Articles | Volume 22, issue 18
https://doi.org/10.5194/bg-22-4903-2025
© Author(s) 2025. This work is distributed under
the Creative Commons Attribution 4.0 License.
the Creative Commons Attribution 4.0 License.
https://doi.org/10.5194/bg-22-4903-2025
© Author(s) 2025. This work is distributed under
the Creative Commons Attribution 4.0 License.
the Creative Commons Attribution 4.0 License.
Research article
| 
24 Sep 2025
Research article |
| 24 Sep 2025
| 24 Sep 2025
European tree cover during the Holocene reconstructed from pollen records
Luke Sweeney
CORRESPONDING AUTHOR
Geography and Environmental Science, School of Archaeology, Geography and Environmental Science (SAGES), University of Reading, Whiteknights, Reading, RG6 6AH, UK
Leverhulme Centre for Wildfires, Environment and Society, Imperial College London, South Kensington, London, SW7 2BW, UK
Sandy P. Harrison
Geography and Environmental Science, School of Archaeology, Geography and Environmental Science (SAGES), University of Reading, Whiteknights, Reading, RG6 6AH, UK
Leverhulme Centre for Wildfires, Environment and Society, Imperial College London, South Kensington, London, SW7 2BW, UK
Marc Vander Linden
Institute for the Modelling of Socio-Environmental Transitions, Bournemouth University, Christchurch House, Talbot Campus, Poole, BH12 5BB, UK
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Yicheng Shen, Luke Sweeney, Mengmeng Liu, Jose Antonio Lopez Saez, Sebastián Pérez-Díaz, Reyes Luelmo-Lautenschlaeger, Graciela Gil-Romera, Dana Hoefer, Gonzalo Jiménez-Moreno, Heike Schneider, I. Colin Prentice, and Sandy P. Harrison
Clim. Past, 18, 1189–1201, https://doi.org/10.5194/cp-18-1189-2022, https://doi.org/10.5194/cp-18-1189-2022, 2022
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We present a method to reconstruct burnt area using a relationship between pollen and charcoal abundances and the calibration of charcoal abundance using modern observations of burnt area. We use this method to reconstruct changes in burnt area over the past 12 000 years from sites in Iberia. We show that regional changes in burnt area reflect known changes in climate, with a high burnt area during warming intervals and low burnt area when the climate was cooler and/or wetter than today.
Sandy P. Harrison, Roberto Villegas-Diaz, Esmeralda Cruz-Silva, Daniel Gallagher, David Kesner, Paul Lincoln, Yicheng Shen, Luke Sweeney, Daniele Colombaroli, Adam Ali, Chéïma Barhoumi, Yves Bergeron, Tatiana Blyakharchuk, Přemysl Bobek, Richard Bradshaw, Jennifer L. Clear, Sambor Czerwiński, Anne-Laure Daniau, John Dodson, Kevin J. Edwards, Mary E. Edwards, Angelica Feurdean, David Foster, Konrad Gajewski, Mariusz Gałka, Michelle Garneau, Thomas Giesecke, Graciela Gil Romera, Martin P. Girardin, Dana Hoefer, Kangyou Huang, Jun Inoue, Eva Jamrichová, Nauris Jasiunas, Wenying Jiang, Gonzalo Jiménez-Moreno, Monika Karpińska-Kołaczek, Piotr Kołaczek, Niina Kuosmanen, Mariusz Lamentowicz, Martin Lavoie, Fang Li, Jianyong Li, Olga Lisitsyna, José Antonio López-Sáez, Reyes Luelmo-Lautenschlaeger, Gabriel Magnan, Eniko Katalin Magyari, Alekss Maksims, Katarzyna Marcisz, Elena Marinova, Jenn Marlon, Scott Mensing, Joanna Miroslaw-Grabowska, Wyatt Oswald, Sebastián Pérez-Díaz, Ramón Pérez-Obiol, Sanna Piilo, Anneli Poska, Xiaoguang Qin, Cécile C. Remy, Pierre J. H. Richard, Sakari Salonen, Naoko Sasaki, Hieke Schneider, William Shotyk, Migle Stancikaite, Dace Šteinberga, Normunds Stivrins, Hikaru Takahara, Zhihai Tan, Liva Trasune, Charles E. Umbanhowar, Minna Väliranta, Jüri Vassiljev, Xiayun Xiao, Qinghai Xu, Xin Xu, Edyta Zawisza, Yan Zhao, Zheng Zhou, and Jordan Paillard
Earth Syst. Sci. Data, 14, 1109–1124, https://doi.org/10.5194/essd-14-1109-2022, https://doi.org/10.5194/essd-14-1109-2022, 2022
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We provide a new global data set of charcoal preserved in sediments that can be used to examine how fire regimes have changed during past millennia and to investigate what caused these changes. The individual records have been standardised, and new age models have been constructed to allow better comparison across sites. The data set contains 1681 records from 1477 sites worldwide.
Kieran M. R. Hunt and Sandy P. Harrison
Clim. Past, 21, 1–26, https://doi.org/10.5194/cp-21-1-2025, https://doi.org/10.5194/cp-21-1-2025, 2025
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In this study, we train machine learning models on tree rings, speleothems, and instrumental rainfall to estimate seasonal monsoon rainfall over India over the last 500 years. Our models highlight multidecadal droughts in the mid-17th and 19th centuries, and we link these to historical famines. Using techniques from explainable AI (artificial intelligence), we show that our models use known relationships between local hydroclimate and the monsoon circulation.
Jierong Zhao, Boya Zhou, Sandy P. Harrison, and I. Colin Prentice
EGUsphere, https://doi.org/10.5194/egusphere-2024-3897, https://doi.org/10.5194/egusphere-2024-3897, 2025
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We used eco-evolutionary optimality modelling to examine how climate and CO2 impacted vegetation at the Last Glacial Maximum (LGM, 21,000 years ago) and the mid-Holocene (MH, 6,000 years ago). Low CO2 at the LGM was as important as climate in reducing tree cover and productivity, and increasing C4 plant abundance. Climate had positive effects on MH vegetation, but the low CO2 was a constraint on plant growth. These results show it is important to consider changing CO2 to model ecosystem changes.
Fang Li, Xiang Song, Sandy P. Harrison, Jennifer R. Marlon, Zhongda Lin, L. Ruby Leung, Jörg Schwinger, Virginie Marécal, Shiyu Wang, Daniel S. Ward, Xiao Dong, Hanna Lee, Lars Nieradzik, Sam S. Rabin, and Roland Séférian
Geosci. Model Dev., 17, 8751–8771, https://doi.org/10.5194/gmd-17-8751-2024, https://doi.org/10.5194/gmd-17-8751-2024, 2024
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This study provides the first comprehensive assessment of historical fire simulations from 19 Earth system models in phase 6 of the Coupled Model Intercomparison Project (CMIP6). Most models reproduce global totals, spatial patterns, seasonality, and regional historical changes well but fail to simulate the recent decline in global burned area and underestimate the fire response to climate variability. CMIP6 simulations address three critical issues of phase-5 models.
