forecasts integrated forecast system, Q. J. Roy.
Meteor. Soc., 152, e70111,
<a href="https://doi.org/10.1002/qj.70111" target="_blank">https://doi.org/10.1002/qj.70111</a>, 2026.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib25"><label>Gibelin et al.(2006)</label><mixed-citation>
      
Gibelin, A.-L., Calvet, J.-C., Roujean, J.-L., Jarlan, L., and Los, S. O.:
Ability of the land surface model ISBA-A-gs to simulate leaf area index at
the global scale: Comparison with satellites products, J. Geophys.
Res.-Atmos., 111, <a href="https://doi.org/10.1029/2005JD006691" target="_blank">https://doi.org/10.1029/2005JD006691</a>, 2006.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib26"><label>Guanter et al.(2015)Guanter, Aben, Tol, Krijger, Hollstein, Köhler,
Damm, Joiner, Frankenberg, and Landgraf</label><mixed-citation>
      
Guanter, L., Aben, I., Tol, P., Krijger, J. M., Hollstein, A., Köhler, P., Damm, A., Joiner, J., Frankenberg, C., and Landgraf, J.: Potential of the TROPOspheric Monitoring Instrument (TROPOMI) onboard the Sentinel-5 Precursor for the monitoring of terrestrial chlorophyll fluorescence, Atmos. Meas. Tech., 8, 1337–1352, <a href="https://doi.org/10.5194/amt-8-1337-2015" target="_blank">https://doi.org/10.5194/amt-8-1337-2015</a>, 2015.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib27"><label>Guanter et al.(2021)Guanter, Bacour, Schneider, Aben, van Kempen,
Maignan, Retscher, Köhler, Frankenberg, Joiner, and Zhang</label><mixed-citation>
      
Guanter, L., Bacour, C., Schneider, A., Aben, I., van Kempen, T. A., Maignan, F., Retscher, C., Köhler, P., Frankenberg, C., Joiner, J., and Zhang, Y.: The TROPOSIF global sun-induced fluorescence dataset from the Sentinel-5P TROPOMI mission, Earth Syst. Sci. Data, 13, 5423–5440, <a href="https://doi.org/10.5194/essd-13-5423-2021" target="_blank">https://doi.org/10.5194/essd-13-5423-2021</a>, 2021.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib28"><label>Hamer et al.(2026)Hamer, Markelj, Rojas-Munoz, Bonan, Calvet,
Marécal, Guenther, Trimmel, Vallejo, Eckhardt, Sousa Santos, Sindelarova,
Simpson, Schmidbauer, and Tarrasón</label><mixed-citation>
      
Hamer, P. D., Markelj, M., Rojas-Munoz, O., Bonan, B., Calvet, J.-C., Marécal, V., Guenther, A., Trimmel, H., Vallejo, I., Eckhardt, S., Sousa Santos, G., Sindelarova, K., Simpson, D., Schmidbauer, N., Hellén, H., Rubli, P., Reimann, S., Claude, A., Kubistin, D., Cozic, J., Dernie, J., and Tarrasón, L.: Two biogenic volatile organic compound emission datasets over Europe based on land surface modelling and satellite data assimilation, Earth Syst. Sci. Data, 18, 3635–3669, <a href="https://doi.org/10.5194/essd-18-3635-2026" target="_blank">https://doi.org/10.5194/essd-18-3635-2026</a>, 2026.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib29"><label>Horowitz(2016)</label><mixed-citation>
      
Horowitz, C. A.: Paris Agreement, International Legal Materials, Cambridge University Press, 55,
740–755, <a href="https://doi.org/10.1017/S0020782900004253" target="_blank">https://doi.org/10.1017/S0020782900004253</a>, 2016.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib30"><label>Jacobs(1994)</label><mixed-citation>
      
Jacobs, C.: Direct impact of atmospheric CO2 enrichment on regional
transpiration, internal phd, wU thesis 1772 Proefschrift Wageningen, Netherlands, ISBN 9789054852506, <a href="https://doi.org/10.18174/206972" target="_blank">https://doi.org/10.18174/206972</a>, 1994.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib31"><label>Jacobs et al.(1996)Jacobs, Van den Hurk, and
De Bruin</label><mixed-citation>
      
Jacobs, C., Van den Hurk, B., and De Bruin, H.: Stomatal behaviour and
photosynthetic rate of unstressed grapevines in semi-arid conditions,
Agr. Forest Meteorol., 80, 111–134, 1996.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib32"><label>Jarlan et al.(2023)</label><mixed-citation>
      
