Department of Marine Microbiology, Netherlands Institute of Ecology – KNAW, P.O. Box 140, 4400 AC Yerseke, The Netherlands

present address: Marine Biological Laboratory, Institute of Biology, University of Copenhagen, Strandpromenaden 5, 3000 Helsingør, Denmark

Marine Biological Laboratory, Institute of Biology, University of Copenhagen, Strandpromenaden 5, 3000 Helsingør, Denmark

Marine Biological Laboratory, Institute of Biology, University of Copenhagen, Strandpromenaden 5, 3000 Helsingør, Denmark

Department of Environmental Biotechnology, TU Delft, Julianalaan 67, 2628 BC Delft, The Netherlands

Department of Environmental Biotechnology, TU Delft, Julianalaan 67, 2628 BC Delft, The Netherlands

Department of Marine Microbiology, Netherlands Institute of Ecology – KNAW, P.O. Box 140, 4400 AC Yerseke, The Netherlands

Department of Marine Microbiology, Netherlands Institute of Ecology – KNAW, P.O. Box 140, 4400 AC Yerseke, The Netherlands

present address: AE3 Consultancy, Fuchsialaan 8, 4401HV Yerseke, The Netherlands