Nikita Kaushal, Franziska A. Lechleitner, Micah Wilhelm, Khalil Azennoud, Janica C. Bühler, Kerstin Braun, Yassine Ait Brahim, Andy Baker, Yuval Burstyn, Laia Comas-Bru, Jens Fohlmeister, Yonaton Goldsmith, Sandy P. Harrison, István G. Hatvani, Kira Rehfeld, Magdalena Ritzau, Vanessa Skiba, Heather M. Stoll, József G. Szűcs, Péter Tanos, Pauline C. Treble, Vitor Azevedo, Jonathan L. Baker, Andrea Borsato, Sakonvan Chawchai, Andrea Columbu, Laura Endres, Jun Hu, Zoltán Kern, Alena Kimbrough, Koray Koç, Monika Markowska, Belen Martrat, Syed Masood Ahmad, Carole Nehme, Valdir Felipe Novello, Carlos Pérez-Mejías, Jiaoyang Ruan, Natasha Sekhon, Nitesh Sinha, Carol V. Tadros, Benjamin H. Tiger, Sophie Warken, Annabel Wolf, Haiwei Zhang, and SISAL Working Group members
Earth Syst. Sci. Data, 16, 1933–1963, https://doi.org/10.5194/essd-16-1933-2024, https://doi.org/10.5194/essd-16-1933-2024, 2024
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Speleothems are a popular, multi-proxy climate archive that provide regional to global insights into past hydroclimate trends with precise chronologies. We present an update to the SISAL (Speleothem Isotopes
Synthesis and AnaLysis) database, SISALv3, which, for the first time, contains speleothem trace element records, in addition to an update to the stable isotope records available in previous versions of the database, cumulatively providing data from 365 globally distributed sites.
Synthesis and AnaLysis) database, SISALv3, which, for the first time, contains speleothem trace element records, in addition to an update to the stable isotope records available in previous versions of the database, cumulatively providing data from 365 globally distributed sites.
Mengmeng Liu, Iain Colin Prentice, and Sandy P. Harrison
Clim. Past Discuss., https://doi.org/10.5194/cp-2024-12, https://doi.org/10.5194/cp-2024-12, 2024
Preprint under review for CP
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Dansgaard-Oeschger events were large and rapid warming events that occurred multiple times during the last ice age. We show that changes in the northern extratropics and the southern extratropics were anti-phased, with warming over most of the north and cooling in the south. The reconstructions do not provide evidence for a change in seasonality in temperature. However, they do indicate that warming was generally accompanied by wetter conditions and cooling by drier conditions.
Huiying Xu, Han Wang, Iain Colin Prentice, and Sandy P. Harrison
Biogeosciences, 20, 4511–4525, https://doi.org/10.5194/bg-20-4511-2023, https://doi.org/10.5194/bg-20-4511-2023, 2023
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Leaf carbon (C) and nitrogen (N) are crucial elements in leaf construction and physiological processes. This study reconciled the roles of phylogeny, species identity, and climate in stoichiometric traits at individual and community levels. The variations in community-level leaf N and C : N ratio were captured by optimality-based models using climate data. Our results provide an approach to improve the representation of leaf stoichiometry in vegetation models to better couple N with C cycling.
Esmeralda Cruz-Silva, Sandy P. Harrison, I. Colin Prentice, Elena Marinova, Patrick J. Bartlein, Hans Renssen, and Yurui Zhang
Clim. Past, 19, 2093–2108, https://doi.org/10.5194/cp-19-2093-2023, https://doi.org/10.5194/cp-19-2093-2023, 2023
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We examined 71 pollen records (12.3 ka to present) in the eastern Mediterranean, reconstructing climate changes. Over 9000 years, winters gradually warmed due to orbital factors. Summer temperatures peaked at 4.5–5 ka, likely declining because of ice sheets. Moisture increased post-11 kyr, remaining high from 10–6 kyr before a slow decrease. Climate models face challenges in replicating moisture transport.
Olivia Haas, Iain Colin Prentice, and Sandy P. Harrison
Biogeosciences, 20, 3981–3995, https://doi.org/10.5194/bg-20-3981-2023, https://doi.org/10.5194/bg-20-3981-2023, 2023
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We quantify the impact of CO2 and climate on global patterns of burnt area, fire size, and intensity under Last Glacial Maximum (LGM) conditions using three climate scenarios. Climate change alone did not produce the observed LGM reduction in burnt area, but low CO2 did through reducing vegetation productivity. Fire intensity was sensitive to CO2 but strongly affected by changes in atmospheric dryness. Low CO2 caused smaller fires; climate had the opposite effect except in the driest scenario.
Giulia Mengoli, Sandy P. Harrison, and I. Colin Prentice
EGUsphere, https://doi.org/10.5194/egusphere-2023-1261, https://doi.org/10.5194/egusphere-2023-1261, 2023
Preprint archived
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Soil water availability affects plant carbon uptake by reducing leaf area and/or by closing stomata, which reduces its efficiency. We present a new formulation of how climatic dryness reduces both maximum carbon uptake and the soil-moisture threshold below which it declines further. This formulation illustrates how plants adapt their water conservation strategy to thrive in dry climates, and is step towards a better representation of soil-moisture effects in climate models.
Mengmeng Liu, Yicheng Shen, Penelope González-Sampériz, Graciela Gil-Romera, Cajo J. F. ter Braak, Iain Colin Prentice, and Sandy P. Harrison
Clim. Past, 19, 803–834, https://doi.org/10.5194/cp-19-803-2023, https://doi.org/10.5194/cp-19-803-2023, 2023
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We reconstructed the Holocene climates in the Iberian Peninsula using a large pollen data set and found that the west–east moisture gradient was much flatter than today. We also found that the winter was much colder, which can be expected from the low winter insolation during the Holocene. However, summer temperature did not follow the trend of summer insolation, instead, it was strongly correlated with moisture.
Yicheng Shen, Luke Sweeney, Mengmeng Liu, Jose Antonio Lopez Saez, Sebastián Pérez-Díaz, Reyes Luelmo-Lautenschlaeger, Graciela Gil-Romera, Dana Hoefer, Gonzalo Jiménez-Moreno, Heike Schneider, I. Colin Prentice, and Sandy P. Harrison
Clim. Past, 18, 1189–1201, https://doi.org/10.5194/cp-18-1189-2022, https://doi.org/10.5194/cp-18-1189-2022, 2022
Short summary
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We present a method to reconstruct burnt area using a relationship between pollen and charcoal abundances and the calibration of charcoal abundance using modern observations of burnt area. We use this method to reconstruct changes in burnt area over the past 12 000 years from sites in Iberia. We show that regional changes in burnt area reflect known changes in climate, with a high burnt area during warming intervals and low burnt area when the climate was cooler and/or wetter than today.
