Department of Biology, San Diego State University, San Diego, CA, USA

Department of Animal and Plant Sciences, University of Sheffield, Western Bank, Sheffield, S10 2TN, UK

Department of Biology, San Diego State University, San Diego, CA, USA

Department of Land, Air and Water Resources, University of California, Davis, CA, 95616-8627, USA

Department of Animal and Plant Sciences, University of Sheffield, Western Bank, Sheffield, S10 2TN, UK

Water and Environmental Research Center, and International Arctic Research Center, University of Alaska Fairbanks, Fairbanks, AK, USA

Graduate School of Life and Environmental Sciences, Osaka Prefecture University, Sakai, Japan

Department of Environmental Science, Policy and Management, University of California Berkeley, Berkeley, CA, USA

Department of Biology, San Diego State University, San Diego, CA, USA