Jarlan, L., Albergel, C., Bonan, B., Calvet, J.-C., De Rosnay, P., Ottlé, C., and Peylin, P.: Assimilation de données de télédétection pour le suivi des surfaces continentales, in: Inversion et assimilation de données de télédétection: Estimation des paramètres géophysiques, edited by: Yan, Y., ISTE, 45–95, <a href="https://doi.org/10.51926/ISTE.9142.ch2" target="_blank">https://doi.org/10.51926/ISTE.9142.ch2</a>, 2023.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib33"><label>Joiner et al.(2016)Joiner, Yoshida, Guanter, and
Middleton</label><mixed-citation>
      
Joiner, J., Yoshida, Y., Guanter, L., and Middleton, E. M.: New methods for the retrieval of chlorophyll red fluorescence from hyperspectral satellite instruments: simulations and application to GOME-2 and SCIAMACHY, Atmos. Meas. Tech., 9, 3939–3967, <a href="https://doi.org/10.5194/amt-9-3939-2016" target="_blank">https://doi.org/10.5194/amt-9-3939-2016</a>, 2016.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib34"><label>Jung et al.(2020)Jung, Schwalm, Migliavacca, Walther, Camps-Valls,
Koirala, Anthoni, Besnard, Bodesheim, Carvalhais, Chevallier, Gans, Goll,
Haverd, Köhler, Ichii, Jain, Liu, Lombardozzi, Nabel, Nelson, O'Sullivan,
Pallandt, Papale, Peters, Pongratz, Rödenbeck, Sitch, Tramontana, Walker,
Weber, and Reichstein</label><mixed-citation>
      
Jung, M., Schwalm, C., Migliavacca, M., Walther, S., Camps-Valls, G., Koirala, S., Anthoni, P., Besnard, S., Bodesheim, P., Carvalhais, N., Chevallier, F., Gans, F., Goll, D. S., Haverd, V., Köhler, P., Ichii, K., Jain, A. K., Liu, J., Lombardozzi, D., Nabel, J. E. M. S., Nelson, J. A., O'Sullivan, M., Pallandt, M., Papale, D., Peters, W., Pongratz, J., Rödenbeck, C., Sitch, S., Tramontana, G., Walker, A., Weber, U., and Reichstein, M.: Scaling carbon fluxes from eddy covariance sites to globe: synthesis and evaluation of the FLUXCOM approach, Biogeosciences, 17, 1343–1365, <a href="https://doi.org/10.5194/bg-17-1343-2020" target="_blank">https://doi.org/10.5194/bg-17-1343-2020</a>, 2020.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib35"><label>Köhler et al.(2015)Köhler, Guanter, and Joiner</label><mixed-citation>
      
Köhler, P., Guanter, L., and Joiner, J.: A linear method for the retrieval of sun-induced chlorophyll fluorescence from GOME-2 and SCIAMACHY data, Atmos. Meas. Tech., 8, 2589–2608, <a href="https://doi.org/10.5194/amt-8-2589-2015" target="_blank">https://doi.org/10.5194/amt-8-2589-2015</a>, 2015.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib36"><label>Köhler et al.(2018)Köhler, Frankenberg, Magney, Guanter, Joiner,
and Landgraf</label><mixed-citation>
      
Köhler, P., Frankenberg, C., Magney, T. S., Guanter, L., Joiner, J., and
Landgraf, J.: Global Retrievals of Solar-Induced Chlorophyll Fluorescence
With TROPOMI: First Results and Intersensor Comparison to OCO-2, Geophys.
Res. Lett., 45, 10456–10463,
<a href="https://doi.org/10.1029/2018GL079031" target="_blank">https://doi.org/10.1029/2018GL079031</a>, 2018.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib37"><label>Leroux et al.(2018)Leroux, Calvet, Munier, and Albergel</label><mixed-citation>
      
Leroux, D. J., Calvet, J.-C., Munier, S., and Albergel, C.: Using
Satellite-Derived Vegetation Products to Evaluate LDAS-Monde over the
Euro-Mediterranean Area, Remote Sens., 10, <a href="https://doi.org/10.3390/rs10081199" target="_blank">https://doi.org/10.3390/rs10081199</a>, 2018.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib38"><label>LOBELIA(2024)</label><mixed-citation>
      
LOBELIA: Revealing emission sources from space, Zenodo, <a href="https://doi.org/10.5281/zenodo.10837172" target="_blank">https://doi.org/10.5281/zenodo.10837172</a>, 2024.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib39"><label>Lunel et al.(2024)Lunel, Boone, and Le Moigne</label><mixed-citation>
      