Sandy P. Harrison, Roberto Villegas-Diaz, Esmeralda Cruz-Silva, Daniel Gallagher, David Kesner, Paul Lincoln, Yicheng Shen, Luke Sweeney, Daniele Colombaroli, Adam Ali, Chéïma Barhoumi, Yves Bergeron, Tatiana Blyakharchuk, Přemysl Bobek, Richard Bradshaw, Jennifer L. Clear, Sambor Czerwiński, Anne-Laure Daniau, John Dodson, Kevin J. Edwards, Mary E. Edwards, Angelica Feurdean, David Foster, Konrad Gajewski, Mariusz Gałka, Michelle Garneau, Thomas Giesecke, Graciela Gil Romera, Martin P. Girardin, Dana Hoefer, Kangyou Huang, Jun Inoue, Eva Jamrichová, Nauris Jasiunas, Wenying Jiang, Gonzalo Jiménez-Moreno, Monika Karpińska-Kołaczek, Piotr Kołaczek, Niina Kuosmanen, Mariusz Lamentowicz, Martin Lavoie, Fang Li, Jianyong Li, Olga Lisitsyna, José Antonio López-Sáez, Reyes Luelmo-Lautenschlaeger, Gabriel Magnan, Eniko Katalin Magyari, Alekss Maksims, Katarzyna Marcisz, Elena Marinova, Jenn Marlon, Scott Mensing, Joanna Miroslaw-Grabowska, Wyatt Oswald, Sebastián Pérez-Díaz, Ramón Pérez-Obiol, Sanna Piilo, Anneli Poska, Xiaoguang Qin, Cécile C. Remy, Pierre J. H. Richard, Sakari Salonen, Naoko Sasaki, Hieke Schneider, William Shotyk, Migle Stancikaite, Dace Šteinberga, Normunds Stivrins, Hikaru Takahara, Zhihai Tan, Liva Trasune, Charles E. Umbanhowar, Minna Väliranta, Jüri Vassiljev, Xiayun Xiao, Qinghai Xu, Xin Xu, Edyta Zawisza, Yan Zhao, Zheng Zhou, and Jordan Paillard
Earth Syst. Sci. Data, 14, 1109–1124, https://doi.org/10.5194/essd-14-1109-2022, https://doi.org/10.5194/essd-14-1109-2022, 2022
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We provide a new global data set of charcoal preserved in sediments that can be used to examine how fire regimes have changed during past millennia and to investigate what caused these changes. The individual records have been standardised, and new age models have been constructed to allow better comparison across sites. The data set contains 1681 records from 1477 sites worldwide.
Alexander Kuhn-Régnier, Apostolos Voulgarakis, Peer Nowack, Matthias Forkel, I. Colin Prentice, and Sandy P. Harrison
Biogeosciences, 18, 3861–3879, https://doi.org/10.5194/bg-18-3861-2021, https://doi.org/10.5194/bg-18-3861-2021, 2021
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Along with current climate, vegetation, and human influences, long-term accumulation of biomass affects fires. Here, we find that including the influence of antecedent vegetation and moisture improves our ability to predict global burnt area. Additionally, the length of the preceding period which needs to be considered for accurate predictions varies across regions.
Sarah E. Parker, Sandy P. Harrison, Laia Comas-Bru, Nikita Kaushal, Allegra N. LeGrande, and Martin Werner
Clim. Past, 17, 1119–1138, https://doi.org/10.5194/cp-17-1119-2021, https://doi.org/10.5194/cp-17-1119-2021, 2021
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Regional trends in the oxygen isotope (δ18O) composition of stalagmites reflect several climate processes. We compare stalagmite δ18O records from monsoon regions and model simulations to identify the causes of δ18O variability over the last 12 000 years, and between glacial and interglacial states. Precipitation changes explain the glacial–interglacial δ18O changes in all monsoon regions; Holocene trends are due to a combination of precipitation, atmospheric circulation and temperature changes.
Masa Kageyama, Sandy P. Harrison, Marie-L. Kapsch, Marcus Lofverstrom, Juan M. Lora, Uwe Mikolajewicz, Sam Sherriff-Tadano, Tristan Vadsaria, Ayako Abe-Ouchi, Nathaelle Bouttes, Deepak Chandan, Lauren J. Gregoire, Ruza F. Ivanovic, Kenji Izumi, Allegra N. LeGrande, Fanny Lhardy, Gerrit Lohmann, Polina A. Morozova, Rumi Ohgaito, André Paul, W. Richard Peltier, Christopher J. Poulsen, Aurélien Quiquet, Didier M. Roche, Xiaoxu Shi, Jessica E. Tierney, Paul J. Valdes, Evgeny Volodin, and Jiang Zhu
Clim. Past, 17, 1065–1089, https://doi.org/10.5194/cp-17-1065-2021, https://doi.org/10.5194/cp-17-1065-2021, 2021
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The Last Glacial Maximum (LGM; ~21 000 years ago) is a major focus for evaluating how well climate models simulate climate changes as large as those expected in the future. Here, we compare the latest climate model (CMIP6-PMIP4) to the previous one (CMIP5-PMIP3) and to reconstructions. Large-scale climate features (e.g. land–sea contrast, polar amplification) are well captured by all models, while regional changes (e.g. winter extratropical cooling, precipitations) are still poorly represented.
Laia Comas-Bru, Kira Rehfeld, Carla Roesch, Sahar Amirnezhad-Mozhdehi, Sandy P. Harrison, Kamolphat Atsawawaranunt, Syed Masood Ahmad, Yassine Ait Brahim, Andy Baker, Matthew Bosomworth, Sebastian F. M. Breitenbach, Yuval Burstyn, Andrea Columbu, Michael Deininger, Attila Demény, Bronwyn Dixon, Jens Fohlmeister, István Gábor Hatvani, Jun Hu, Nikita Kaushal, Zoltán Kern, Inga Labuhn, Franziska A. Lechleitner, Andrew Lorrey, Belen Martrat, Valdir Felipe Novello, Jessica Oster, Carlos Pérez-Mejías, Denis Scholz, Nick Scroxton, Nitesh Sinha, Brittany Marie Ward, Sophie Warken, Haiwei Zhang, and SISAL Working Group members
Earth Syst. Sci. Data, 12, 2579–2606, https://doi.org/10.5194/essd-12-2579-2020, https://doi.org/10.5194/essd-12-2579-2020, 2020
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This paper presents an updated version of the SISAL (Speleothem Isotope Synthesis and Analysis) database. This new version contains isotopic data from 691 speleothem records from 294 cave sites and new age–depth models, including their uncertainties, for 512 speleothems.
Chris M. Brierley, Anni Zhao, Sandy P. Harrison, Pascale Braconnot, Charles J. R. Williams, David J. R. Thornalley, Xiaoxu Shi, Jean-Yves Peterschmitt, Rumi Ohgaito, Darrell S. Kaufman, Masa Kageyama, Julia C. Hargreaves, Michael P. Erb, Julien Emile-Geay, Roberta D'Agostino, Deepak Chandan, Matthieu Carré, Partrick J. Bartlein, Weipeng Zheng, Zhongshi Zhang, Qiong Zhang, Hu Yang, Evgeny M. Volodin, Robert A. Tomas, Cody Routson, W. Richard Peltier, Bette Otto-Bliesner, Polina A. Morozova, Nicholas P. McKay, Gerrit Lohmann, Allegra N. Legrande, Chuncheng Guo, Jian Cao, Esther Brady, James D. Annan, and Ayako Abe-Ouchi
Clim. Past, 16, 1847–1872, https://doi.org/10.5194/cp-16-1847-2020, https://doi.org/10.5194/cp-16-1847-2020, 2020
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This paper provides an initial exploration and comparison to climate reconstructions of the new climate model simulations of the mid-Holocene (6000 years ago). These use state-of-the-art models developed for CMIP6 and apply the same experimental set-up. The models capture several key aspects of the climate, but some persistent issues remain.
Cited articles
Abis, B. and Brovkin, V.: Environmental conditions for alternative tree-cover states in high latitudes, Biogeosciences, 14, 511–527, https://doi.org/10.5194/bg-14-511-2017, 2017.
Akaike, H.: A New Look at the Statistical Model Identification, IEEE T. Automat. Contr., 19, 716–723, https://doi.org/10.1109/TAC.1974.1100705, 1974.
Alkama, R. and Cescatti, A.: Biophysical climate impacts of recent changes in global forest cover, Science, 351, 600–604, https://doi.org/10.1126/science.aac8083, 2016.