Lunel, T., Boone, A. A., and Le Moigne, P.: Irrigation strongly influences  near-surface conditions and induces breeze circulation: Observational and  model-based evidence, Q. J. Roy. Meteor. Soc., 150, <a href="https://doi.org/10.1002/qj.4736" target="_blank">https://doi.org/10.1002/qj.4736</a>, 2024.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib40"><label>Mahfouf and Bilodeau(2009)</label><mixed-citation>
      
Mahfouf, J.-F. and Bilodeau, B.: A simple strategy for linearizing complex
moist convective schemes, Q. J. Roy. Meteor. Soc., 135, 953–962, <a href="https://doi.org/10.1002/qj.427" target="_blank">https://doi.org/10.1002/qj.427</a>, 2009.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib41"><label>Mahfouf et al.(2009)Mahfouf, Bergaoui, Draper, Bouyssel, Taillefer,
and Taseva</label><mixed-citation>
      
Mahfouf, J.-F., Bergaoui, K., Draper, C., Bouyssel, F., Taillefer, F., and
Taseva, L.: A comparison of two off-line soil analysis schemes for
assimilation of screen level observations, J. Geophys. Res.-Atmos., 114, <a href="https://doi.org/10.1029/2008JD011077" target="_blank">https://doi.org/10.1029/2008JD011077</a>, 2009.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib42"><label>Masson et al.(2013)Masson, Le Moigne, Martin, Faroux, Alias, Alkama,
Belamari, Barbu, Boone, Bouyssel et al.</label><mixed-citation>
      
Masson, V., Le Moigne, P., Martin, E., Faroux, S., Alias, A., Alkama, R., Belamari, S., Barbu, A., Boone, A., Bouyssel, F., Brousseau, P., Brun, E., Calvet, J.-C., Carrer, D., Decharme, B., Delire, C., Donier, S., Essaouini, K., Gibelin, A.-L., Giordani, H., Habets, F., Jidane, M., Kerdraon, G., Kourzeneva, E., Lafaysse, M., Lafont, S., Lebeaupin Brossier, C., Lemonsu, A., Mahfouf, J.-F., Marguinaud, P., Mokhtari, M., Morin, S., Pigeon, G., Salgado, R., Seity, Y., Taillefer, F., Tanguy, G., Tulet, P., Vincendon, B., Vionnet, V., and Voldoire, A.: The SURFEXv7.2 land and ocean surface platform for coupled or offline simulation of earth surface variables and fluxes, Geosci. Model Dev., 6, 929–960, <a href="https://doi.org/10.5194/gmd-6-929-2013" target="_blank">https://doi.org/10.5194/gmd-6-929-2013</a>, 2013.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib43"><label>Moya et al.(2004)Moya, Camenen, Evain, Goulas, Cerovic, Latouche,
Flexas, and Ounis</label><mixed-citation>
      
Moya, I., Camenen, L., Evain, S., Goulas, Y., Cerovic, Z., Latouche, G.,
Flexas, J., and Ounis, A.: A new instrument for passive remote sensing: 1.
Measurements of sunlight-induced chlorophyll fluorescence, Remote Sens.
Environ., 91, 186–197, <a href="https://doi.org/10.1016/j.rse.2004.02.012" target="_blank">https://doi.org/10.1016/j.rse.2004.02.012</a>,
2004.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib44"><label>Nelson et al.(2024)Nelson, Walther, Gans, Kraft, Weber, Novick,
Buchmann, Migliavacca, Wohlfahrt, Šigut, Ibrom, Papale, Göckede,
Duveiller, Knohl, Hörtnagl, Scott, Dušek, Zhang, Hamdi, Reichstein,
Aranda-Barranco, Ardö, Op de Beeck, Billesbach, Bowling, Bracho, Brümmer,
Camps-Valls, Chen, Cleverly, Desai, Dong, El-Madany, Euskirchen,
Feigenwinter, Galvagno, Gerosa, Gielen, Goded, Goslee, Gough, Heinesch,
Ichii, Jackowicz-Korczynski, Klosterhalfen, Knox, Kobayashi, Kohonen,
Korkiakoski, Mammarella, Gharun, Marzuoli, Matamala, Metzger, Montagnani,
Nicolini, O'Halloran, Ourcival, Peichl, Pendall, Ruiz Reverter, Roland,
Sabbatini, Sachs, Schmidt, Schwalm, Shekhar, Silberstein, Silveira, Spano,
Tagesson, Tramontana, Trotta, Turco, Vesala, Vincke, Vitale, Vivoni, Wang,
Woodgate, Yepez, Zhang, Zona, and Jung</label><mixed-citation>
      