Alley, R. B.: The Younger Dryas cold interval as viewed from central Greenland, Quaternary Sci. Rev., 19, 213–226, https://doi.org/10.1016/S0277-3791(99)00062-1, 2000.
Alley, R. B. and Ágústsdóttir, A. M.: The 8k event: Cause and consequences of a major Holocene abrupt climate change, Quaternary Sci. Rev., 24, 1123–1149, https://doi.org/10.1016/j.quascirev.2004.12.004, 2005.
Alley, R. B., Mayewski, P. A., Sowers, T., Stuiver, M., Taylor, K. C., and Clark, P. U.: Holocene climatic instability: A prominent, widespread event 8200 yr ago, Geology, 25, 483–486, https://doi.org/10.1130/0091-7613(1997)025<0483:HCIAPW>2.3.CO;2, 1997.
Bartlein, P. J., Harrison, S. P., and Izumi, K.: Underlying causes of Eurasian midcontinental aridity in simulations of mid-Holocene climate, Geophys. Res. Lett., 44, 9020–9028, https://doi.org/10.1002/2017GL074476, 2017.
Baston, D.: exactextractr: Fast Extraction from Raster Datasets using Polygons, exactextractr version 0.10.0, https://isciences.gitlab.io/exactextractr/ (last access: 25 April 2025), 2023.
Bini, M., Zanchetta, G., Perşoiu, A., Cartier, R., Català, A., Cacho, I., Dean, J. R., Di Rita, F., Drysdale, R. N., Finnè, M., Isola, I., Jalali, B., Lirer, F., Magri, D., Masi, A., Marks, L., Mercuri, A. M., Peyron, O., Sadori, L., Sicre, M.-A., Welc, F., Zielhofer, C., and Brisset, E.: The 4.2 ka BP Event in the Mediterranean region: an overview, Clim. Past, 15, 555–577, https://doi.org/10.5194/cp-15-555-2019, 2019.
Binney, H., Edwards, M., Macias-Fauria, M., Lozhkin, A., Anderson, P., Kaplan, J. O., Andreev, A., Bezrukova, E., Blyakharchuk, T., Jankovska, V., Khazina, I., Krivonogov, S., Kremenetski, K., Nield, J., Novenko, E., Ryabogina, N., Solovieva, N., Willis, K., and Zernitskaya, V.: Vegetation of Eurasia from the last glacial maximum to present: Key biogeographic patterns, Quaternary Sci. Rev., 157, 80–97, https://doi.org/10.1016/j.quascirev.2016.11.022, 2017.
Blaauw, M. and Christen, J. A.: Flexible paleoclimate age-depth models using an autoregressive gamma process, rbacon version 3.4.2, Bayesian Anal., 6, https://doi.org/10.1214/11-BA618, 2011.
Blaschek, M. and Renssen, H.: The Holocene thermal maximum in the Nordic Seas: the impact of Greenland Ice Sheet melt and other forcings in a coupled atmosphere–sea-ice–ocean model, Clim. Past, 9, 1629–1643, https://doi.org/10.5194/cp-9-1629-2013, 2013.
Bonan, G. B.: Forests and Climate Change: Forcings, Feedbacks, and the Climate Benefits of Forests, Science, 320, 1444–1449, https://doi.org/10.1126/science.1155121, 2008.
Bottema, S.: The Younger Dryas in the Eastern Mediterranean, Quaternay Sci. Rev., 14, 883–891, https://doi.org/10.1016/0277-3791(95)00069-0, 1995.
Braconnot, P., Zhu, D., Marti, O., and Servonnat, J.: Strengths and challenges for transient Mid- to Late Holocene simulations with dynamical vegetation, Clim. Past, 15, 997–1024, https://doi.org/10.5194/cp-15-997-2019, 2019.
Bradshaw, R. H. W. and Webb, T.: Relationships between Contemporary Pollen and Vegetation Data from Wisconsin and Michigan, USA, Ecology, 66, 721–737, https://doi.org/10.2307/1940533, 1985.
Buchhorn, M., Smets, B., Bertels, L., de Roo, B., Lesiv, M., Tsendbazar, N.-E., Li, L., and Tarko, A.: Copernicus Global Land Service: Land Cover 100 m: collection 3: epoch 2015: Globe, Zenodo [data set], https://doi.org/10.5281/zenodo.3939038, 2020a.
Buchhorn, M., Smets, B., Bertels, L., De Roo, B., Lesiv, M., Tsendbazar, N.-E., Herold, M., and Fritz, S.: Copernicus Global Land Service: Land Cover 100 m: collection 3: epoch 2016: Globe, Zenodo [data set], https://doi.org/10.5281/zenodo.3518026, 2020b.
Buchhorn, M., Smets, B., Bertels, L., De Roo, B., Lesiv, M., Tsendbazar, N.-E., Herold, M., and Fritz, S.: Copernicus Global Land Service: Land Cover 100 m: collection 3: epoch 2017: Globe, Zenodo [data set], https://doi.org/10.5281/zenodo.3518036, 2020c.
Buchhorn, M., Smets, B., Bertels, L., De Roo, B., Lesiv, M., Tsendbazar, N.-E., Herold, M., and Fritz, S.: Copernicus Global Land Service: Land Cover 100 m: collection 3: epoch 2018: Globe, Zenodo [data set], https://doi.org/10.5281/zenodo.3518038, 2020d.
Buchhorn, M., Smets, B., Bertels, L., De Roo, B., Lesiv, M., Tsendbazar, N.-E., Herold, M., and Fritz, S.: Copernicus Global Land Service: Land Cover 100 m: collection 3: epoch 2019: Globe, Zenodo [data set], https://doi.org/10.5281/zenodo.3939050, 2020e.
Bunting, M. J. and Farrell, M.: Do Local Habitat Conditions Affect Estimates of Relative Pollen Productivity and Source Area in Heathlands?, Front. Ecol. Evol., 10, 787345, https://doi.org/10.3389/fevo.2022.787345, 2022.
Bunting, M. J., Twiddle, C. L., and Middleton, R.: Using models of pollen dispersal and deposition in hilly landscapes: Some possible approaches, Palaeogeogr. Palaeocl., 259, 77–91, https://doi.org/10.1016/j.palaeo.2007.03.051, 2008.
Cheng, H., Zhang, H., Spötl, C., Baker, J., Sinha, A., Li, H., Bartolomeì, M., Moreno, A., Kathayat, G., Zhao, J., Dong, X., Li, Y., Ning, Y., Jia, X., Zong, B., Ait Brahim, Y., Pérez-Mejías, C., Cai, Y., Novello, V. F., Cruz, F. W., Severinghaus, J. P., An, Z., and Edwards, R. L.: Timing and structure of the Younger Dryas event and its underlying climate dynamics, P. Natl. Acad. Sci. USA, 117, 23408–23417, https://doi.org/10.1073/pnas.2007869117, 2020.
Cordova, C. E., Harrison, S. P., Mudie, P. J., Riehl, S., Leroy, S. A. G., and Ortiz, N.: Pollen, plant macrofossil and charcoal records for palaeovegetation reconstruction in the Mediterranean-Black Sea Corridor since the Last Glacial Maximum, Quaternary Int., 197, 12–26, https://doi.org/10.1016/j.quaint.2007.06.015, 2009.
Cox, D. R. and Snell, E. J.: Analysis of Binary Data, 2nd Edn., Chapman & Hall, ISBN 9780412306204, 1989.
Cribari-Neto, F. and Zeileis, A.: Beta Regression in R, betareg version 3.2-2, J. Stat. Softw., 34, https://doi.org/10.18637/jss.v034.i02, 2010.
Cruz-Silva, E., Harrison, S. P., Prentice, I. C., Marinova, E., Bartlein, P. J., Renssen, H., and Zhang, Y.: Pollen-based reconstructions of Holocene climate trends in the eastern Mediterranean region, Clim. Past, 19, 2093–2108, https://doi.org/10.5194/cp-19-2093-2023, 2023.
Dallmeyer, A., Claussen, M., Lorenz, S. J., and Shanahan, T.: The end of the African humid period as seen by a transient comprehensive Earth system model simulation of the last 8000 years, Clim. Past, 16, 117–140, https://doi.org/10.5194/cp-16-117-2020, 2020.
Davis, B. A. S., Brewer, S., Stevenson, A. C., and Guiot, J.: The temperature of Europe during the Holocene reconstructed from pollen data, Quaternary Sci. Rev., 22, 1701–1716, https://doi.org/10.1016/S0277-3791(03)00173-2, 2003.
Davis, B. A. S., Collins, P. M., and Kaplan, J. O.: The age and post-glacial development of the modern European vegetation: a plant functional approach based on pollen data, Veg. Hist. Archaeobot., 24, 303–317, https://doi.org/10.1007/s00334-014-0476-9, 2015.
Duffin, K. I. and Bunting, M. J.: Relative pollen productivity and fall speed estimates for southern African savanna taxa, Veg. Hist. Archaeobot., 17, 507–525, https://doi.org/10.1007/s00334-007-0101-2, 2008.
Eastwood, W. J., Roberts, N., Lamb, H. F., and Tibby, J. C.: Holocene environmental change in southwest Turkey: a palaeoecological record of lake and catchment-related changes, Quaternary Sci. Rev., 18, 671–695, https://doi.org/10.1016/S0277-3791(98)00104-8, 1999.
European Environment Agency: Biogeographical regions, https://sdi.eea.europa.eu/catalogue/srv/api/records/11db8d14-f167-4cd5-9205-95638dfd9618 (last access: 17 August 2023), 2016.
Fagerlind, F.: The real signification of pollen diagrams, Botaniska Notiser, 105, 185–224, 1952.
Fall, P. L.: Spatial patterns of atmospheric pollen dispersal in the Colorado Rocky Mountains, USA, Rev. Palaeobot. Palyno., 74, 293–313, https://doi.org/10.1016/0034-6667(92)90013-7, 1992.
FAO: Global Forest Resources Assessment 2020, FAO, https://doi.org/10.4060/ca9825en, 2020.
Fyfe, R. M., Woodbridge, J., and Roberts, N.: From forest to farmland: pollen-inferred land cover change across Europe using the pseudobiomization approach, Glob. Change Biol., 21, 1197–1212, https://doi.org/10.1111/gcb.12776, 2015.
Gaillard, M.-J., Sugita, S., Mazier, F., Trondman, A.-K., Broström, A., Hickler, T., Kaplan, J. O., Kjellström, E., Kokfelt, U., Kuneš, P., Lemmen, C., Miller, P., Olofsson, J., Poska, A., Rundgren, M., Smith, B., Strandberg, G., Fyfe, R., Nielsen, A. B., Alenius, T., Balakauskas, L., Barnekow, L., Birks, H. J. B., Bjune, A., Björkman, L., Giesecke, T., Hjelle, K., Kalnina, L., Kangur, M., van der Knaap, W. O., Koff, T., Lagerås, P., Latałowa, M., Leydet, M., Lechterbeck, J., Lindbladh, M., Odgaard, B., Peglar, S., Segerström, U., von Stedingk, H., and Seppä, H.: Holocene land-cover reconstructions for studies on land cover-climate feedbacks, Clim. Past, 6, 483–499, https://doi.org/10.5194/cp-6-483-2010, 2010.
Gaillard, M.-J., Githumbi, E., Achoundong, G., Lézine, A.-M., Hély, C., Lebamba, J., Marquer, L., Mazier, F., Li, F., and Sugita, S.: The challenge of pollen-based quantitative reconstruction of Holocene plant cover in tropical regions: A pilot study in Cameroon, in: Quaternary Vegetation Dynamics: the African Pollen Database, edited by: Runge, J., Gosling, W., Lézine, A.-M., and Scott, L., CRC Press, London, 259–279, ISBN 9780367755089, 2021.
Gil-Romera, G., García Antón, M., and Calleja, J. A.: The late Holocene palaeoecological sequence of Serranía de las Villuercas (southern Meseta, western Spain), Veg. Hist. Archaeobot., 17, 653–666, https://doi.org/10.1007/s00334-008-0146-x, 2008.
Githumbi, E., Fyfe, R., Gaillard, M.-J., Trondman, A.-K., Mazier, F., Nielsen, A.-B., Poska, A., Sugita, S., Woodbridge, J., Azuara, J., Feurdean, A., Grindean, R., Lebreton, V., Marquer, L., Nebout-Combourieu, N., Stančikaitė, M., Tanţău, I., Tonkov, S., Shumilovskikh, L., and LandClimII data contributors: European pollen-based REVEALS land-cover reconstructions for the Holocene: methodology, mapping and potentials, Earth Syst. Sci. Data, 14, 1581–1619, https://doi.org/10.5194/essd-14-1581-2022, 2022.
Gudmundsson, L., Bremnes, J. B., Haugen, J. E., and Engen-Skaugen, T.: Technical Note: Downscaling RCM precipitation to the station scale using statistical transformations – a comparison of methods, qmap version 1.0-6, Hydrol. Earth Syst. Sci., 16, 3383–3390, https://doi.org/10.5194/hess-16-3383-2012, 2012.
Guiot, J.: Methodology of the last climatic cycle reconstruction in France from pollen data, Palaeogeogr. Palaeocl., 80, 49–69, https://doi.org/10.1016/0031-0182(90)90033-4, 1990.
Hansen, M. C., Potapov, P. V., Moore, R., Hancher, M., Turubanova, S. A., Tyukavina, A., Thau, D., Stehman, S. V., Goetz, S. J., Loveland, T. R., Kommareddy, A., Egorov, A., Chini, L., Justice, C. O., and Townshend, J. R. G.: High-Resolution Global Maps of 21st-Century Forest Cover Change, Science, 342, 850–853, https://doi.org/10.1126/science.1244693, 2013.
Harrison, S. P. and Marinova, E.: EMBSeCBIO modern pollen biomisation, University of Reading [data set], https://doi.org/10.17864/1947.109, 2017.
Harrison, S. P., Gaillard, M.-J., Stocker, B. D., Vander Linden, M., Klein Goldewijk, K., Boles, O., Braconnot, P., Dawson, A., Fluet-Chouinard, E., Kaplan, J. O., Kastner, T., Pausata, F. S. R., Robinson, E., Whitehouse, N. J., Madella, M., and Morrison, K. D.: Development and testing scenarios for implementing land use and land cover changes during the Holocene in Earth system model experiments, Geosci. Model Dev., 13, 805–824, https://doi.org/10.5194/gmd-13-805-2020, 2020.
Harrison, S. P., Marinova, E., and Cruz-Silva, E.: EMBSeCBIO pollen database, University of Reading [data set], https://doi.org/10.17864/1947.309, 2021.
Harrison, S., Egbudom, M.-A., Liu, M., Cruz-Silva, E., and Sweeney, L.: The SPECIAL EPD Database: an expanded resource to document changes in vegetation and climate from pollen records from Europe, the Middle East and western Eurasia, University of Reading [data set], https://doi.org/10.17864/1947.001295, 2024.