Nelson, J. A., Walther, S., Gans, F., Kraft, B., Weber, U., Novick, K., Buchmann, N., Migliavacca, M., Wohlfahrt, G., Šigut, L., Ibrom, A., Papale, D., Göckede, M., Duveiller, G., Knohl, A., Hörtnagl, L., Scott, R. L., Dušek, J., Zhang, W., Hamdi, Z. M., Reichstein, M., Aranda-Barranco, S., Ardö, J., Op de Beeck, M., Billesbach, D., Bowling, D., Bracho, R., Brümmer, C., Camps-Valls, G., Chen, S., Cleverly, J. R., Desai, A., Dong, G., El-Madany, T. S., Euskirchen, E. S., Feigenwinter, I., Galvagno, M., Gerosa, G. A., Gielen, B., Goded, I., Goslee, S., Gough, C. M., Heinesch, B., Ichii, K., Jackowicz-Korczynski, M. A., Klosterhalfen, A., Knox, S., Kobayashi, H., Kohonen, K.-M., Korkiakoski, M., Mammarella, I., Gharun, M., Marzuoli, R., Matamala, R., Metzger, S., Montagnani, L., Nicolini, G., O'Halloran, T., Ourcival, J.-M., Peichl, M., Pendall, E., Ruiz Reverter, B., Roland, M., Sabbatini, S., Sachs, T., Schmidt, M., Schwalm, C. R., Shekhar, A., Silberstein, R., Silveira, M. L., Spano, D., Tagesson, T., Tramontana, G., Trotta, C., Turco, F., Vesala, T., Vincke, C., Vitale, D., Vivoni, E. R., Wang, Y., Woodgate, W., Yepez, E. A., Zhang, J., Zona, D., and Jung, M.: X-BASE: the first terrestrial carbon and water flux products from an extended data-driven scaling framework, FLUXCOM-X, Biogeosciences, 21, 5079–5115, <a href="https://doi.org/10.5194/bg-21-5079-2024" target="_blank">https://doi.org/10.5194/bg-21-5079-2024</a>, 2024.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib45"><label>Noilhan and Mahfouf(1996)</label><mixed-citation>
      
Noilhan, J. and Mahfouf, J.-F.: The ISBA land surface parameterisation scheme,
Global Planet. Change, 13, 145–159,
<a href="https://doi.org/10.1016/0921-8181(95)00043-7" target="_blank">https://doi.org/10.1016/0921-8181(95)00043-7</a>, 1996.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib46"><label>Noilhan and Planton(1989)</label><mixed-citation>
      
Noilhan, J. and Planton, S.: A Simple Parameterization of Land Surface
Processes for Meteorological Models, Mon. Weather Rev., 117, 536–549,
<a href="https://doi.org/10.1175/1520-0493(1989)117&lt;0536:ASPOLS&gt;2.0.CO;2" target="_blank">https://doi.org/10.1175/1520-0493(1989)117&lt;0536:ASPOLS&gt;2.0.CO;2</a>, 1989.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib47"><label>Rascher et al.(2015)</label><mixed-citation>
      
Rascher, U., Alonso, L., Burkart, A., Cilia, C., Cogliati, S., Colombo, R., Damm, A., Drusch, M., Guanter, L., Hanus, J., Hyvärinen, T., Julitta, T., Jussila, J., Kataja, K., Kokkalis, P., Kraft, S., Kraska, T., Matveeva, M., Moreno, J., Muller, O., Panigada, C., Pikl, M., Pinto, F., Prey, L., Pude, R., Rossini, M., Schickling, A., Schurr, U., Schüttemeyer, D., Verrelst, J., and Zemek, F.: Sun-induced fluorescence – a new probe of photosynthesis: First maps from the imaging spectrometer HyPlant, Glob. Change Biol., 21, 4673–4684, <a href="https://doi.org/10.1111/gcb.13017" target="_blank">https://doi.org/10.1111/gcb.13017</a>, 2015.