Heaton, T. J., Köhler, P., Butzin, M., Bard, E., Reimer, R. W., Austin, W. E. N., Bronk Ramsey, C., Grootes, P. M., Hughen, K. A., Kromer, B., Reimer, P. J., Adkins, J., Burke, A., Cook, M. S., Olsen, J., and Skinner, L. C.: Marine20 – The Marine Radiocarbon Age Calibration Curve (0–55,000 cal BP), Radiocarbon, 62, 779–820, https://doi.org/10.1017/RDC.2020.68, 2020.
Hill, M. O.: Diversity and Evenness: A Unifying Notation and Its Consequences, Ecology, 54, 427–432, https://doi.org/10.2307/1934352, 1973.
Hill, T. R., Duthie, T. J., and Bunting, J.: Pollen productivity estimates from KwaZulu-Natal Drakensberg, South Africa, in: Quaternary Vegetation Dynamics: the African Pollen Database, edited by: Runge, J., Gosling, W., Lézine, A.-M., and Scott, L., CRC Press, London, 259–274, ISBN 9780367755089, 2021.
Jackson, S. T. and Williams, J. W.: Modern Analogs in Quaternary Paleoecology: Here Today, Gone Yesterday, Gone Tomorrow?, Annu. Rev. Earth Pl. Sc., 32, 495–537, https://doi.org/10.1146/annurev.earth.32.101802.120435, 2004.
Klein Goldewijk, K., Beusen, A., and Janssen, P.: Long-term dynamic modeling of global population and built-up area in a spatially explicit way: HYDE 3.1, Holocene, 20, 565–573, https://doi.org/10.1177/0959683609356587, 2010.
Klein Goldewijk, K., Beusen, A., Doelman, J., and Stehfest, E.: Anthropogenic land use estimates for the Holocene – HYDE 3.2, Earth Syst. Sci. Data, 9, 927–953, https://doi.org/10.5194/essd-9-927-2017, 2017.
Kling, M. M. and Ackerly, D. D.: Global wind patterns shape genetic differentiation, asymmetric gene flow, and genetic diversity in trees, P. Natl. Acad. Sci. USA, 118, https://doi.org/10.1073/pnas.2017317118, 2021.
Kuosmanen, N., Fang, K., Bradshaw, R. H., Clear, J. L., and Seppä, H.: Role of forest fires in Holocene stand-scale dynamics in the unmanaged taiga forest of northwestern Russia, Holocene, 24, 1503–1514, https://doi.org/10.1177/0959683614544065, 2014.
Leite-Filho, A. T., Costa, M. H., and Fu, R.: The southern Amazon rainy season: The role of deforestation and its interactions with large-scale mechanisms, Int. J. Climatol, 40, 2328–2341, https://doi.org/10.1002/joc.6335, 2020.
Liu, M., Shen, Y., González-Sampériz, P., Gil-Romera, G., ter Braak, C. J. F., Prentice, I. C., and Harrison, S. P.: Holocene climates of the Iberian Peninsula: pollen-based reconstructions of changes in the west–east gradient of temperature and moisture, Clim. Past, 19, 803–834, https://doi.org/10.5194/cp-19-803-2023, 2023.
Liu, Z., Otto-Bliesner, B. L., He, F., Brady, E. C., Tomas, R., Clark, P. U., Carlson, A. E., Lynch-Stieglitz, J., Curry, W., Brook, E., Erickson, D., Jacob, R., Kutzbach, J., and Cheng, J.: Transient Simulation of Last Deglaciation with a New Mechanism for Bølling-Allerød Warming, Science, 325, 310–314, https://doi.org/10.1126/science.1171041, 2009.
Loader, C.: Locfit: Local regression, likelihood and density estimation, locfit version 1.5-9.12, http://cran.r-project.org/package=locfit%5Cnpapers2://publication/uuid/E5C68958-7122-4093-A8E2-3108F9DF726A (last access: 25 April 2025), 2025.
Marinova, E., Harrison, S. P., Bragg, F., Connor, S., de Laet, V., Leroy, S. A. G., Mudie, P., Atanassova, J., Bozilova, E., Caner, H., Cordova, C., Djamali, M., Filipova-Marinova, M., Gerasimenko, N., Jahns, S., Kouli, K., Kotthoff, U., Kvavadze, E., Lazarova, M., Novenko, E., Ramezani, E., Röpke, A., Shumilovskikh, L., Tanţau, I., and Tonkov, S.: Pollen-derived biomes in the Eastern Mediterranean–Black Sea–Caspian-Corridor, J Biogeogr., 45, 484–499, https://doi.org/10.1111/jbi.13128, 2018.
Markgraf, V.: Pollen Dispersal in a Mountain Area, Grana, 19, 127–146, https://doi.org/10.1080/00173138009424995, 1980.
Marlon, J. R., Bartlein, P. J., Daniau, A.-L., Harrison, S. P., Maezumi, S. Y., Power, M. J., Tinner, W., and Vannière, B.: Global biomass burning: a synthesis and review of Holocene paleofire records and their controls, Quaternary Sci. Rev., 65, 5–25, https://doi.org/10.1016/j.quascirev.2012.11.029, 2013.
Marquer, L., Gaillard, M.-J., Sugita, S., Trondman, A.-K., Mazier, F., Nielsen, A. B., Fyfe, R. M., Odgaard, B. V., Alenius, T., Birks, H. J. B., Bjune, A. E., Christiansen, J., Dodson, J., Edwards, K. J., Giesecke, T., Herzschuh, U., Kangur, M., Lorenz, S., Poska, A., Schult, M., and Seppä, H.: Holocene changes in vegetation composition in northern Europe: why quantitative pollen-based vegetation reconstructions matter, Quaternary Sci. Rev., 90, 199–216, https://doi.org/10.1016/j.quascirev.2014.02.013, 2014.
Marquer, L., Gaillard, M.-J., Sugita, S., Poska, A., Trondman, A.-K., Mazier, F., Nielsen, A. B., Fyfe, R. M., Jönsson, A. M., Smith, B., Kaplan, J. O., Alenius, T., Birks, H. J. B., Bjune, A. E., Christiansen, J., Dodson, J., Edwards, K. J., Giesecke, T., Herzschuh, U., Kangur, M., Koff, T., Latałowa, M., Lechterbeck, J., Olofsson, J., and Seppä, H.: Quantifying the effects of land use and climate on Holocene vegetation in Europe, Quaternary Sci. Rev., 171, 20–37, https://doi.org/10.1016/j.quascirev.2017.07.001, 2017.
Marquer, L., Mazier, F., Sugita, S., Galop, D., Houet, T., Faure, E., Gaillard, M.-J., Haunold, S., de Munnik, N., Simonneau, A., De Vleeschouwer, F., and Le Roux, G.: Pollen-based reconstruction of Holocene land-cover in mountain regions: Evaluation of the Landscape Reconstruction Algorithm in the Vicdessos valley, northern Pyrenees, France, Quaternary Sci. Rev., 228, 106049, https://doi.org/10.1016/j.quascirev.2019.106049, 2020.
Moreira, F., Rego, F. C., and Ferreira, P. G.: Temporal (1958–1995) pattern of change in a cultural landscape of northwestern Portugal: implications for fire occurrence, Landsc. Ecol., 16, 557–567, https://doi.org/10.1023/A:1013130528470, 2001.
Mountain Research Initiative EDW Working Group: Elevation-dependent warming in mountain regions of the world, Nat. Clim. Change, 5, 424–430, https://doi.org/10.1038/nclimate2563, 2015.
Moyes, A. B., Germino, M. J., and Kueppers, L. M.: Moisture rivals temperature in limiting photosynthesis by trees establishing beyond their cold-edge range limit under ambient and warmed conditions, New Phytol., 207, 1005–1014, https://doi.org/10.1111/nph.13422, 2015.
Nielsen, A. B., Giesecke, T., Theuerkauf, M., Feeser, I., Behre, K.-E., Beug, H.-J., Chen, S.-H., Christiansen, J., Dörfler, W., Endtmann, E., Jahns, S., de Klerk, P., Kühl, N., Latałowa, M., Odgaard, B. V., Rasmussen, P., Stockholm, J. R., Voigt, R., Wiethold, J., and Wolters, S.: Quantitative reconstructions of changes in regional openness in north-central Europe reveal new insights into old questions, Quaternary Sci. Rev., 47, 131–149, https://doi.org/10.1016/j.quascirev.2012.05.011, 2012.
Ortu, E., de Beaulieu, J. L., Caramiello, R., and Siniscalco, C.: Late Glacial and Holocene vegetation dynamics at various altitudes in the Ellero Valley, Maritime Alps, northwestern Italy, Ecoscience, 15, 200–212, https://doi.org/10.2980/15-2-2926, 2008.
Ortu, E., David, F., and Peyron, O.: Pollen-inferred palaeoclimate reconstruction in the Alps during the Lateglacial and the early Holocene: how to estimate the effect of elevation and local parameters, J. Quaternary Sci., 25, 651–661, https://doi.org/10.1002/jqs.1335, 2010.
Overpeck, J. T., Webb, T., and Prentice, I. C.: Quantitative Interpretation of Fossil Pollen Spectra: Dissimilarity Coefficients and the Method of Modern Analogs, Quaternary Res., 23, 87–108, https://doi.org/10.1016/0033-5894(85)90074-2, 1985.
Piraquive Bermúdez, D., Theuerkauf, M., and Giesecke, T.: Towards quantifying changes in forest cover in the Araucaria forest-grassland mosaic in southern Brazil, Veg. Hist. Archaeobot., 31, 107–122, https://doi.org/10.1007/s00334-021-00841-2, 2022.
Pirzamanbein, B., Lindström, J., Poska, A., Sugita, S., Trondman, A.-K., Fyfe, R., Mazier, F., Nielsen, A. B., Kaplan, J. O., Bjune, A. E., Birks, H. J. B., Giesecke, T., Kangur, M., Latałowa, M., Marquer, L., Smith, B., and Gaillard, M.-J.: Creating spatially continuous maps of past land cover from point estimates: A new statistical approach applied to pollen data, Ecol. Complex., 20, 127–141, https://doi.org/10.1016/j.ecocom.2014.09.005, 2014.
Pouzet, P., Maanan, M., Piotrowska, N., Baltzer, A., Stéphan, P., and Robin, M.: Chronology of Holocene storm events along the European Atlantic coast, Prog. Phys. Geogr., 42, 431–450, https://doi.org/10.1177/0309133318776500, 2018.
Prentice, C.: Records of vegetation in time and space: the principles of pollen analysis, in: Vegetation history, Springer Netherlands, Dordrecht, 17–42, https://doi.org/10.1007/978-94-009-3081-0_2, 1988.
Prentice, I. C.: Pollen Representation, Source Area, and Basin Size: Toward a Unified Theory of Pollen Analysis, Quaternary Res., 23, 76–86, https://doi.org/10.1016/0033-5894(85)90073-0, 1985.
Prentice, I. C. and Webb, T.: Pollen percentages, tree abundances and the Fagerlind effect, J. Quaternary Sci., 1, 35–43, https://doi.org/10.1002/jqs.3390010105, 1986.
R Core Team: R: A Language and Environment for Statistical Computing, R Foundation for Statistical Computing, Vienna, Austria, version 4.3.3, https://www.r-project.org/ (last access: 25 April 2025), 2024.
Reimer, P. J., Austin, W. E. N., Bard, E., Bayliss, A., Blackwell, P. G., Bronk Ramsey, C., Butzin, M., Cheng, H., Edwards, R. L., Friedrich, M., Grootes, P. M., Guilderson, T. P., Hajdas, I., Heaton, T. J., Hogg, A. G., Hughen, K. A., Kromer, B., Manning, S. W., Muscheler, R., Palmer, J. G., Pearson, C., van der Plicht, J., Reimer, R. W., Richards, D. A., Scott, E. M., Southon, J. R., Turney, C. S. M., Wacker, L., Adolphi, F., Büntgen, U., Capano, M., Fahrni, S. M., Fogtmann-Schulz, A., Friedrich, R., Köhler, P., Kudsk, S., Miyake, F., Olsen, J., Reinig, F., Sakamoto, M., Sookdeo, A., and Talamo, S.: The IntCal20 Northern Hemisphere Radiocarbon Age Calibration Curve (0–55 cal kBP), Radiocarbon, 62, 1–33, https://doi.org/10.1017/rdc.2020.41, 2020.
Renssen, H., Seppä, H., Heiri, O., Roche, D. M., Goosse, H., and Fichefet, T.: The spatial and temporal complexity of the Holocene thermal maximum, Nat. Geosci., 2, 411–414, https://doi.org/10.1038/ngeo513, 2009.
Roberts, C. N., Woodbridge, J., Palmisano, A., Bevan, A., Fyfe, R., and Shennan, S.: Mediterranean landscape change during the Holocene: Synthesis, comparison and regional trends in population, land cover and climate, Holocene, 29, 923–937, https://doi.org/10.1177/0959683619826697, 2019.
Roberts, N., Fyfe, R. M., Woodbridge, J., Gaillard, M.-J., Davis, B. A. S., Kaplan, J. O., Marquer, L., Mazier, F., Nielsen, A. B., Sugita, S., Trondman, A.-K., and Leydet, M.: Europe's lost forests: a pollen-based synthesis for the last 11,000 years, Sci. Rep., 8, 716, https://doi.org/10.1038/s41598-017-18646-7, 2018.
Senf, C. and Seidl, R.: Mapping the forest disturbance regimes of Europe, Nat. Sustain., 4, 63–70, https://doi.org/10.1038/s41893-020-00609-y, 2020.
Simas, A. B., Barreto-Souza, W., and Rocha, A. V.: Improved estimators for a general class of beta regression models, Comput. Stat. Data Anal., 54, 348–366, https://doi.org/10.1016/j.csda.2009.08.017, 2010.
Sjöström, J. K., Gyllencreutz, R., Martínez Cortizas, A., Nylund, A., Piilo, S. R., Schenk, F., McKeown, M., Ryberg, E. E., and Kylander, M. E.: Holocene storminess dynamics in northwestern Ireland: Shifts in storm duration and frequency between the mid- and late Holocene, Quaternary Sci. Rev., 337, 108803, https://doi.org/10.1016/j.quascirev.2024.108803, 2024.
Staal, A., Tuinenburg, O. A., Bosmans, J. H. C., Holmgren, M., van Nes, E. H., Scheffer, M., Zemp, D. C., and Dekker, S. C.: Forest-rainfall cascades buffer against drought across the Amazon, Nat. Clim. Change, 8, 539–543, https://doi.org/10.1038/s41558-018-0177-y, 2018.