    </mixed-citation></ref-html>
<ref-html id="bib1.bib48"><label>Ricart et al.(2016)Ricart, Ribas, and Pavón</label><mixed-citation>
      
Ricart, S., Ribas, A., and Pavón, D.: Qualifying irrigation system
sustainability by means of stakeholder perceptions and concerns: lessons from
the Segarra-Garrigues Canal, Spain, Nat. Resour. Forum, 40, 77–90,
<a href="https://doi.org/10.1111/1477-8947.12097" target="_blank">https://doi.org/10.1111/1477-8947.12097</a>, 2016.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib49"><label>Richards(1931)</label><mixed-citation>
      
Richards, L. A.: Capillary conduction of liquids through porous mediums,
Physics, 1, 318–333, 1931.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib50"><label>Rüdiger et al.(2010)Rüdiger, Albergel, Mahfouf, Calvet, and
Walker</label><mixed-citation>
      
Rüdiger, C., Albergel, C., Mahfouf, J.-F., Calvet, J.-C., and Walker, J. P.:
Evaluation of the observation operator Jacobian for leaf area index data
assimilation with an extended Kalman filter, J. Geophys. Res.-Atmos., 115, <a href="https://doi.org/10.1029/2009JD012912" target="_blank">https://doi.org/10.1029/2009JD012912</a>, 2010.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib51"><label>Sabater et al.(2008)Sabater, Rüdiger, Calvet, Fritz, Jarlan, and
Kerr</label><mixed-citation>
      
Sabater, J. M., Rüdiger, C., Calvet, J.-C., Fritz, N., Jarlan, L., and Kerr,
Y.: Joint assimilation of surface soil moisture and LAI observations into a
land surface model, Agr. Forest Meteorol., 148, 1362–1373,
<a href="https://doi.org/10.1016/j.agrformet.2008.04.003" target="_blank">https://doi.org/10.1016/j.agrformet.2008.04.003</a>, 2008.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib52"><label>Trimmel et al.(2023)Trimmel, Hamer, Mayer, Schreier, Weihs,
Eitzinger, Sandén, Fitzky, Richter, Calvet, Bonan, Meurey, Vallejo,
Eckhardt, Santos, Oumami, Arteta, Marécal, Tarrasón, Karl, and
Rieder</label><mixed-citation>
      
Trimmel, H., Hamer, P., Mayer, M., Schreier, S. F., Weihs, P., Eitzinger, J.,
Sandén, H., Fitzky, A. C., Richter, A., Calvet, J.-C., Bonan, B., Meurey,
C., Vallejo, I., Eckhardt, S., Santos, G. S., Oumami, S., Arteta, J.,
Marécal, V., Tarrasón, L., Karl, T., and Rieder, H. E.: The influence of
vegetation drought stress on formaldehyde and ozone distributions over a
central European city, Atmos. Environ., 304, 119768,
<a href="https://doi.org/10.1016/j.atmosenv.2023.119768" target="_blank">https://doi.org/10.1016/j.atmosenv.2023.119768</a>, 2023.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib53"><label>Vanderbecken(2026a)Vanderbecken</label><mixed-citation>
      
Vanderbecken, P. J.: WP4 – supplementary data – Using deep learning to assimilate sun-induced fluorescence satellite observations in the ISBA land surface model: Datasets, Zenodo [data set], <a href="https://doi.org/10.5281/zenodo.18668100" target="_blank">https://doi.org/10.5281/zenodo.18668100</a>, 2026a.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib54"><label>Vanderbecken(2026b)Vanderbecken</label><mixed-citation>
      
Vanderbecken, P. J.: WP4 – supplementary data – Using deep learning to assimilate sun-induced fluorescence satellite observations in the ISBA land surface model: model, Zenodo [code], <a href="https://doi.org/10.5281/zenodo.18669437" target="_blank">https://doi.org/10.5281/zenodo.18669437</a>, 2026b.

    </mixed-citation></ref-html>
<ref-html id="bib1.bib55"><label>Veefkind et al.(2012)Veefkind, Aben, McMullan, Förster, de Vries,
Otter, Claas, Eskes, de Haan, Kleipool, van Weele, Hasekamp, Hoogeveen,
Landgraf, Snel, Tol, Ingmann, Voors, Kruizinga, Vink, Visser, and
Levelt</label><mixed-citation>
      
Veefkind, J., Aben, I., McMullan, K., Förster, H., de Vries, J., Otter, G.,
Claas, J., Eskes, H., de Haan, J., Kleipool, Q., van Weele, M., Hasekamp,
O., Hoogeveen, R., Landgraf, J., Snel, R., Tol, P., Ingmann, P., Voors, R.,
Kruizinga, B., Vink, R., Visser, H., and Levelt, P.: TROPOMI on the ESA
Sentinel-5 Precursor: A GMES mission for global observations of the
atmospheric composition for climate, air quality and ozone layer
applications, Remote Sens. Environ., 120, 70–83,
<a href="https://doi.org/10.1016/j.rse.2011.09.027" target="_blank">https://doi.org/10.1016/j.rse.2011.09.027</a>, 2012.

    </mixed-citation></ref-html>--></article>