Sugita, S.: A Model of Pollen Source Area for an Entire Lake Surface, Quaternary Res., 39, 239–244, https://doi.org/10.1006/qres.1993.1027, 1993.
Sugita, S.: Pollen Representation of Vegetation in Quaternary Sediments: Theory and Method in Patchy Vegetation, J. Ecol., 82, 881, https://doi.org/10.2307/2261452, 1994.
Sugita, S.: Theory of quantitative reconstruction of vegetation I: Pollen from large sites REVEALS regional vegetation composition, Holocene, https://doi.org/10.1177/0959683607075837, 2007a.
Sugita, S.: Theory of quantitative reconstruction of vegetation II: all you need is LOVE, Holocene, 17, 243–257, https://doi.org/10.1177/0959683607075838, 2007b.
Sweeney, L.: sweeney-l/European_Holocene_tree_cover: 25/04/25 (v3.0.0), Zenodo [code, data set], https://doi.org/10.5281/zenodo.15283096, 2025.
Tabares, X., Ratzmann, G., Kruse, S., Theuerkauf, M., Mapani, B., and Herzschuh, U.: Relative pollen productivity estimates of savanna taxa from southern Africa and their application to reconstruct shrub encroachment during the last century, Holocene, 31, 1100–1111, https://doi.org/10.1177/09596836211003193, 2021.
Thorley, A.: Pollen Analytical Evidence Relating to the Vegetation History of the Chalk, J. Biogeogr., 8, 93–106, https://doi.org/10.2307/2844552, 1981.
Tong, Z.-Y., Wu, L.-Y., Feng, H.-H., Zhang, M., Armbruster, W. S., Renner, S. S., and Huang, S.-Q.: New calculations indicate that 90 % of flowering plant species are animal-pollinated, Natl. Sci. Rev., 10, https://doi.org/10.1093/nsr/nwad219, 2023.
Trondman, A.-K., Gaillard, M.-J., Mazier, F., Sugita, S., Fyfe, R. M., Nielsen, A. B., Twiddle, C., Barratt, P., Birks, H. J. B., Bjune, A. E., Björkman, L., Broström, A., Caseldine, C., David, R., Dodson, J., Dörfler, W., Fischer, E., Geel, B., Giesecke, T., Hultberg, T., Kalnina, L., Kangur, M., Knaap, P., Koff, T., Kuneš, P., Lagerås, P., Latałowa, M., Lechterbeck, J., Leroyer, C., Leydet, M., Lindbladh, M., Marquer, L., Mitchell, F. J. G., Odgaard, B. V., Peglar, S. M., Persson, T., Poska, A., Rösch, M., Seppä, H., Veski, S., and Wick, L.: Pollen-based quantitative reconstructions of Holocene regional vegetation cover (plant-functional types and land-cover types) in Europe suitable for climate modelling, Glob. Change Biol., 21, 676–697, https://doi.org/10.1111/gcb.12737, 2015.
Turubanova, S., Potapov, P., Hansen, M. C., Li, X., Tyukavina, A., Pickens, A. H., Hernandez-Serna, A., Arranz, A. P., Guerra-Hernandez, J., Senf, C., Häme, T., Valbuena, R., Eklundh, L., Brovkina, O., Navrátilová, B., Novotný, J., Harris, N., and Stolle, F.: Tree canopy extent and height change in Europe, 2001–2021, quantified using Landsat data archive, Remote Sens. Environ., 298, 113797, https://doi.org/10.1016/j.rse.2023.113797, 2023.
Viedma, O., Moity, N., and Moreno, J. M.: Changes in landscape fire-hazard during the second half of the 20th century: Agriculture abandonment and the changing role of driving factors, Agr. Ecosyst. Environ., 207, 126–140, https://doi.org/10.1016/j.agee.2015.04.011, 2015.
Villegas-Diaz, R. and Harrison, S. P.: The SPECIAL Modern Pollen Data Set for Climate Reconstructions, version 2 (SMPDSv2), University of Reading [data set], https://doi.org/10.17864/1947.000389, 2022.
Villegas-Diaz, R., Cruz-Silva, E., and Harrison, S. P.: ageR: Supervised Age Models, ageR version 0.2.0.900, Zenodo, https://doi.org/10.5281/zenodo.4636716, 2021.
Wei, D., González-Sampériz, P., Gil-Romera, G., Harrison, S. P., and Prentice, I. C.: Seasonal temperature and moisture changes in interior semi-arid Spain from the last interglacial to the Late Holocene, Quaternary Res., 101, 143–155, https://doi.org/10.1017/qua.2020.108, 2021.
Weiss, H., Courty, M.-A., Wetterstrom, W., Guichard, F., Senior, L., Meadow, R., and Curnow, A.: The Genesis and Collapse of Third Millennium North Mesopotamian Civilization, Science, 261, 995–1004, https://doi.org/10.1126/science.261.5124.995, 1993.
Whitney, B. S., Smallman, T. L., Mitchard, E. T., Carson, J. F., Mayle, F. E., and Bunting, M. J.: Constraining pollen-based estimates of forest cover in the Amazon: A simulation approach, Holocene, 29, 262–270, https://doi.org/10.1177/0959683618810394, 2018.
Wieczorek, M. and Herzschuh, U.: Compilation of relative pollen productivity (RPP) estimates and taxonomically harmonised RPP datasets for single continents and Northern Hemisphere extratropics, Earth Syst. Sci. Data, 12, 3515–3528, https://doi.org/10.5194/essd-12-3515-2020, 2020.
Wörl, V., Jetschni, J., and Jochner-Oette, S.: Birch Pollen Deposition and Transport along an Altitudinal Gradient in the Bavarian Alps – A Case Study Using Gravimetric Pollen Traps in the Pollen Season 2020, Atmosphere, 13, 2007, https://doi.org/10.3390/atmos13122007, 2022.
Zanon, M., Davis, B. A. S., Marquer, L., Brewer, S., and Kaplan, J. O.: European forest cover during the past 12,000 years: A palynological reconstruction based on modern analogs and remote sensing, Front. Plant Sci., 9, 1–25, https://doi.org/10.3389/fpls.2018.00253, 2018.
Zapolska, A., Serge, M. A., Mazier, F., Quiquet, A., Renssen, H., Vrac, M., Fyfe, R., and Roche, D. M.: More than agriculture: Analysing time-cumulative human impact on European land-cover of second half of the Holocene, Quaternary Sci. Rev., 314, 108227, https://doi.org/10.1016/j.quascirev.2023.108227, 2023.
Zhang, Y., Renssen, H., and Seppä, H.: Effects of melting ice sheets and orbital forcing on the early Holocene warming in the extratropical Northern Hemisphere, Clim. Past, 12, 1119–1135, https://doi.org/10.5194/cp-12-1119-2016, 2016.
Zhang, Y., Renssen, H., Seppä, H., and Valdes, P. J.: Holocene temperature trends in the extratropical Northern Hemisphere based on inter-model comparisons, J. Quaternary Sci., 33, 464–476, https://doi.org/10.1002/jqs.3027, 2018.
Short summary
Changes in tree cover across Europe during the Holocene are reconstructed from fossil pollen data using a model developed with modern observations of tree cover and modern pollen assemblages. There is a rapid increase in tree cover after the last glacial period, with maximum cover during the mid-Holocene and a decline thereafter; the timing of the maximum and the speed of the increase and subsequent decrease vary regionally, likely reflecting differences in climate trajectories and human influence.
Changes in tree cover across Europe during the Holocene are reconstructed from fossil pollen...
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